DOCSLIB.ORG

This Article Appeared in a Journal Published by Elsevier. the Attached Copy Is Furnished to the Author for Internal Non-Commerci

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
This Article Appeared in a Journal Published by Elsevier. the Attached Copy Is Furnished to the Author for Internal Non-Commerci This article appeared in a journal published by Elsevier. The attached copy is furnished to the author for internal non-commercial research and education use, including for instruction at the authors institution and sharing with colleagues. Other uses, including reproduction and distribution, or selling or licensing copies, or posting to personal, institutional or third party websites are prohibited. In most cases authors are permitted to post their version of the article (e.g. in Word or Tex form) to their personal website or institutional repository. Authors requiring further information regarding Elsevier’s archiving and manuscript policies are encouraged to visit: http://www.elsevier.com/copyright Author's personal copy Cretaceous Research 30 (2009) 1217–1222 Contents lists available at ScienceDirect Cretaceous Research journal homepage: www.elsevier.com/locate/CretRes Ascalochrysidae – a new lacewing family from the Mesozoic of China (Insecta: Neuroptera: Chrysopoidea) Dong Ren a, Vladimir N. Makarkin b,* a College of Life Science, Capital Normal University, Beijing 100048, China b Institute of Biology and Soil Sciences, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok 690022, Russian Federation article info abstract Article history: Ascalochrysa megaptera gen. et sp. nov. is described from the Yixian Formation (Lower Cretaceous) of Received 3 March 2009 China. It is treated as belonging to the new family Ascalochrysidae fam. nov., closely related to Meso- Accepted in revised form 19 June 2009 chrysopidae but easily distinguished from it in the hindwing venation features: e.g., numerous subcostal Available online 1 July 2009 crossveins; branches of Rs widely, irregularly spaced and deeply branched; crossvenation rich, irregular; convex vein-like fold (‘M5’) before concave CuA present. The presence of well-developed ‘M5’ in the hind Keywords: wing is interpreted to be plesiomorphic condition within the order. Ascalochrysidae are considered the Ascalochrysidae sister group of Mesochrysopidae (s.l.). Mesochrysopidae Cretaceous Ó 2009 Elsevier Ltd. All rights reserved. Yixian Formation China 1. Introduction 2. Materials and methods The taxonomic diversity of the Order Neuroptera was greatest in The specimen examined was collected near Chaomidian Village the Mesozoic. Several new neuropteran families have been in Liaoning Province, NE China from the deposits of the Yixian described from this era in the last decade, i.e., Makarkiniidae Formation. All photographs were taken with a Nikon Digital (Martins-Neto, 2000), Grammolingiidae (Ren, 2002), Limaiidae, Camera DXM1200C attached to a Leica MZ12.5 stereomicroscope. Tachinymphidae, Liassochrysidae (Nel et al., 2005), Aetheog- Drawing was done directly from the fossil with a camera lucida rammatidae (Ren and Engel, 2008a). Although at least two of these mounted on a Leica MZ12.5 stereomicroscope. are probably synonyms of other families (e.g., Makarkiniidae is We follow here the traditional (sensu Wootton, 2003) venational a probable synonym of Kalligrammatidae: Makarkin and Archibald, terminology of Comstock (1918) with the current interpretation of 2003; Liassochrysidae is a synonym of Mantispidae: Wedmann and Oswald (1993), Makarkin and Menon (2005), Archibald and Makarkin, 2007), the description of new Mesozoic families should Makarkin (2006) and Wedmann and Makarkin (2007). Terminology be expected in the future. of wing spaces follows Oswald (1993). In this paper we describe a new genus and species of a large Venational abbreviations are as follows: Sc, subcosta; R1, first neuropteran from the Lower Cretaceous of