DOCSLIB.ORG

On the New Fossil Genera and Species of Neuroptera (Insecta) from the Late Jurassic of Northeast China *

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text

Load more

mZ1;f1: m4M ~4&J?t~*1i Vol. 21. No.4 19961f,10fj ACTA ZOOTAXONOMICA SINICA Oct. , 1996 ON THE NEW FOSSIL GENERA AND SPECIES OF NEUROPTERA (INSECTA) FROM THE LATE JURASSIC OF NORTHEAST CHINA * REN Dong GUO Zi-guang (Geological Museum 0/ China. Beijing 100034) Abstract Five new genera and nine new species of the Upper Jurassic Neuroptera are de­ scribed and illustrated: Kalligramma liaoningensis sp. nov .• Sophogramma papilionacea gen. et sp. nov. , S. eucalla gen. et sp. nov. ,·S. plecophlebia gen. et sp. nov. Call belonging to the Kalligrammatidae); Lasiosmylus newi gen. et sp. nov. COsmylidae); Oloberotha sinica gen. et sp. nov. (Berothidae); Lembochrysa miniscula gen. et sp. nov., L. polyneura gen. et sp. nov. (both belonging to Chrysopidae); Choromyrmeleon othneius gen. et sp. nov. (Myrmeleon­ tidae). All the specimens described here were collected from the Yixian Formation of Beipiao City, Liaoning Province, P. R. China. Key words Neuroptera, Kalligrammatidae, Osmylidae, Berothidae, Chrysopidae, Myrmeleontidae, fossil, new genus, new species. A large collection of well-preserved fossil insects has recently been recovered by the au­ thors from the Late Jurassic Yixian Formation, located in Chaomidian Village, about 25km south-east of Beipiao City, in the west of Liaoning Province, China. Among which a large number of remains are assigned to the Neuroptera, being very common in the lacustrine sed­ iments of Yixian Formation from western Liaoning. This paper is to describe five new gen­ era and nine new species of fossil Neuroptera and to compare their characters. most of the wing venations, with homologous characters of extinct and extant groups. The insect-bear­ ing strata are of Late Jurassic age, their population and geographical distribution have been published (Ren, 1994a; Ren,1994b; Ren and Hong, 1994; Ren et at. , 1995). All the specimens described in this paper are deposited in the Geological Museum of China, Beijing. Family Kalligrammatidae Handlirsch, 1906 Genus Sophogramma gen. nov. Type species: Sophogramma papilionacea gen. et sp. nov. * This project is supported by the National Natural Science Foundation of China (NSFC Grant No. 49302019). Accepted Oct. 2,1995. 461 462 21 !(i: Diagnosis Forewing eye-spot absent. Humeral recurrent vein (Vr) present. All costal veinlets sinuate and forked. Rs with about 12-15 primary branches, many of them deeply forked. MA with 5 parallel pectinate branches. A basal cell (be) present between MP1 and MP2 at base of wing. MP with basal fork far proximal and with no additional forkings before vicinity of wing margin. CuA pectinately forked and terminated at hind margin of wing. CuP single, with a few branches near wing margin. lA running parallel to CuP for a long distance and with numerous branches. Species composition: three species. Discussion This new genus undoubtedly should be assigned to family Kalligram­ matidae (Carpenter, 1992; Martynova, 1947; Ponomarenko, 1984; Panfilov, 1968, 1980) according to the following characters: forewing with costal area of moderate width; vein Sc joining R near wing apex; MA extensively branched, sending several main branches anteri­ orly; crossveins numerous over entire wing. Up to now, 6 previously described extinct genera belong to this family as follows: Kalligramma Walther. 1904; Kalligrammula Handlirsch, 1919; Meioneurites Handlirsch, 1906 (all from the Jurassic of Germany and Kazakhstan); Lithogramma Panfilov, 1968; Kalligrammins Panfilov, 1980 (both from the Upper Jurassic of Kazakhstan); Angara­ gramma Ponomarenko, 1984 (from the Middle and Upper Jurassic of TransbaikaD. This new genus is distinguished from all the known genera by the combination of diag­ nosis of characters mentioned above ,especially by a obvious recurrent vein which also devel­ oped in Berothidae (Macleod and Adams, 1967 ), Psychopsidae ( Macleod, 1970; New, 1988), Polystoechotidae (Borror et al. , 1976). As far as now known, the new genus is the first recorded fossil Kalligrammtids from China. Sophogramma papilionacea gen. et sp. nov. (Fig. 1,10: 1) Holotype: a rather well-preserved almost complete forewing, registration No. LB95001. Horizon and type locality: Yixian Formation, Upper Jurassic, near Chaomidian Village, Beipiao City, Liaoning Province. People's Republic of China. Etymology From the Greek "papilion-"-"butterfly-like". Description Left forewing length 7 5mm, width 43mm. Rs with 12 branches, most of them forked deeply. MA curved and pectinately forked. MP forked near the base. Anterior and posterior MP parallel, both forked apically. MP 1 and MP 2 forming a loop or cell (be) basally. CuA pectinately forked, with 4 main branches. CuP forked apically. lA with 6 pectinately branches. 