DOCSLIB.ORG

A Revision of the Family Rhachiberothidae Tjeder, 1959, Stat.N. {Neuroptera)

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
A Revision of the Family Rhachiberothidae Tjeder, 1959, Stat.N. {Neuroptera) Systematic Entomology (1994) 19, 181-206 A revision of the family Rhachiberothidae Tjeder, 1959, stat.n. {Neuroptera) U . ASP 6 CK and M . W . MANSELL* Naturhistorisches Museum Wien, Vienna, Austria, and *Plant Protection Research Institute, Pretoria, South Africa Abstract. The subfamily Rhachiberothinae Tjeder, 1959, originally included in the Berothidae and recently transferred to the Mantispidae, is revised and elevated to family rank. A historical review and redescriptions of the family Rhachiberothidae stat.n. and of the genera Rhachiberotha Tjeder, 1959, and Mucroberotha Tjeder, 1959, are presented. Five new species, R.ingwe, R.sheilae, M.aethiopica, M.angolana and M.minteri, are described and differ­ entiated from the five hitherto-known species. The hypothesis of a sister-group relationship of the Rhachiberothidae to the Berothidae is re-established. The phylogenetic position of the Dilaridae as an adelphotaxon of the monophyletic group ([Rhachiberothidae + Berothidae] + Mantispidae) is discussed. Introduction This paper includes a historical review of the subject, a revision of the Rhachiberothidae with descriptions of five The Rhachiberothidae are an intriguing and enigmafic new species and a discussion of the phylogenetic relation­ group of insects whose phylogenetic relationships and ships of the Rhachiberothidae, Berothidae, Mantispidae systematic status have evoked liveJy debate since they and Dilaridae, and their relevant autapomorphies and were firstdescribed by Tjeder in 1959. Although originally synapomorphies. tentatively placed as a subfamily of the Berothidae, their The study is based on 170 specimens, including eight striking similarity to symphrasine Mantispidae raised holotypes, in the following collections: The Natural questions about their possible affinities with mantispids. History Museum, London, England (BMNH); Cali­ The identity and monophyly of both these families had fornia Academy of Sciences, San Francisco, California never been questioned before Tjeder's (1959) paper, (CASC); H. & U. Aspock Collection, Vienna, Austria despite the lack of definedautapomorphic characters which (HUAC); L. R. Minter Collection, Pietersburg, South the discovery of Rhachiberothinae demanded. Africa (LRMC); Naturhistorisches Museum Wien, Vienna, The present study was initiated to address these issues, Austria (NHMV); National Museum of Zimbabwe, and to document the new taxa and data which have accumu­ Bulawayo, Zimbabwe (NMBZ); Natal Museum, Pieter­ lated since Tjeder's (1959, 1968) studies. During the maritzburg, South Africa (NMSA); P. Ohm Collection, course of our investigations all available specimens of Kiel, Germany (POCG); R. B. Miller Collection, Project Rhachiberothinae, including the types, and data were City, California, U.S.A. (RBMC); National Collection of studied. Insects, Pretoria, South Africa (SANC, accession code However, before this work could be published, Willmann AcNE); Staatliches Museum filr Naturkunde, Stuttgart, (1990) transferred the Rhachiberothinae to the Mantis­ Germany (SMSG); National Museum of Natural History, pidae after studying only the holotype of Mucroberotha Smithsonian Institution, Washington, D.C. (USNM); fasciata Tjeder. His arguments have been carefully asses­ Zoological Institute, Lund University, Lund, Sweden sed, relative to the comprehensive material now available (ZILS); Institut und Zoologisches Museum, Hamburg, to us, and are discussed below. We still argue in favour of Germany (ZMUH). a sister-group relationship with the Berothidae, based upon Terminology and abbreviations are as follows: Al -A3, synapomorphies of larvae and adults which are more con­ anal veins; be, bursa copulatrix; C, costa; c, ninth gono­ vincing than the raptorial forelegs in terms of parsimony. coxite; cat, catalogue; charact, characterization; chor, We further propose that the Rhachiberothinae be elevated chorology; com, comment; CuA, anterior cubitus; CuP, to full family rank. posterior cubitus; cua-cup, crossvein; descr, description; Correspondence: Dr U. Aspi:ick, Zweite Zoologisch Abteilung, e, ectoproct; f, penisfilum;fe, femur; fig,figure; foss, fossil; Naturhistorisches Museum Wien, Burgring 7, A-1014 Wien, g, gonarcus; gl, gonapophysis lateralis; h, hypandrium Postfach 417, Austria. intemum; M, median; MA, anterior median; MP, posterior 181 .
