Nitidulidae 317 Nomina Insecta Nearctica
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Elytra Reduction May Affect the Evolution of Beetle Hind Wings
Zoomorphology https://doi.org/10.1007/s00435-017-0388-1 ORIGINAL PAPER Elytra reduction may affect the evolution of beetle hind wings Jakub Goczał1 · Robert Rossa1 · Adam Tofilski2 Received: 21 July 2017 / Revised: 31 October 2017 / Accepted: 14 November 2017 © The Author(s) 2017. This article is an open access publication Abstract Beetles are one of the largest and most diverse groups of animals in the world. Conversion of forewings into hardened shields is perceived as a key adaptation that has greatly supported the evolutionary success of this taxa. Beetle elytra play an essential role: they minimize the influence of unfavorable external factors and protect insects against predators. Therefore, it is particularly interesting why some beetles have reduced their shields. This rare phenomenon is called brachelytry and its evolution and implications remain largely unexplored. In this paper, we focused on rare group of brachelytrous beetles with exposed hind wings. We have investigated whether the elytra loss in different beetle taxa is accompanied with the hind wing shape modification, and whether these changes are similar among unrelated beetle taxa. We found that hind wings shape differ markedly between related brachelytrous and macroelytrous beetles. Moreover, we revealed that modifications of hind wings have followed similar patterns and resulted in homoplasy in this trait among some unrelated groups of wing-exposed brachelytrous beetles. Our results suggest that elytra reduction may affect the evolution of beetle hind wings. Keywords Beetle · Elytra · Evolution · Wings · Homoplasy · Brachelytry Introduction same mechanism determines wing modification in all other insects, including beetles. However, recent studies have The Coleoptera order encompasses almost the quarter of all provided evidence that formation of elytra in beetles is less currently known animal species (Grimaldi and Engel 2005; affected by Hox gene than previously expected (Tomoyasu Hunt et al. -
A Catalogue of Coleoptera Specimens with Potential Forensic Interest in the Goulandris Natural History Museum Collection
ENTOMOLOGIA HELLENICA Vol. 25, 2016 A catalogue of Coleoptera specimens with potential forensic interest in the Goulandris Natural History Museum collection Dimaki Maria Goulandris Natural History Museum, 100 Othonos St. 14562 Kifissia, Greece Anagnou-Veroniki Maria Makariou 13, 15343 Aghia Paraskevi (Athens), Greece Tylianakis Jason Zoology Department, University of Canterbury, Private Bag 4800, Christchurch, New Zealand http://dx.doi.org/10.12681/eh.11549 Copyright © 2017 Maria Dimaki, Maria Anagnou- Veroniki, Jason Tylianakis To cite this article: Dimaki, M., Anagnou-Veroniki, M., & Tylianakis, J. (2016). A catalogue of Coleoptera specimens with potential forensic interest in the Goulandris Natural History Museum collection. ENTOMOLOGIA HELLENICA, 25(2), 31-38. doi:http://dx.doi.org/10.12681/eh.11549 http://epublishing.ekt.gr | e-Publisher: EKT | Downloaded at 27/12/2018 06:22:38 | ENTOMOLOGIA HELLENICA 25 (2016): 31-38 Received 15 March 2016 Accepted 12 December 2016 Available online 3 February 2017 A catalogue of Coleoptera specimens with potential forensic interest in the Goulandris Natural History Museum collection MARIA DIMAKI1’*, MARIA ANAGNOU-VERONIKI2 AND JASON TYLIANAKIS3 1Goulandris Natural History Museum, 100 Othonos St. 14562 Kifissia, Greece 2Makariou 13, 15343 Aghia Paraskevi (Athens), Greece 3Zoology Department, University of Canterbury, Private Bag 4800, Christchurch, New Zealand ABSTRACT This paper presents a catalogue of the Coleoptera specimens in the Goulandris Natural History Museum collection that have potential forensic interest. Forensic entomology can help to estimate the time elapsed since death by studying the necrophagous insects collected on a cadaver and its surroundings. In this paper forty eight species (369 specimens) are listed that belong to seven families: Silphidae (3 species), Staphylinidae (6 species), Histeridae (11 species), Anobiidae (4 species), Cleridae (6 species), Dermestidae (14 species), and Nitidulidae (4 species). -
Keys to Families of Beetles in America North of Mexico