China. We interpret this branch of radius (R); Rs, radial sector; Rs1, most proximal branch of as a representative of the new family Ascalochrysidae fam. nov. Rs; M, media; MA, media anterior; MP, media posterior; CuP, poste- belonging to Chrysopoidea. This superfamily was recently estab- rior cubitus; CuA, anterior cubitus; 1A–3A, first to third anal veins. lished to include several families, mainly extinct (Nel et al., 2005). Institutional abbreviation. CNU, Capital Normal University, We discuss the systematic position of Ascalochrysidae fam. nov. Beijing, China. among chrysopoids, the family composition of this superfamily, and some important character states found in Ascalochrysa gen. 3. Locality and stratigraphy nov., e.g., the presence of a convex vein-like fold before CuA. Fossils occurring in the Yixian and Jiufotang Formations of northeastern China constitute the Jehol Biota, an assemblage which is thought to have been widely distributed in eastern Asia during * Corresponding author. the Early Cretaceous (Chen, 1988; Chen et al., 1998; Zhou et al., E-mail address: [email protected] (V.N. Makarkin). 2003). The Yixian Formation consists mainly of lacustrine 0195-6671/$ – see front matter Ó 2009 Elsevier Ltd. All rights reserved. doi:10.1016/j.cretres.2009.06.004 Author's personal copy 1218 D. Ren, V.N. Makarkin / Cretaceous Research 30 (2009) 1217–1222 sediments intercalated with volcaniclastics (Ren et al., 1995). It is crossveins throughout wing remigium numerous, irregular, not well known by a large number of exceptionally well-preserved forming gradate series or regular reticulation. fossils, including numerous insects, conchostracans, ostracods, gastropods, bivalves, fish, salamanders, turtles, lizards, feathered Comparison. The venational pattern of Ascalochrysidae fam. nov. dinosaurs, primitive birds (Confuciusornis Hou et al. and Liao- is most similar to that of Mesochrysopidae (s.l.) and strongly ningornis Hou) and angiosperm (Archaefructus Sun et al.) (Sun et al., different from that of all other families of the order. The new family 1998; Hou et al., 1999; Ding et al., 2001; Zhou et al., 2003). Some shares with Mesochrysopidae (s.l.) the following hind wing apo- Yixian Neuroptera have been described by Ren et al. (1995), Ren morphic character states: (1) the reduction of CuP to a part of and Guo (1996), Ren and Yin (2002), Ren (2003a), Nel et al. (2005) apparent basal crossvein between CuA and 1A; (2) the entire loss and Ren and Engel (2008a, b). Palaeobotanical data from paly- (or strong reduction) of 2A and 3A as a result of the reduction of nomorphs and macrofossils provide contradictory evidence on the cubito-anal area as a whole; (3) the loss or poor development of the climatic conditions in the Jehol times. On the one hand, the data humeral lobe and frenulum of coupling apparatus (the latter shared indicate a warm and humid Yixian climate (Ding et al., 2001), and with Chrysopidae). Other shared apomorphic character states are on the other, many plants suggest arid or semi-arid climatic the absence of trichosors, nygmata and the basal sinuous crossvein conditions (Zhou et al., 2003; Barrett and Hilton, 2006). Fu¨ rsich r-m, and the distal fusion of Sc and R1, but these features occur in et al. (2007) suggest that the climate was semi-arid with dry and other families of different lineages. Moreover, both families have wet seasons. distinctive structure of M and CuA. Ascalochrysidae fam. nov. are The age of the Yixian Formation was controversial for a long easily distinguished from Mesochrysopidae (s.l.) by the following time, with estimates varying from Late Jurassic to Early Cretaceous features [character states of Mesochrysopidae are given in (Sha, 2007a,b for summary). Now, an Early Cretaceous age of the brackets]: Sc þ R1 entering wing margin well beyond apex [at or Yixian Formation is considered to be well supported by radiometric (rarer) before wing apex]; subcostal crossveins numerous [only one dating; using different radioactive decay series, from 133.46 Æ 0.18 basal crossvein at most]; crossvenation over remigium irregular for the lowest beds (Chen et al., 2004), 126.1 Æ1.7 to 124.6 Æ 0.1 Ma [regular, forming gradate series or reticulation]; ‘M5’ present for the second Member containing insect fossils (Swisher et al., [absent]; M branched very close to the wing base [far to the wing 1999, 2002; Wang et al., 2001; Chen et al., 2004; He et al., 2006)to base]; Banksian folds absent [two of these folds present]. 