Load more

Recommended publications
  • Head Anatomy of Adult Nevrorthus Apatelios and Basal Splitting Events in Neuroptera (Neuroptera: Nevrorthidae)
    72 (2): 111 – 136 27.7.2014 © Senckenberg Gesellschaft für Naturforschung, 2014. Head anatomy of adult Nevrorthus apatelios and basal splitting events in Neuroptera (Neuroptera: Nevrorthidae) Susanne Randolf *, 1, 2, Dominique Zimmermann 1, 2 & Ulrike Aspöck 1, 2 1 Natural History Museum Vienna, 2nd Zoological Department, Burgring 7, 1010 Vienna, Austria — 2 University of Vienna, Department of In- tegrative Zoology, Althanstrasse 14, 1090 Vienna, Austria; Susanne Randolf * [[email protected]]; Dominique Zimmermann [[email protected]]; Ulrike Aspöck [[email protected]] — * Corresponding author Accepted 22.v.2014. Published online at www.senckenberg.de/arthropod-systematics on 18.vii.2014. Abstract External and internal features of the head of adult Nevrorthus apatelios are described in detail. The results are compared with data from literature. The mouthpart muscle M. stipitalis transversalis and a hypopharyngeal transverse ligament are newly described for Neuroptera and herewith reported for the first time in Endopterygota. A submental gland with multiporous opening is described for Nevrorthidae and Osmylidae and is apparently unique among insects. The parsimony analysis indicates that Sisyridae is the sister group to all remaining Neuroptera. This placement is supported by the development of 1) a transverse division of the galea in two parts in all Neuroptera exclud­ ing Sisyridae, 2) the above mentioned submental gland in Nevrorthidae and Osmylidae, and 3) a poison system in all neuropteran larvae except Sisyridae. Implications for the phylogenetic relationships from the interpretation of larval character evolution, specifically the poison system, cryptonephry and formation of the head capsule are discussed. Key words Head anatomy, cladistic analysis, phylogeny, M.
    [Show full text]
  • UFRJ a Paleoentomofauna Brasileira
    Anuário do Instituto de Geociências - UFRJ www.anuario.igeo.ufrj.br A Paleoentomofauna Brasileira: Cenário Atual The Brazilian Fossil Insects: Current Scenario Dionizio Angelo de Moura-Júnior; Sandro Marcelo Scheler & Antonio Carlos Sequeira Fernandes Universidade Federal do Rio de Janeiro, Programa de Pós-Graduação em Geociências: Patrimônio Geopaleontológico, Museu Nacional, Quinta da Boa Vista s/nº, São Cristóvão, 20940-040. Rio de Janeiro, RJ, Brasil. E-mails: [email protected]; [email protected]; [email protected] Recebido em: 24/01/2018 Aprovado em: 08/03/2018 DOI: http://dx.doi.org/10.11137/2018_1_142_166 Resumo O presente trabalho fornece um panorama geral sobre o conhecimento da paleoentomologia brasileira até o presente, abordando insetos do Paleozoico, Mesozoico e Cenozoico, incluindo a atualização das espécies publicadas até o momento após a última grande revisão bibliográica, mencionando ainda as unidades geológicas em que ocorrem e os trabalhos relacionados. Palavras-chave: Paleoentomologia; insetos fósseis; Brasil Abstract This paper provides an overview of the Brazilian palaeoentomology, about insects Paleozoic, Mesozoic and Cenozoic, including the review of the published species at the present. It was analiyzed the geological units of occurrence and the related literature. Keywords: Palaeoentomology; fossil insects; Brazil Anuário do Instituto de Geociências - UFRJ 142 ISSN 0101-9759 e-ISSN 1982-3908 - Vol. 41 - 1 / 2018 p. 142-166 A Paleoentomofauna Brasileira: Cenário Atual Dionizio Angelo de Moura-Júnior; Sandro Marcelo Schefler & Antonio Carlos Sequeira Fernandes 1 Introdução Devoniano Superior (Engel & Grimaldi, 2004). Os insetos são um dos primeiros organismos Algumas ordens como Blattodea, Hemiptera, Odonata, Ephemeroptera e Psocopera surgiram a colonizar os ambientes terrestres e aquáticos no Carbonífero com ocorrências até o recente, continentais (Engel & Grimaldi, 2004).