816 · Key to Families Keys to Families of Beetles in America North of Mexico by Michael A. Ivie hese keys are specifically designed for North American and, where possible, overly long lists of options, but when nec- taxa and may lead to incorrect identifications of many essary, I have erred on the side of directing the user to a correct Ttaxa from outside this region. They are aimed at the suc- identification. cessful family placement of all beetles in North America north of No key will work on all specimens because of abnormalities Mexico, and as such will not always be simple to use. A key to the of development, poor preservation, previously unknown spe- most common 50% of species in North America would be short cies, sexes or variation, or simple errors in characterization. Fur- and simple to use. However, after an initial learning period, most thermore, with more than 30,000 species to be considered, there coleopterists recognize those groups on sight, and never again are undoubtedly rare forms that escaped my notice and even key them out. It is the odd, the rare and the exceptional that make possibly some common and easily collected species with excep- a complex key necessary, and it is in its ability to correctly place tional characters that I overlooked. While this key should work those taxa that a key is eventually judged. Although these keys for at least 95% of specimens collected and 90% of North Ameri- build on many previous successful efforts, especially those of can species, the specialized collector who delves into unique habi- Crowson (1955), Arnett (1973) and Borror et al. -
Coleoptera: Scarabaeidae)
Systematic Entomology (2005), 31, 113–144 DOI: 10.1111/j.1365-3113.2005.00307.x The phylogeny of Sericini and their position within the Scarabaeidae based on morphological characters (Coleoptera: Scarabaeidae) DIRK AHRENS Deutsches Entomologisches Institut im Zentrum fu¨r Agrarlandschafts- und Landnutzungsforschung Mu¨ncheberg, Germany Abstract. To reconstruct the phylogeny of the Sericini and their systematic position among the scarabaeid beetles, cladistic analyses were performed using 107 morphological characters from the adults and larvae of forty-nine extant scarabaeid genera. Taxa represent most ‘traditional’ subfamilies of coprophagous and phytophagous Scarabaeidae, with emphasis on the Sericini and other melo- lonthine lineages. Several poorly studied exoskeletal features have been examined, including the elytral base, posterior wing venation, mouth parts, endosternites, coxal articulation, and genitalia. The results of the analysis strongly support the monophyly of the ‘orphnine group’ þ ‘melolonthine group’ including phytopha- gous scarabs such as Dynastinae, Hopliinae, Melolonthinae, Rutelinae, and Cetoniinae. This clade was identified as the sister group to the ‘dung beetle line’ represented by Aphodius þ Copris. The ‘melolonthine group’ is comprised in the strict consensus tree by two major clades and two minor lineages, with the included taxa of Euchirinae, Rutelinae, and Dynastinae nested together in one of the major clades (‘melolonthine group I’). Melolonthini, Cetoniinae, and Rutelinae are strongly supported, whereas Melolonthinae and Pachydemini appear to be paraphyletic. Sericini þ Ablaberini were identified to be sister taxa nested within the second major melolonthine clade (‘melolonthine group II’). As this clade is distributed primarily in the southern continents, one could assume that Sericini þ Ablaberini are derived from a southern lineage. -
Coleoptera: Tenebrionoidea)
ACTA ENTOMOLOGICA MUSEI NATIONALIS PRAGAE Published 30.vi.2010 Volume 50(1), pp. 157–166 ISSN 0374-1036 A review of Ripiphoridae in the Arabian Peninsula (Coleoptera: Tenebrionoidea) Jan BATELKA Nad Vodovodem 16, CZ-100 00 Praha 10, Czech Republic; e-mail: [email protected] Abstract. Distribution of the Ripiphoridae (Coleoptera: Tenebrionoidea) in the Arabian Peninsula is evaluated. Six species belonging to the genera Macrosiagon Hentz, 1830 and Ripiphorus Bosc, 1791 are fi gured and keyed, and the distribution of each species is mapped. Including new and previously published records, the Ripiphoridae are now reported from 19 localities of the Arabian Peninsula and offshore islands. Coordinates for each exact locality are given. Key words. Coleoptera, Tenebrionoidea, Ripiphoridae, Ripiphorinae, Macrosia- gon, Ripiphorus, faunistics, Arabian Peninsula, Palaearctic Region Introduction The Ripiphoridae (Coleoptera: Tenebrionoidea) are