121 Æ0.2 Ma for overlying lava layers and intrusive volcanics Superficially, the genus Ascalochrysa gen. nov. resembles the (Smith et al., 1995), i.e., ranging from the late Hauterivian to early genera of Ascalaphidae in general appearance, but they are easily Aptian (geologic time scale of Gradstein et al., 2005). Recently, an distinguished from the new genus in the absence of subcostal Early Cretaceous age of the Yixian Formation was supported by new crossveins and ‘M5’, the distal position of the origin of Rs, and the evidence (Li and Batten 2007; Yang et al., 2007; Zhu et al., 2007). configuration of M. Determination of the exact age of the Yixian Formation is very Comments on some character states.Scþ R1 entering wing important as it bears great evolutionary consequences, for example, margin beyond apex as found in Ascalochrysa gen. nov. is charac- the origin of angiosperms and the early radiation of birds (Barrett, teristic of the Myrmeleontoidea families (i.e., Myrmeleontidae, 2000). Some palaeontologists interpret this formation (treated as Ascalaphidae, Nymphidae, Palaeoleontidae, Nemopteridae, Babin- Early Cretaceous in age) as a refugium of Jurassic terrestrial relicts skaiidae) for which this character state is certainly a synapomor- in East Asia, based on the occurrences of compsognathid thero- phy. This condition is probably convergent in Ascalochrysidae fam. pod dinosaurs, ‘rhamphorhynchoid’ pterosaurs,
Load more

Recommended publications
  • The New Fossil Lacewings of Grammolingiidae (Neuroptera) from the Jurassic of Central Asia and Mongolia, with Notes on Biogeography of the Family
    Zootaxa 3478: 297–308 (2012) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA Copyright © 2012 · Magnolia Press Article ISSN 1175-5334 (online edition) urn:lsid:zoobank.org:pub:5EA14648-EF42-4CE2-9C68-CA0892C24E3E The new fossil lacewings of Grammolingiidae (Neuroptera) from the Jurassic of Central Asia and Mongolia, with notes on biogeography of the family ALEXANDER V. KHRAMOV Paleontological institute of the Russian Academy of Science, Profsouznaya str. 123, 117997, Moscow, Russia. [email protected] Abstract A new genus and three new species of the family Grammolingiidae are described: Protolingia mira gen. et sp. nov. and Litholingia longa sp. nov. from the Sai-Sagul locality (Kyrgyzstan, upper Lower Jurassic–lower Middle Jurassic) and Lep- tolingia oblonga sp. nov. from the Houtiyn-Hotgor locality (Mongolia, Upper Jurassic). Grammolingiidae are recorded from the Karatau locality (Kazakhstan, Upper Jurassic). This fossil lacewing family occurred in the South of Central Asia and in East Asia during the Middle and Upper Jurassic; its distribution was limited by Mongol-Okhotsk and Turgai seas. Grammolingiidae from Sai-Sagul is the oldest record of this family. They demonstrate unusual characteristics for the fam- ily, such as the pectinate CuP and the distal fusion of Sc and R1. Key words: Neuroptera, Grammolingiidae, Jurassic, Mongol-Okhotsk sea Introduction Grammolingiidae is an extinct Jurassic family of Neuroptera, which currently includes four genera and fourteen species. Thirteen species came from the Middle Jurassic Daohugou locality, China, Inner Mongolia (Ren 2002; Shi et al. 2011; Liu et al. 2011; Shi et al. 2012), and only one was found outside Daohugou, in the Upper Jurassic Shar- Teg locality, Mongolia (Khramov 2010).