    [Show full text]
  • André Nel Sixtieth Anniversary Festschrift
    Palaeoentomology 002 (6): 534–555 ISSN 2624-2826 (print edition) https://www.mapress.com/j/pe/ PALAEOENTOMOLOGY PE Copyright © 2019 Magnolia Press Editorial ISSN 2624-2834 (online edition) https://doi.org/10.11646/palaeoentomology.2.6.1 http://zoobank.org/urn:lsid:zoobank.org:pub:25D35BD3-0C86-4BD6-B350-C98CA499A9B4 André Nel sixtieth anniversary Festschrift DANY AZAR1, 2, ROMAIN GARROUSTE3 & ANTONIO ARILLO4 1Lebanese University, Faculty of Sciences II, Department of Natural Sciences, P.O. Box: 26110217, Fanar, Matn, Lebanon. Email: [email protected] 2State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China. 3Institut de Systématique, Évolution, Biodiversité, ISYEB-UMR 7205-CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France. 4Departamento de Biodiversidad, Ecología y Evolución, Facultad de Biología, Universidad Complutense, Madrid, Spain. FIGURE 1. Portrait of André Nel. During the last “International Congress on Fossil Insects, mainly by our esteemed Russian colleagues, and where Arthropods and Amber” held this year in the Dominican several of our members in the IPS contributed in edited volumes honoring some of our great scientists. Republic, we unanimously agreed—in the International This issue is a Festschrift to celebrate the 60th Palaeoentomological Society (IPS)—to honor our great birthday of Professor André Nel (from the ‘Muséum colleagues who have given us and the science (and still) national d’Histoire naturelle’, Paris) and constitutes significant knowledge on the evolution of fossil insects a tribute to him for his great ongoing, prolific and his and terrestrial arthropods over the years.
    [Show full text]
  • From Chewing to Sucking Via Phylogeny—From Sucking to Chewing Via Ontogeny: Mouthparts of Neuroptera
    Chapter 11 From Chewing to Sucking via Phylogeny—From Sucking to Chewing via Ontogeny: Mouthparts of Neuroptera Dominique Zimmermann, Susanne Randolf, and Ulrike Aspöck Abstract The Neuroptera are highly heterogeneous endopterygote insects. While their relatives Megaloptera and Raphidioptera have biting mouthparts also in their larval stage, the larvae of Neuroptera are characterized by conspicuous sucking jaws that are used to imbibe fluids, mostly the haemolymph of prey. They comprise a mandibular and a maxillary part and can be curved or straight, long or short. In the pupal stages, a transformation from the larval sucking to adult biting and chewing mouthparts takes place. The development during metamorphosis indicates that the larval maxillary stylet contains the Anlagen of different parts of the adult maxilla and that the larval mandibular stylet is a lateral outgrowth of the mandible. The mouth- parts of extant adult Neuroptera are of the biting and chewing functional type, whereas from the Mesozoic era forms with siphonate mouthparts are also known. Various food sources are used in larvae and in particular in adult Neuroptera. Morphological adaptations of the mouthparts of adult Neuroptera to the feeding on honeydew, pollen and arthropods are described in several examples. New hypoth- eses on the diet of adult Nevrorthidae and Dilaridae are presented. 11.1 Introduction The order Neuroptera, comprising about 5820 species (Oswald and Machado 2018), constitutes together with its sister group, the order Megaloptera (about 370 species), and their joint sister group Raphidioptera (about 250 species) the superorder Neuropterida. Neuroptera, formerly called Planipennia, are distributed worldwide and comprise 16 families of extremely heterogeneous insects.