2A appearing dichotomously forked. 3A simple. The characters of forewing venation as shown in Fig. 1 in detail. Hindwings and other body parts unknown. Sophogramma plecophlebia gen. et sp. nov. (Fig. 2,11: 1) Holotype one rather well preserved complete forewing ,registration No. LB95009. 463 c lOmm MP, MP, CuA CuP L.. Fig. 1 Sophogramma papilionacea gen. et sp. nov. forewing, holotype, LB95001 Horizon and type l-ocality: same as Sophogramma papilionacea gen. et sp. nov. Etymology From the Greek "plec-" and "-phlebia"-"net vein". Description Forewing length 62mm, width 35mm. Posterior margin of wing mlSS- MA Fig. 2 Sophogrammaplecophlebia gen. et sp."nov. forewing, holotype, LB95009 464 21 ~ ing. Humeral recurrent vein distinctly present. Costal veinlets mostly forked. Rs with 12 branches, mostly forked. The basal and second branches of Rs forked distad to forking point of MA. CuA only with (as preserved) 1 branch. The details of forewing venation as shown in Fig. 2. Discussion This new species can be easily distinguished from the S. papilionacea gen. et sp. nov. by the basal and second branches of Rs forked more distally and by the CuA with 1 branch. Sophogramma eucalla gen. et sp. nov. (Fig. 3,11: 3) Holotype: two rather well-preserved incomplete forewing, registration No. LB9 5002, LB95003, part and counterpart. Horizon and type locality same as Sophogramma pacilionacea gen. et sp. nov. Etymology From the Greek "eucall-" - "wonderful". Description Forewing length, as preserved, 7Smm, width 35mm. Forewing with a rounded but slightly pointed apex, without maculations. The distal part of posterior margin of wing missing. Sc and Rl fused apically and thence curved posterially to enter margin be­ fore wing apex. Rs with 15 branches, except first one, the 2nd to 5th branches forked deeply. The first and 2nd pectinate branches of MA forked deeply. CuA with more than 7 pectinate branches. The characters of forewing venation as shown in Fig. 3. CuP CuA MP, MP, Fig. 3 Sophogramma eucalla gen. et sp. nov. forewing, holotype, LB95002 Discussion The new species differs from the type species S. papilionacea mainly by the first basal branch of Rs forked later, wider CuA area with more than 7 pectinate branches. It also can be separated from the S. plecophlebia by Rs with 15 branches, and the number and arrangement of pectinate branches of CuA. 465 Genus Kalligramma Wal ther, 1904 Kalligramma liaoningensis sp. nov. (Fig. 4,10: 2) Holotype two rather well-preserved complete forewing. registration No. LB95004, LB95005, part and counterpart. Horizon and type locality same as Sophogramma papilionacea gen. et sp. nov. Etymology From the fossil locality "Liaoning Province". Description Forewing length 73mm. width 43mm. Crossveins in costal space mostly simple, but a -few forked. Sc+ Rl simple. Rs with 9 branches, which dichotomously forked near the margin. MA dichotomously forked at the centre of wing. MP pectinately forked, with 5 branches. Cu forked near the base. CuA dichotomously forked. CuP pectinately forked, with 6 main branches. 1A appeared 3 main pectinate branches. 2A with at least 8 main pectinate branches. Wing eye-spot present. ~"~Sc+RI Rs MA CuP CuA Fig. 4 Kalligramma liaoningensis sp. nov. forewing, holotype, LB95004 Discussion 5 species of Kalligramma Walther. 1904 have been described as follow: K. haeckeli Walther, 1904 (hindwing from Upper Jurassic of Germany); K. turutanovae Martynova, 1947 (forewing from Upper Jurassic of Kazakhstan); K. multinerve Panfilov, 1968 (forewing). K. ftexuosum Panfilov. 1968 (hindwing) and K. sharovi Panfilov, 1968 (hindwing) (all from the Upper Jurassic of Kazakhstan). The new species is much similar to K. sharovi, but differs from the latter in 9 branches of Rs, the shape of wing eye-spot and CuA with only 2 branches. 466 21 ~ Family Osmylidae Leach, 1815 (Sensu Pafilov, 1980) Subfamily Spilosmylinae Kruger, 1913 Genus Lasiosmylus gen. nov. Type species: Lasiosmylus newi gen. et sp. nov. Diagnosis Pronotum . cordiform. Wings slender, not falcate, with relatively a few crossveins. Vein let in basal and middle part of costal area not forked. Crossveins absent be­ tween Rl and Rs. Rs with 7 main branches, forewing with 2 subcostal crossveins. MP forked opposite to the separation of MA from Rs. Species composition: type species. Discussion According to Panfilov (980), the Osmylidae contains Osmylitidae Mar­ tynova, 1949 and Mesopolystoechotidae Martynova, 1949. Up to the present, only 4 os­ mylitids and 1 previously known Mesopolystoechotid have been reported: Osmylites Haase, 1890; Kirgisellodes Martynov. 1937; Mesomylina Bode, 1953 and Mesopolystoechus Mar­ tynov, 1937 (all of them from Jurassic of Europe); Sinosmylites Hong, 1983 (from the Middle Jurassic of China). The new genus differs from them by the following combination of characters:
Recommended publications
  • UFRJ a Paleoentomofauna Brasileira