a cosmopolitan group of parasitoids, whose biogeography is only poorly understood. Only scarce distributional data are usually available because of their cryptic way of life in larval stages and short-lived adults. The Arabian Peninsula plays an important role in the understanding of their distribution in the Old World, as it is a transitional zone among three main zoogeographical realms: Afrotropical, Oriental and Palaearctic. The aim of this paper is to provide basis for further studies of the Ripiphoridae in this part of Asia and to make further research easier for those students who are not familiar with these rarely collected beetles. Each species is keyed and fi gured based on specimens collected in the Arabian Peninsula, with emphasis on colour variability and also on sexual dimorphism where appropriate. -
The Beetle Fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): Diversity and Distribution
INSECTA MUNDI, Vol. 20, No. 3-4, September-December, 2006 165 The beetle fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): Diversity and distribution Stewart B. Peck Department of Biology, Carleton University, 1125 Colonel By Drive, Ottawa, Ontario K1S 5B6, Canada stewart_peck@carleton. ca Abstract. The beetle fauna of the island of Dominica is summarized. It is presently known to contain 269 genera, and 361 species (in 42 families), of which 347 are named at a species level. Of these, 62 species are endemic to the island. The other naturally occurring species number 262, and another 23 species are of such wide distribution that they have probably been accidentally introduced and distributed, at least in part, by human activities. Undoubtedly, the actual numbers of species on Dominica are many times higher than now reported. This highlights the poor level of knowledge of the beetles of Dominica and the Lesser Antilles in general. Of the species known to occur elsewhere, the largest numbers are shared with neighboring Guadeloupe (201), and then with South America (126), Puerto Rico (113), Cuba (107), and Mexico-Central America (108). The Antillean island chain probably represents the main avenue of natural overwater dispersal via intermediate stepping-stone islands. The distributional patterns of the species shared with Dominica and elsewhere in the Caribbean suggest stages in a dynamic taxon cycle of species origin, range expansion, distribution contraction, and re-speciation. Introduction windward (eastern) side (with an average of 250 mm of rain annually). Rainfall is heavy and varies season- The islands of the West Indies are increasingly ally, with the dry season from mid-January to mid- recognized as a hotspot for species biodiversity June and the rainy season from mid-June to mid- (Myers et al. -
From Characters of the Female Reproductive Tract
Phylogeny and Classification of Caraboidea Mus. reg. Sci. nat. Torino, 1998: XX LCE. (1996, Firenze, Italy) 107-170 James K. LIEBHERR and Kipling W. WILL* Inferring phylogenetic relationships within Carabidae (Insecta, Coleoptera) from characters of the female reproductive tract ABSTRACT Characters of the female reproductive tract, ovipositor, and abdomen are analyzed using cladi stic parsimony for a comprehensive representation of carabid beetle tribes. The resulting cladogram is rooted at the family Trachypachidae. No characters of the female reproductive tract define the Carabidae as monophyletic. The Carabidac exhibit a fundamental dichotomy, with the isochaete tri bes Metriini and Paussini forming the adelphotaxon to the Anisochaeta, which includes Gehringiini and Rhysodini, along with the other groups considered member taxa in Jeannel's classification. Monophyly of Isochaeta is supported by the groundplan presence of a securiform helminthoid scle rite at the spermathecal base, and a rod-like, elongate laterotergite IX leading to the explosion cham ber of the pygidial defense glands. Monophyly of the Anisochaeta is supported by the derived divi sion of gonocoxa IX into a basal and apical portion. Within Anisochaeta, the evolution of a secon dary spermatheca-2, and loss ofthe primary spermathcca-I has occurred in one lineage including the Gehringiini, Notiokasiini, Elaphrini, Nebriini, Opisthiini, Notiophilini, and Omophronini. This evo lutionary replacement is demonstrated by the possession of both spermatheca-like structures in Gehringia olympica Darlington and Omophron variegatum (Olivier). The adelphotaxon to this sper matheca-2 clade comprises a basal rhysodine grade consisting of Clivinini, Promecognathini, Amarotypini, Apotomini, Melaenini, Cymbionotini, and Rhysodini. The Rhysodini and Clivinini both exhibit a highly modified laterotergite IX; long and thin, with or without a clavate lateral region. -