    [Show full text]
  • A New Genus and Species of Thorny Lacewing from Upper Cretaceous Kuji Amber, Northeastern Japan (Neuroptera, Rhachiberothidae)
    A peer-reviewed open-access journal ZooKeys 802: 109–120 (2018) A new genus and species of thorny lacewing... 109 doi: 10.3897/zookeys.802.28754 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research A new genus and species of thorny lacewing from Upper Cretaceous Kuji amber, northeastern Japan (Neuroptera, Rhachiberothidae) Hiroshi Nakamine1, Shûhei Yamamoto2 1 Minoh Park Insect Museum, Minoh Park 1-18, Minoh City, Osaka, 562-0002, Japan 2 Integrative Research Center, Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA Corresponding author: Hiroshi Nakamine ([email protected]) Academic editor: S. Winterton | Received 31 July 2018 | Accepted 15 October 2018 | Published 4 December 2018 http://zoobank.org/407331A3-C2B3-4FDF-BA4D-D47ACDDA9BEF Citation: Nakamine H, Yamamoto S (2018) A new genus and species of thorny lacewing from Upper Cretaceous Kuji amber, northeastern Japan (Neuroptera, Rhachiberothidae). ZooKeys 802: 109–120. https://doi.org/10.3897/ zookeys.802.28754 Abstract Kujiberotha teruyukii gen. et sp. n., a remarkable new genus and species of Rhachiberothidae, is described from Upper Cretaceous amber from the Kuji area in northeastern Japan. This discovery represents the first record of this family both from Japan and from East Asia. This fossil taxon has the largest foreleg in the subfamily Paraberothinae found to date and its discovery implies that this group had higher morphologi- cal diversity in the Cretaceous than it does now. This finding also stresses the importance of the insect inclusions in Kuji amber, which have not been well explored in spite of their potential abundance.
    [Show full text]
  • X Revisiok of the Nearctic Hemerobiidae, Berothidae, Sisyridae, Polys'i'oechotidae and Dilaridae (Neuroptera)
    X REVISIOK OF THE NEARCTIC HEMEROBIIDAE, BEROTHIDAE, SISYRIDAE, POLYS'I'OECHOTIDAE AND DILARIDAE (NEUROPTERA) Received February 28, 1940 Presented March 13, 1940 The insects treated in this revision are among and Professor H. 13. Hungerford, University of the most typical of the Seuroptera (Planipennia). Kansas. I am under special obligation to Pro- Khen the family Hemerobiidae was established by fessor R. C. Smith, of Kansas State College, for Leach (1815), it included nearly all of the insects the loan of his extensive private collection of these now comprising the order. Since the beginning of insects, and for the opportunity of seeing many the present century, however, and subsequent to small collections sent to him for identification dur- the publication of Banks' "Revision of the Ne- ing the preparation of this revision. I am deeply ~irctic Hemerobiidae" (1905b), various genera grateful to llr. D. E. Kimmins of the British have been removed from the Hemerobiidae and lluseum of Satural History for placing at my placed in separate families. The groips thus disposal the type specimens in the Lfuseum col- formed (Berothidae, Sisyridae, Polystoechotidae, lection, and for making detailed comparisons with Dilnridae) have little in common with the re- Illclachlan's types, which were received at the stricted family Hemerobiidae; but in order that RIuseum after my visit there in 1938. As on the scope of the present revision be kept identical previous occasions I am indebted to Professor nith that of Banks', these families have also been Banks for many helpful suggestions and criticisms. included. The morphology of the Xeuroptera in general During the course of this revisional study, I and of most of the families occurring in the Ne- have examined somewhat more than eight thou- arctic region has been extensively treated by Kil- sand individuals of the families mentioned.