    UFRJ a Paleoentomofauna Brasileira

    Anuário do Instituto de Geociências - UFRJ www.anuario.igeo.ufrj.br A Paleoentomofauna Brasileira: Cenário Atual The Brazilian Fossil Insects: Current Scenario Dionizio Angelo de Moura-Júnior; Sandro Marcelo Scheler & Antonio Carlos Sequeira Fernandes Universidade Federal do Rio de Janeiro, Programa de Pós-Graduação em Geociências: Patrimônio Geopaleontológico, Museu Nacional, Quinta da Boa Vista s/nº, São Cristóvão, 20940-040. Rio de Janeiro, RJ, Brasil. E-mails: [email protected]; [email protected]; [email protected] Recebido em: 24/01/2018 Aprovado em: 08/03/2018 DOI: http://dx.doi.org/10.11137/2018_1_142_166 Resumo O presente trabalho fornece um panorama geral sobre o conhecimento da paleoentomologia brasileira até o presente, abordando insetos do Paleozoico, Mesozoico e Cenozoico, incluindo a atualização das espécies publicadas até o momento após a última grande revisão bibliográica, mencionando ainda as unidades geológicas em que ocorrem e os trabalhos relacionados. Palavras-chave: Paleoentomologia; insetos fósseis; Brasil Abstract This paper provides an overview of the Brazilian palaeoentomology, about insects Paleozoic, Mesozoic and Cenozoic, including the review of the published species at the present. It was analiyzed the geological units of occurrence and the related literature. Keywords: Palaeoentomology; fossil insects; Brazil Anuário do Instituto de Geociências - UFRJ 142 ISSN 0101-9759 e-ISSN 1982-3908 - Vol. 41 - 1 / 2018 p. 142-166 A Paleoentomofauna Brasileira: Cenário Atual Dionizio Angelo de Moura-Júnior; Sandro Marcelo Schefler & Antonio Carlos Sequeira Fernandes 1 Introdução Devoniano Superior (Engel & Grimaldi, 2004). Os insetos são um dos primeiros organismos Algumas ordens como Blattodea, Hemiptera, Odonata, Ephemeroptera e Psocopera surgiram a colonizar os ambientes terrestres e aquáticos no Carbonífero com ocorrências até o recente, continentais (Engel & Grimaldi, 2004).
  • From Chewing to Sucking Via Phylogeny—From Sucking to Chewing Via Ontogeny: Mouthparts of Neuroptera

    From Chewing to Sucking Via Phylogeny—From Sucking to Chewing Via Ontogeny: Mouthparts of Neuroptera