The Evolution and Genomic Basis of Beetle Diversity
The evolution and genomic basis of beetle diversity Duane D. McKennaa,b,1,2, Seunggwan Shina,b,2, Dirk Ahrensc, Michael Balked, Cristian Beza-Bezaa,b, Dave J. Clarkea,b, Alexander Donathe, Hermes E. Escalonae,f,g, Frank Friedrichh, Harald Letschi, Shanlin Liuj, David Maddisonk, Christoph Mayere, Bernhard Misofe, Peyton J. Murina, Oliver Niehuisg, Ralph S. Petersc, Lars Podsiadlowskie, l m l,n o f l Hans Pohl , Erin D. Scully , Evgeny V. Yan , Xin Zhou , Adam Slipinski , and Rolf G. Beutel aDepartment of Biological Sciences, University of Memphis, Memphis, TN 38152; bCenter for Biodiversity Research, University of Memphis, Memphis, TN 38152; cCenter for Taxonomy and Evolutionary Research, Arthropoda Department, Zoologisches Forschungsmuseum Alexander Koenig, 53113 Bonn, Germany; dBavarian State Collection of Zoology, Bavarian Natural History Collections, 81247 Munich, Germany; eCenter for Molecular Biodiversity Research, Zoological Research Museum Alexander Koenig, 53113 Bonn, Germany; fAustralian National Insect Collection, Commonwealth Scientific and Industrial Research Organisation, Canberra, ACT 2601, Australia; gDepartment of Evolutionary Biology and Ecology, Institute for Biology I (Zoology), University of Freiburg, 79104 Freiburg, Germany; hInstitute of Zoology, University of Hamburg, D-20146 Hamburg, Germany; iDepartment of Botany and Biodiversity Research, University of Wien, Wien 1030, Austria; jChina National GeneBank, BGI-Shenzhen, 518083 Guangdong, People’s Republic of China; kDepartment of Integrative Biology, Oregon State -
Of the Galapagos Islands, Ecuador
Belgian Journal ofEntomology 5 (2003) : 89-102 A review of the Oedemeridae (Coleoptera) of the Galapagos Islands, Ecuador Stewart B. PECK and Joyce COOK Department of Biology, Carleton University, 1125 Colonel By Drive, Ottawa, K1S 5B6, Canada (e-mail: ste'[email protected]). Abstract Extensive new collections contribute new information on the identity and distribution of the oedemerid beetles of the Galiipagos Islands. Specimens previously recorded as near Oxacis pilosa CHAMPION are descn'bed as Oxycopis galapagoensis sp. n. Oxacis pilosa CHAMPION of Guatemala and Nicaragua is transferred to the genus Oxycopis. Hypasclera collenettei (BLAIR) is the most common and widespread species in the islands, and is variable in that it shows significant differences in aedeagus morphology between separate islands. Alloxacis hoodi V AN DYKE is found be a synonym of H. collenettei. H. seymourensis (MUTCHLER) is known only from the central islands. Paroxacis galapagoensis (LINELL) is also widespread. All four Galapagos species are presently considered to be endemic, and each represents a separate ancestral colonization of the archipelago. Keywords: · Hypasclera, Oxycopis, Paroxacis, island insects, endemic species, colonization. Introduction Members of the beetle family Oedemeridae are commonly called the false blister beetles. Adults are found frequently at lights or by sweeping vegetation, and they are obligate pollen feeders (AR.NETT, 1984). Larvae may feed on plant roots or may be inhabitants of moist decaying wood and some may live in salt-soaked driftwood (ARNETT, 1984, KrusKA, 2002). Oedemerids have been described and reported from the Galapagos by several workers: BLAIR (1928; 1933); F'RANZ (1985); LINELL (1898); MUTCHLER (1938); and VAN DYKE (1953). -
Diversidad De Cantharidae, Lampyridae