    [Show full text]
  • Insecta: Neuroptera) from Subsaharan Africa
    Ann. Naturhist. Mus. Wien 99 B 1 - 20 Wien, Dezember 1997 Studies on new and poorly-known Rhachiberothidae (Insecta: Neuroptera) from subsaharan Africa Abstract Three new species and one new genus of the family Rhachiberothidae are described and figured (wings, legs, d and/or p genitalia, partly colour photographs of living specimens). Rhachiberotha pulchra sp.n. was detected in the north of Namibia (W Grootfontein: d, p) and in Northern Transvaal (NW Potgietersrus: Q); apparently it represents the sister species of R. sheilae U. ASPOCK& MANSELL,1994. Mucroberotha copelandi sp.n. was found in the south of Kenya (Kajiado: p) and in Tanzania (Mkomazi Game Reserve: d, p); its position within the genus seems isolated. Hoelzeliella manselli gen.n. sp.n. was discovered in the Western Cape (Groot-Swartberge: p). The systematic position of Hoelzeliella within the Rhachiberothidae is uncertain; possibly the genus is more closely related to Mucroberotha TJEDER,1959, than to Rhachiberotha TJEDER, 1959, a better assessment may, however, be expected as soon as the d has been found. Further records of Mucroberotha vesicaria TJEDER,1959, in South Africa as well as Namibia are presented; the variability of the species (mainly on the basis of the vesicae) is discussed. The distribution of the family is mapped. The discovery of H. manselli has enlarged the known distribution considerably to the south. The distribution of the family Rhachiberothidae covers at least a large part of the Afrotropis, so far altogether thirteen species are known. Key words: Rhachiberothidae, Rhachiberotha pulchra, Mucroberotha vesicaria, Mucroberotha copelandi, Hoelzeliella manselli, new species, new genus, South Africa, Namibia, Tanzania, Kenya, Afrotropis, variability.
    [Show full text]
  • The Dustywings in Cretaceous Burmese Amber (Insecta: Neuroptera: Coniopterygidae)
    Journal of Systematic Palaeontology 2 (2): 133–136 Issued 23 July 2004 DOI: 10.1017/S1477201904001191 Printed in the United Kingdom C The Natural History Museum The Dustywings in Cretaceous Burmese amber (Insecta: Neuroptera: Coniopterygidae) Michael S. Engel Division of Entomology, Natural History Museum and Department of Ecology and Evolutionary Biology, Snow Hall, 1460 Jayhawk Boulevard, University of Kansas, Lawrence, KS 66045-7523, USA SYNOPSIS The dustywing fauna (Neuroptera: Coniopterygidae) of Upper Albian Burmese amber is revised. Two species are recognised, one belonging to the subfamily Aleuropteryginae and one to the Coniopteryginae. The aleuropterygine species is placed in the genus Glaesoconis (Glaesoconis baliopteryx sp. nov.), a previously known fontenelleine genus from New Jersey and Siberian ambers. The apparent coniopterygine differs in several features of wing venation and is therefore placed in its own tribe: Phthanoconini nov. (Phthanoconis burmitica gen. et sp. nov.). A revised key to Cretaceous dustywing genera is provided. KEY WORDS Aleuropteryginae, Coniopteryginae, Myanmar, Neuropterida, Planipennia, taxonomy Introduction arthropods such as mites living on conifers and deciduous trees or shrubs (Meinander 1972). The earliest fossil of the The Neuropterida (Neuroptera, Megaloptera and Raphidiop- family is Juraconiopteryx from the Upper Jurassic Karatau tera) are one of the most distinctive and ancient of endop- deposits in southern Kazakhstan (Meinander 1975) and, al- terygote lineages. Stem-group neuropterids occurred during though placed in Aleuropteryginae, little is preserved so that the Lower Permian with putative basal members of the or- assignment must be considered tentative. The earliest defin- ders Neuroptera and Megaloptera appearing shortly there- itive members of the family are those in Lower Cretaceous after and the earliest records of Raphidioptera coming from amber from Lebanon (Whalley 1980; Azar et al.