    Chapter 11 From Chewing to Sucking via Phylogeny—From Sucking to Chewing via Ontogeny: Mouthparts of Neuroptera Dominique Zimmermann, Susanne Randolf, and Ulrike Aspöck Abstract The Neuroptera are highly heterogeneous endopterygote insects. While their relatives Megaloptera and Raphidioptera have biting mouthparts also in their larval stage, the larvae of Neuroptera are characterized by conspicuous sucking jaws that are used to imbibe fluids, mostly the haemolymph of prey. They comprise a mandibular and a maxillary part and can be curved or straight, long or short. In the pupal stages, a transformation from the larval sucking to adult biting and chewing mouthparts takes place. The development during metamorphosis indicates that the larval maxillary stylet contains the Anlagen of different parts of the adult maxilla and that the larval mandibular stylet is a lateral outgrowth of the mandible. The mouth- parts of extant adult Neuroptera are of the biting and chewing functional type, whereas from the Mesozoic era forms with siphonate mouthparts are also known. Various food sources are used in larvae and in particular in adult Neuroptera. Morphological adaptations of the mouthparts of adult Neuroptera to the feeding on honeydew, pollen and arthropods are described in several examples. New hypoth- eses on the diet of adult Nevrorthidae and Dilaridae are presented. 11.1 Introduction The order Neuroptera, comprising about 5820 species (Oswald and Machado 2018), constitutes together with its sister group, the order Megaloptera (about 370 species), and their joint sister group Raphidioptera (about 250 species) the superorder Neuropterida. Neuroptera, formerly called Planipennia, are distributed worldwide and comprise 16 families of extremely heterogeneous insects.

  • A New Genus and Species of Thorny Lacewing from Upper Cretaceous Kuji Amber, Northeastern Japan (Neuroptera, Rhachiberothidae)

    A peer-reviewed open-access journal ZooKeys 802: 109–120 (2018) A new genus and species of thorny lacewing... 109 doi: 10.3897/zookeys.802.28754 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research A new genus and species of thorny lacewing from Upper Cretaceous Kuji amber, northeastern Japan (Neuroptera, Rhachiberothidae) Hiroshi Nakamine1, Shûhei Yamamoto2 1 Minoh Park Insect Museum, Minoh Park 1-18, Minoh City, Osaka, 562-0002, Japan 2 Integrative Research Center, Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA Corresponding author: Hiroshi Nakamine ([email protected]) Academic editor: S. Winterton | Received 31 July 2018 | Accepted 15 October 2018 | Published 4 December 2018 http://zoobank.org/407331A3-C2B3-4FDF-BA4D-D47ACDDA9BEF Citation: Nakamine H, Yamamoto S (2018) A new genus and species of thorny lacewing from Upper Cretaceous Kuji amber, northeastern Japan (Neuroptera, Rhachiberothidae). ZooKeys 802: 109–120. https://doi.org/10.3897/ zookeys.802.28754 Abstract Kujiberotha teruyukii gen. et sp. n., a remarkable new genus and species of Rhachiberothidae, is described from Upper Cretaceous amber from the Kuji area in northeastern Japan. This discovery represents the first record of this family both from Japan and from East Asia. This fossil taxon has the largest foreleg in the subfamily Paraberothinae found to date and its discovery implies that this group had higher morphologi- cal diversity in the Cretaceous than it does now. This finding also stresses the importance of the insect inclusions in Kuji amber, which have not been well explored in spite of their potential abundance.
  • X Revisiok of the Nearctic Hemerobiidae, Berothidae, Sisyridae, Polys'i'oechotidae and Dilaridae (Neuroptera)

    X Revisiok of the Nearctic Hemerobiidae, Berothidae, Sisyridae, Polys'i'oechotidae and Dilaridae (Neuroptera)