Revista Mexicana de Biodiversidad 80: 675- 686, 2009 Diversidad de Cantharidae, Lampyridae, Lycidae, Phengodidae y Telegeusidae (Coleoptera: Elateroidea) en un bosque tropical caducifolio de la sierra de San Javier, Sonora, México Diversity of Cantharidae, Lampyridae, Lycidae, Phengodidae and Telegeusidae (Coleoptera: Elateroidea) in a tropical dry forest of the Sierra San Javier, Sonora, Mexico Santiago Zaragoza-Caballero1* y Enrique Ramírez-García2 1Laboratorio de Entomología, Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México. Apartado postal 70-153, 04510 México D. F., México. 2Estación de Biología Chamela, Instituto de Biología, Universidad Nacional Autónoma de México. Apartado postal 21, San Patricio 48980 Jalisco, México. *Correspondencia: [email protected] Resumen. Se presenta un estudio de la diversidad faunística de las familias Cantharidae, Lampyridae, Lycidae, Phengodidae y Telegeusidae (Coleoptera: Elateroidea), presentes en un bosque tropical caducifolio de la sierra de San Javier, Sonora, México, que corresponde al límite boreal de este biotopo en América. La recolección incluyó trampas de atracción luminosa y red entomológica aérea, se realizó en noviembre de 2003, febrero y abril de 2004, y de julio a octubre de ese mismo año, durante 5 días de cada mes. Comprende la época lluviosa (julio-octubre) y la temporada seca (noviembre-abril). Se capturó un total de 1 501 individuos que representan 30 especies. La familia más abundante fue Cantharidae con 696 individuos, seguida de Lycidae con 561, Lampyridae con 166, Phengodidae con 66 y Telegeusidae con 12. La más rica en especies fue Lycidae con 12, seguida de Cantharidae con 11, Lampyridae con 3, Phengodidae con 3 y Telegeusidae con 1. -
Coleoptera: Nitidulidae) De Coahuila, México
Escarabajos de la savia (Coleoptera: Nitidulidae) de Coahuila, México. HERMELINDO HERNÁNDEZ TORRES TESIS PRESENTADA COMO REQUISITO PARCIAL PARA OBTENER EL GRADO DE MAESTRO EN CIENCIAS EN PARASITOLOGÍA AGRÍCOLA UNIVERSIDAD AUTÓNOMA AGRARIA ANTONIO NARRO Buenavista, Saltillo, Coahuila, México Marzo, 2013 i ii DEDICATORIA A DIOS TODOPODEROSO. Por ser mi padre y confidente y regalarme cada maravilloso día para cumplir cada uno de mis propósitos y por permitirme culminar con éxito el esfuerzo de estos años de estudio. Para Él mi agradecimiento infinito. iii A LA MEMORIA DE MI MADRE ANGELA HERNÀNDEZ CASTILLO Que desde el Cielo está conmigo y que siempre recordaré, amaré y llevaré en mi corazón. A mi familia: Pedro Hernández Reyes Mario Hernández Castillo Álvaro Hernández Castillo Adela Hernández Hernández Y el pequeño Michel. Por los agradables momentos que pasamos juntos. A La M.C. Ave María Hernández López por su compañía, amor y respeto. A LA UNIVERSIDAD Y A MIS CATEDRÁTICOS. Especialmente al Dr. Oswaldo García Martínez, con afecto, respeto y admiración. iv AGRADECIMIENTOS Agradezco a Dios por protegerme durante todo mi camino y darme fuerzas para superar obstáculos y dificultades a lo largo de toda mi vida. Al Dr. Oswaldo García Martínez. Primeramente por confiar en mí, por brindarme su apoyo incondicional en la realización de esta investigación y sus grandes enseñanzas recibidas. Gracias, Dios lo bendiga siempre. A la M.C. Ave María Hernández López. Gracias por estar a mi lado siempre, por tu apoyo incondicional y respeto. Al M.C. Víctor M. Sánchez V., M.C. Jorge Corrales R. y M.C. Sofía Comparan S. -
Kenai National Wildlife Refuge Species List, Version 2018-07-24
Kenai National Wildlife Refuge Species List, version 2018-07-24 Kenai National Wildlife Refuge biology staff July 24, 2018 2 Cover image: map of 16,213 georeferenced occurrence records included in the checklist. Contents Contents 3 Introduction 5 Purpose............................................................ 5 About the list......................................................... 5 Acknowledgments....................................................... 5 Native species 7 Vertebrates .......................................................... 7 Invertebrates ......................................................... 55 Vascular Plants........................................................ 91 Bryophytes ..........................................................164 Other Plants .........................................................171 Chromista...........................................................171 Fungi .............................................................173 Protozoans ..........................................................186 Non-native species 187 Vertebrates ..........................................................187 Invertebrates .........................................................187 Vascular Plants........................................................190 Extirpated species 207 Vertebrates ..........................................................207 Vascular Plants........................................................207 Change log 211 References 213 Index 215 3 Introduction Purpose to avoid implying