    [Show full text]
  • Changes to the Fossil Record of Insects Through Fifteen Years of Discovery
    This is a repository copy of Changes to the Fossil Record of Insects through Fifteen Years of Discovery. White Rose Research Online URL for this paper: https://eprints.whiterose.ac.uk/88391/ Version: Published Version Article: Nicholson, David Blair, Mayhew, Peter John orcid.org/0000-0002-7346-6560 and Ross, Andrew J (2015) Changes to the Fossil Record of Insects through Fifteen Years of Discovery. PLosOne. e0128554. https://doi.org/10.1371/journal.pone.0128554 Reuse Items deposited in White Rose Research Online are protected by copyright, with all rights reserved unless indicated otherwise. They may be downloaded and/or printed for private study, or other acts as permitted by national copyright laws. The publisher or other rights holders may allow further reproduction and re-use of the full text version. This is indicated by the licence information on the White Rose Research Online record for the item. Takedown If you consider content in White Rose Research Online to be in breach of UK law, please notify us by emailing [email protected] including the URL of the record and the reason for the withdrawal request. [email protected] https://eprints.whiterose.ac.uk/ RESEARCH ARTICLE Changes to the Fossil Record of Insects through Fifteen Years of Discovery David B. Nicholson1,2¤*, Peter J. Mayhew1, Andrew J. Ross2 1 Department of Biology, University of York, York, United Kingdom, 2 Department of Natural Sciences, National Museum of Scotland, Edinburgh, United Kingdom ¤ Current address: Department of Earth Sciences, The Natural History Museum, London, United Kingdom * [email protected] Abstract The first and last occurrences of hexapod families in the fossil record are compiled from publications up to end-2009.
    [Show full text]
  • (Neuroptera: Berothidae) from Baltic Amber
    Zootaxa 3946 (3): 401–415 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2015 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3946.3.7 http://zoobank.org/urn:lsid:zoobank.org:pub:237D932A-88EB-42D6-B3F3-C97C3BB9ECEF An important new fossil genus of Berothinae (Neuroptera: Berothidae) from Baltic amber VLADIMIR N. MAKARKIN1,3 & MICHAEL OHL2 1Institute of Biology and Soil Sciences, Far East Branch of the Russian Academy of Sciences, Vladivostok, 960022, Russia 2Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse, 43, D-10115 Berlin, Germany. E-mail: [email protected] 3Corresponding author. E-mail: [email protected] Abstract Elektroberotha groehni gen. et sp. nov. (Neuroptera: Berothidae) is described from Baltic amber. The genus is assigned to Berothinae based on female terminalia that have the following characteristics: long hypocaudae on gonocoxite 9; ster- nite 7 that is medially divided into a pair of lateral sclerites; and gonocoxite 8 with a medial process that is very similar to that of some extant berothine genera. The new genus is the first described fossil genus of the subfamily Berothinae. It is noteworthy for the possession of a fully-developed CuP in the hind wing, a vein that is strongly reduced in all other species of the subfamily. The genus Spiroberotha is considered to belong to the Berothinae. Key words: Baltic amber, Eocene, Berothidae, Berothinae Introduction The Berothidae is a small family (approximately 130 extant species, including Rhachiberothinae) that occurs in most warm-temperate to tropical regions of the world, and very rarely in temperate regions (Oswald 2013).