    X REVISIOK OF THE NEARCTIC HEMEROBIIDAE, BEROTHIDAE, SISYRIDAE, POLYS'I'OECHOTIDAE AND DILARIDAE (NEUROPTERA) Received February 28, 1940 Presented March 13, 1940 The insects treated in this revision are among and Professor H. 13. Hungerford, University of the most typical of the Seuroptera (Planipennia). Kansas. I am under special obligation to Pro- Khen the family Hemerobiidae was established by fessor R. C. Smith, of Kansas State College, for Leach (1815), it included nearly all of the insects the loan of his extensive private collection of these now comprising the order. Since the beginning of insects, and for the opportunity of seeing many the present century, however, and subsequent to small collections sent to him for identification dur- the publication of Banks' "Revision of the Ne- ing the preparation of this revision. I am deeply ~irctic Hemerobiidae" (1905b), various genera grateful to llr. D. E. Kimmins of the British have been removed from the Hemerobiidae and lluseum of Satural History for placing at my placed in separate families. The groips thus disposal the type specimens in the Lfuseum col- formed (Berothidae, Sisyridae, Polystoechotidae, lection, and for making detailed comparisons with Dilnridae) have little in common with the re- Illclachlan's types, which were received at the stricted family Hemerobiidae; but in order that RIuseum after my visit there in 1938. As on the scope of the present revision be kept identical previous occasions I am indebted to Professor nith that of Banks', these families have also been Banks for many helpful suggestions and criticisms. included. The morphology of the Xeuroptera in general During the course of this revisional study, I and of most of the families occurring in the Ne- have examined somewhat more than eight thou- arctic region has been extensively treated by Kil- sand individuals of the families mentioned.
  • (Neuroptera: Berothidae) from Baltic Amber

    (Neuroptera: Berothidae) from Baltic Amber

    Zootaxa 3946 (3): 401–415 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2015 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3946.3.7 http://zoobank.org/urn:lsid:zoobank.org:pub:237D932A-88EB-42D6-B3F3-C97C3BB9ECEF An important new fossil genus of Berothinae (Neuroptera: Berothidae) from Baltic amber VLADIMIR N. MAKARKIN1,3 & MICHAEL OHL2 1Institute of Biology and Soil Sciences, Far East Branch of the Russian Academy of Sciences, Vladivostok, 960022, Russia 2Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse, 43, D-10115 Berlin, Germany. E-mail: [email protected] 3Corresponding author. E-mail: [email protected] Abstract Elektroberotha groehni gen. et sp. nov. (Neuroptera: Berothidae) is described from Baltic amber. The genus is assigned to Berothinae based on female terminalia that have the following characteristics: long hypocaudae on gonocoxite 9; ster- nite 7 that is medially divided into a pair of lateral sclerites; and gonocoxite 8 with a medial process that is very similar to that of some extant berothine genera. The new genus is the first described fossil genus of the subfamily Berothinae. It is noteworthy for the possession of a fully-developed CuP in the hind wing, a vein that is strongly reduced in all other species of the subfamily. The genus Spiroberotha is considered to belong to the Berothinae. Key words: Baltic amber, Eocene, Berothidae, Berothinae Introduction The Berothidae is a small family (approximately 130 extant species, including Rhachiberothinae) that occurs in most warm-temperate to tropical regions of the world, and very rarely in temperate regions (Oswald 2013).
  • 1 Universidade Federal Do Ceará Centro De Ciências

    1 Universidade Federal Do Ceará Centro De Ciências

    1 UNIVERSIDADE FEDERAL DO CEARÁ CENTRO DE CIÊNCIAS DEPARTAMENTO DE GEOLOGIA PROGRAMA DE PÓS-GRADUAÇÃO EM GEOLOGIA LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL FORTALEZA 2019 2 LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL Tese apresentada ao Programa de Pós- Graduação em Geologia da Universidade Federal do Ceará, como requisito parcial à obtenção do título de doutor em Geologia. Área de concentração: Geologia Sedimentar e Paleontologia. Orientador: Prof. Dr. Geraldo Jorge Barbosa de Moura. Coorientador: Prof. Dr. César Ulisses Vieira Veríssimo. FORTALEZA 2019 3 4 LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL Tese apresentada ao Programa de Pós- Graduação em Geologia da Universidade Federal do Ceará, como requisito parcial à obtenção do título de doutor em Geologia. Área de concentração: Geologia Sedimentar e Paleontologia. Aprovada em: 18/01/2019. BANCA EXAMINADORA ________________________________________ Prof. Dr. Geraldo Jorge Barbosa de Moura (Orientador) Universidade Federal Rural de Pernambuco (UFRPE) _________________________________________ Prof. Dr. Marcio Mendes Universidade Federal do Ceará (UFC) _________________________________________ Prof. Dr. Marcos Antônio Leite do Nascimento Universidade Federal do Rio Grande do Norte (UFRN) _________________________________________ Prof. Dr Kleberson de Oliveira Porpino Universidade do Estado do Rio Grande do Norte (UERN) ________________________________________ Dra Pâmela Moura Universidade Federal do Ceará (UFC) 5 A Deus.

  • Neuroptera, Berothidae) from Australia 1 Doi: 10.3897/Zookeys.64.403 RESEARCH ARTICLE Launched to Accelerate Biodiversity Research

    A peer-reviewed open-access journal ZooKeys 64: 1–8A (2010) new species of Stenobiella Tillyard (Neuroptera, Berothidae) from Australia 1 doi: 10.3897/zookeys.64.403 RESEARCH ARTICLE www.pensoftonline.net/zookeys Launched to accelerate biodiversity research A new species of Stenobiella Tillyard (Neuroptera, Berothidae) from Australia Shaun L. Winterton California State Collection of Arthropods, California Department of Food & Agriculture, 3294 Meadowview Rd. Sacramento, CA, USA 95832-1148 urn:lsid:zoobank.org:author:37F5AC48-EC3A-47ED-902B-2BD1467CCA72 Corresponding author: Shaun L. Winterton ( [email protected] ) Academic editor: A. Contreras-Ramos | Received 2 February 2010 | Accepted 25 August 2010 | Published 22 October 2010 urn:lsid:zoobank.org:pub:2DC68952-8896-4E71-A4FE-15A3D25D9D47 Citation: Winterton SL (2010) A new species of Stenobiella Tillyard (Neuroptera, Berothidae) from Australia. ZooKeys 64 : 1 – 8 . doi: 10.3897/zookeys.64.403 Abstract Stenobiella variola sp. n., a new species of beaded lacewing (Neuroptera: Berothidae), is described and fi gured from south-eastern Australia. A preliminary key to Stenobiella species is presented. Keywords Berothidae, Neuroptera, lacewing Introduction Beaded lacewings (Berothidae) are a small family of Neuroptera comprising approxi- mately 100 species occurring throughout most biogeographical regions. Members of the family are recognised by elongation of the pronotum, female usually with hypocau- dae and substantial cubital veins in both wings. Th e larvae are associated with termites, and undergo a degree of hypermetamorphosis during development (Brushwein, 1987). Four subfamilies of Berothidae are recognised: Rhachiberothinae, Cyrenoberothi- nae, Berothinae and Nosybinae (Aspöck, 1986; MacLeod and Adams, 1967; New, Copyright Shaun L. Winterton. This is an open access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
  • Юрские Сетчатокрылые (Insecta: Neuroptera) Центральной Азии

    Юрские Сетчатокрылые (Insecta: Neuroptera) Центральной Азии

    РОССИЙСКАЯ АКАДЕМИЯ НАУК ПАЛЕОНТОЛОГИЧЕСКИЙ ИНСТИТУТ им. А.А. Борисяка на правах рукописи Храмов Александр Валерьевич ЮРСКИЕ СЕТЧАТОКРЫЛЫЕ (INSECTA: NEUROPTERA) ЦЕНТРАЛЬНОЙ АЗИИ 25.00.02 Палеонтология и стратиграфия Диссертация на соискание ученой степени кандидата биологических наук Научный руководитель: доктор биологических наук Пономаренко Александр Георгиевич Москва - 2014 Оглавление ВВЕДЕНИЕ............................................................................................................................. стр. 4 Глава 1. История изучения юрских Neuroptera................................................................стр.7 Глава 2. Отряд Neuroptera..................................................................................................стр. 11 2.1. Система и биология современных Neuroptera....................................................... стр. 11 2.2. Строение крыльев и номенклатура жилкования Neuroptera............................. стр. 14 2.3. Палеонтологическая летопись Neuroptera.............................................................. стр. 17 Глава 3. Материалы и методы.......................................................................................... стр. 31 3.1. Коллекции юрских Neuroptera и их обработка...................................................... стр. 31 3.2. Описание местонахождений юрских Neuroptera Центральной Азии................ стр. 32 Глава 4. Обзор фаун юрских Neuroptera Центральной Азии..................................... стр. 42 4.1. Согюты (Киргизия).....................................................................................................
  • A Revision of the Family Rhachiberothidae Tjeder, 1959, Stat.N. {Neuroptera)

    A Revision of the Family Rhachiberothidae Tjeder, 1959, Stat.N. {Neuroptera)