    [Show full text]
  • 1 Universidade Federal Do Ceará Centro De Ciências
    1 UNIVERSIDADE FEDERAL DO CEARÁ CENTRO DE CIÊNCIAS DEPARTAMENTO DE GEOLOGIA PROGRAMA DE PÓS-GRADUAÇÃO EM GEOLOGIA LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL FORTALEZA 2019 2 LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL Tese apresentada ao Programa de Pós- Graduação em Geologia da Universidade Federal do Ceará, como requisito parcial à obtenção do título de doutor em Geologia. Área de concentração: Geologia Sedimentar e Paleontologia. Orientador: Prof. Dr. Geraldo Jorge Barbosa de Moura. Coorientador: Prof. Dr. César Ulisses Vieira Veríssimo. FORTALEZA 2019 3 4 LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL Tese apresentada ao Programa de Pós- Graduação em Geologia da Universidade Federal do Ceará, como requisito parcial à obtenção do título de doutor em Geologia. Área de concentração: Geologia Sedimentar e Paleontologia. Aprovada em: 18/01/2019. BANCA EXAMINADORA ________________________________________ Prof. Dr. Geraldo Jorge Barbosa de Moura (Orientador) Universidade Federal Rural de Pernambuco (UFRPE) _________________________________________ Prof. Dr. Marcio Mendes Universidade Federal do Ceará (UFC) _________________________________________ Prof. Dr. Marcos Antônio Leite do Nascimento Universidade Federal do Rio Grande do Norte (UFRN) _________________________________________ Prof. Dr Kleberson de Oliveira Porpino Universidade do Estado do Rio Grande do Norte (UERN) ________________________________________ Dra Pâmela Moura Universidade Federal do Ceará (UFC) 5 A Deus.
    [Show full text]
  • Neuroptera, Berothidae) from Australia 1 Doi: 10.3897/Zookeys.64.403 RESEARCH ARTICLE Launched to Accelerate Biodiversity Research
    A peer-reviewed open-access journal ZooKeys 64: 1–8A (2010) new species of Stenobiella Tillyard (Neuroptera, Berothidae) from Australia 1 doi: 10.3897/zookeys.64.403 RESEARCH ARTICLE www.pensoftonline.net/zookeys Launched to accelerate biodiversity research A new species of Stenobiella Tillyard (Neuroptera, Berothidae) from Australia Shaun L. Winterton California State Collection of Arthropods, California Department of Food & Agriculture, 3294 Meadowview Rd. Sacramento, CA, USA 95832-1148 urn:lsid:zoobank.org:author:37F5AC48-EC3A-47ED-902B-2BD1467CCA72 Corresponding author: Shaun L. Winterton ( [email protected] ) Academic editor: A. Contreras-Ramos | Received 2 February 2010 | Accepted 25 August 2010 | Published 22 October 2010 urn:lsid:zoobank.org:pub:2DC68952-8896-4E71-A4FE-15A3D25D9D47 Citation: Winterton SL (2010) A new species of Stenobiella Tillyard (Neuroptera, Berothidae) from Australia. ZooKeys 64 : 1 – 8 . doi: 10.3897/zookeys.64.403 Abstract Stenobiella variola sp. n., a new species of beaded lacewing (Neuroptera: Berothidae), is described and fi gured from south-eastern Australia. A preliminary key to Stenobiella species is presented. Keywords Berothidae, Neuroptera, lacewing Introduction Beaded lacewings (Berothidae) are a small family of Neuroptera comprising approxi- mately 100 species occurring throughout most biogeographical regions. Members of the family are recognised by elongation of the pronotum, female usually with hypocau- dae and substantial cubital veins in both wings. Th e larvae are associated with termites, and undergo a degree of hypermetamorphosis during development (Brushwein, 1987). Four subfamilies of Berothidae are recognised: Rhachiberothinae, Cyrenoberothi- nae, Berothinae and Nosybinae (Aspöck, 1986; MacLeod and Adams, 1967; New, Copyright Shaun L. Winterton. This is an open access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
    [Show full text]