    Systematic Entomology (1994) 19, 181-206 A revision of the family Rhachiberothidae Tjeder, 1959, stat.n. {Neuroptera) U . ASP 6 CK and M . W . MANSELL* Naturhistorisches Museum Wien, Vienna, Austria, and *Plant Protection Research Institute, Pretoria, South Africa Abstract. The subfamily Rhachiberothinae Tjeder, 1959, originally included in the Berothidae and recently transferred to the Mantispidae, is revised and elevated to family rank. A historical review and redescriptions of the family Rhachiberothidae stat.n. and of the genera Rhachiberotha Tjeder, 1959, and Mucroberotha Tjeder, 1959, are presented. Five new species, R.ingwe, R.sheilae, M.aethiopica, M.angolana and M.minteri, are described and differ­ entiated from the five hitherto-known species. The hypothesis of a sister-group relationship of the Rhachiberothidae to the Berothidae is re-established. The phylogenetic position of the Dilaridae as an adelphotaxon of the monophyletic group ([Rhachiberothidae + Berothidae] + Mantispidae) is discussed. Introduction This paper includes a historical review of the subject, a revision of the Rhachiberothidae with descriptions of five The Rhachiberothidae are an intriguing and enigmafic new species and a discussion of the phylogenetic relation­ group of insects whose phylogenetic relationships and ships of the Rhachiberothidae, Berothidae, Mantispidae systematic status have evoked liveJy debate since they and Dilaridae, and their relevant autapomorphies and were firstdescribed by Tjeder in 1959. Although originally synapomorphies. tentatively placed as a subfamily of the Berothidae, their The study is based on 170 specimens, including eight striking similarity to symphrasine Mantispidae raised holotypes, in the following collections: The Natural questions about their possible affinities with mantispids. History Museum, London, England (BMNH); Cali­ The identity and monophyly of both these families had fornia Academy of Sciences, San Francisco, California never been questioned before Tjeder's (1959) paper, (CASC); H.
  • NOTA / NOTE Echoes from the Past: Rediscovering Isoscelipteron Fulvum Costa, 1863 (Neuroptera: Berothidae) in Italy

    NOTA / NOTE Echoes from the Past: Rediscovering Isoscelipteron Fulvum Costa, 1863 (Neuroptera: Berothidae) in Italy

    ISSN: 1989-6581 Letardi (2017) www.aegaweb.com/arquivos_entomoloxicos ARQUIVOS ENTOMOLÓXICOS, 18: 279-282 NOTA / NOTE Echoes from the past: rediscovering Isoscelipteron fulvum Costa, 1863 (Neuroptera: Berothidae) in Italy. Agostino Letardi ENEA, C.R.Casaccia, STUDI IDR, Via Anguillarese 301, IT-00123, Roma, ITALY. e-mail: [email protected] Abstract: A male and a female of Isoscelipteron fulvum Costa, 1863 (Neuroptera: Berothidae) have been collected in the Catena Costiera mountains (Calabria, Cosenza province). This represents the rediscovery of that species in Italy after the description of the taxon made by Achille Costa in 1863. Aspects of the morphological variability of this species are discussed. Key words: Neuroptera, Berothidae, Isoscelipteron fulvum, morphological variability, faunistics, new record, Italy. Resumen: Ecos del pasado: redescubriendo Isoscelipteron fulvum Costa, 1863 (Neuroptera: Berothidae) en Italia. Se han capturado un macho y una hembra de Isoscelipteron fulvum Costa, 1863 (Neuroptera: Berothidae) en los montes de la Catena Costiera (Calabria, provincia de Cosenza). Esto representa el redescubrimiento de dicha especie en Italia después de la descripción llevada a cabo por Achille Costa en 1863. Se discuten algunas cuestiones relacionadas con la variabilidad morfológica de la especie. Palabras clave: Isoscelipteron fulvum Neuroptera, Berothidae, , variabilidad morfológica, faunística, nueva cita, Italia. Recibido: 23 de noviembre de 2017 Publicado on-line: 14 de diciembre de 2017 Aceptado: 1 de diciembre de 2017 Introduction In 1859 one of the keenest Italian naturalists, Achille Costa (1823-1898) collected in southern Calabria (in the Aspromonte mountain region) a specimen of a new genus of Neuroptera, described in his paper on entomological aspects of southern Calabria (Costa, 1863; see also Pantaleoni, 2012).
  • New Mantises (Insecta: Mantodea) in Cretaceous Ambers from Lebanon, Spain, and Myanmar

    New Mantises (Insecta: Mantodea) in Cretaceous Ambers from Lebanon, Spain, and Myanmar

    Cretaceous Research 60 (2016) 91e108 Contents lists available at ScienceDirect Cretaceous Research journal homepage: www.elsevier.com/locate/CretRes New mantises (Insecta: Mantodea) in Cretaceous ambers from Lebanon, Spain, and Myanmar * Xavier Delclos a, , Enrique Penalver~ b, Antonio Arillo c, Michael S. Engel d, AndreNel e, Dany Azar f, g, Andrew Ross h a Departament d'Estratigrafia, Paleontologia i Geociencies Marines, and Institut de Recerca de la Biodiversitat (IRBio), Facultat de Geologia, Universitat de Barcelona, Martí i Franques s/n, 08028 Barcelona, Spain b Museo Geominero, Instituto Geologico y Minero de Espana,~ Ríos Rosas 23, 28003 Madrid, Spain c Departamento de Zoología y Antropología Física, Facultad de Biología, Universidad Complutense, 28040 Madrid, Spain d Division of Entomology (Paleoentomology), Natural History Museum, and Department of Ecology & Evolutionary Biology, 1501 Crestline Drive e Suite 140, University of Kansas, Lawrence, Kansas 66045, USA e Institut de Systematique, Evolution, Biodiversite, ISYEB e UMR 7205 e CNRS, MNHN, UPMC, EPHE, Museum national d'Histoire naturelle, Sorbonne Universites, 57 rue Cuvier, CP 50, Entomologie, 75005 Paris, France f Lebanese University, Faculty of Sciences II, Department of Natural Sciences, Fanar, Matn P.O. Box 26110217, Lebanon g Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, PR China h Department of Natural Sciences, National Museums Scotland, Chambers St., Edinburgh EH1 1JF, UK article info abstract Article history: Diverse new material of mantises found in the Cretaceous amber-bearing deposits from Lebanon (Bar- Received 28 July 2015 remian), Spain (Albian), and Myanmar (AlbianeCenomanian) are described and figured. The Lebanese Received in revised form and Spanish forms are nymphs; while the one from Myanmar is an adult specimen.
  • Neuroptera, Hemerobiidae)

    Neuroptera, Hemerobiidae)

    Dtsch. Entomol. Z. 67 (2) 2020, 141–149 | DOI 10.3897/dez.67.56008 Anchored between heaven and earth – a new flightless brown lacewing from Peru (Neuroptera, Hemerobiidae) Ulrike Aspöck1,2, Horst Aspöck3, Axel Gruppe4 1 Natural History Museum, 2nd Zoological Department, Burgring 7, A-1010 Vienna, Austria 2 Department of Evolutionary Biology, University of Vienna, Althanstraße 14, A-1090 Vienna, Austria 3 Institute of Specific Prophylaxis and Tropical Medicine, Medical University Vienna (MUW), Kinderspitalgasse 15, A-1090 Vienna, Austria 4 Chair of Zoology – Entomological Division, Technical University of Munich, Hans-Carl-von-Carlowitz-Platz 2, D-85354 Freising, Germany http://zoobank.org/28F525EF-C030-4E04-9706-74C64A59225F Corresponding author: Axel Gruppe ([email protected]) Academic editor: Susanne Randolf ♦ Received 30 June 2020 ♦ Accepted 27 July 2020 ♦ Published 14 August 2020 Abstract Male and female of Nusalala peruana sp. nov., a flightless hemerobiid from the Andes mountain range of northern Peru, at a height of almost 4000 m, are described, figured and documented as the first record of a brachypterous, flightless species Nusalalaof Navás, 1913, from this country. The other two congeneric, brachypterous species are from high altitudes in Colombia and Costa Rica and have been described in the male sex only – the females remain unknown. The coriaceous domed forewings are shared by all three brachypterous Nusalala species. The ribbon-like hindwings of the male of N. peruana sp. nov. are unique, since those of the other brachypterous males are scale-like, as are the hindwings of the female of N. peruana sp. nov. Distribution and evolutionary back- grounds of brachyptery and flightlessness in Neuropterida are discussed.