    [Show full text]
  • UFRJ a Paleoentomofauna Brasileira
    Anuário do Instituto de Geociências - UFRJ www.anuario.igeo.ufrj.br A Paleoentomofauna Brasileira: Cenário Atual The Brazilian Fossil Insects: Current Scenario Dionizio Angelo de Moura-Júnior; Sandro Marcelo Scheler & Antonio Carlos Sequeira Fernandes Universidade Federal do Rio de Janeiro, Programa de Pós-Graduação em Geociências: Patrimônio Geopaleontológico, Museu Nacional, Quinta da Boa Vista s/nº, São Cristóvão, 20940-040. Rio de Janeiro, RJ, Brasil. E-mails: [email protected]; [email protected]; [email protected] Recebido em: 24/01/2018 Aprovado em: 08/03/2018 DOI: http://dx.doi.org/10.11137/2018_1_142_166 Resumo O presente trabalho fornece um panorama geral sobre o conhecimento da paleoentomologia brasileira até o presente, abordando insetos do Paleozoico, Mesozoico e Cenozoico, incluindo a atualização das espécies publicadas até o momento após a última grande revisão bibliográica, mencionando ainda as unidades geológicas em que ocorrem e os trabalhos relacionados. Palavras-chave: Paleoentomologia; insetos fósseis; Brasil Abstract This paper provides an overview of the Brazilian palaeoentomology, about insects Paleozoic, Mesozoic and Cenozoic, including the review of the published species at the present. It was analiyzed the geological units of occurrence and the related literature. Keywords: Palaeoentomology; fossil insects; Brazil Anuário do Instituto de Geociências - UFRJ 142 ISSN 0101-9759 e-ISSN 1982-3908 - Vol. 41 - 1 / 2018 p. 142-166 A Paleoentomofauna Brasileira: Cenário Atual Dionizio Angelo de Moura-Júnior; Sandro Marcelo Schefler & Antonio Carlos Sequeira Fernandes 1 Introdução Devoniano Superior (Engel & Grimaldi, 2004). Os insetos são um dos primeiros organismos Algumas ordens como Blattodea, Hemiptera, Odonata, Ephemeroptera e Psocopera surgiram a colonizar os ambientes terrestres e aquáticos no Carbonífero com ocorrências até o recente, continentais (Engel & Grimaldi, 2004).
    [Show full text]
  • From Chewing to Sucking Via Phylogeny—From Sucking to Chewing Via Ontogeny: Mouthparts of Neuroptera
    Chapter 11 From Chewing to Sucking via Phylogeny—From Sucking to Chewing via Ontogeny: Mouthparts of Neuroptera Dominique Zimmermann, Susanne Randolf, and Ulrike Aspöck Abstract The Neuroptera are highly heterogeneous endopterygote insects. While their relatives Megaloptera and Raphidioptera have biting mouthparts also in their larval stage, the larvae of Neuroptera are characterized by conspicuous sucking jaws that are used to imbibe fluids, mostly the haemolymph of prey. They comprise a mandibular and a maxillary part and can be curved or straight, long or short. In the pupal stages, a transformation from the larval sucking to adult biting and chewing mouthparts takes place. The development during metamorphosis indicates that the larval maxillary stylet contains the Anlagen of different parts of the adult maxilla and that the larval mandibular stylet is a lateral outgrowth of the mandible. The mouth- parts of extant adult Neuroptera are of the biting and chewing functional type, whereas from the Mesozoic era forms with siphonate mouthparts are also known. Various food sources are used in larvae and in particular in adult Neuroptera. Morphological adaptations of the mouthparts of adult Neuroptera to the feeding on honeydew, pollen and arthropods are described in several examples. New hypoth- eses on the diet of adult Nevrorthidae and Dilaridae are presented. 11.1 Introduction The order Neuroptera, comprising about 5820 species (Oswald and Machado 2018), constitutes together with its sister group, the order Megaloptera (about 370 species), and their joint sister group Raphidioptera (about 250 species) the superorder Neuropterida. Neuroptera, formerly called Planipennia, are distributed worldwide and comprise 16 families of extremely heterogeneous insects.
    [Show full text]
  • A New Genus and Species of Thorny Lacewing from Upper Cretaceous Kuji Amber, Northeastern Japan (Neuroptera, Rhachiberothidae)
    A peer-reviewed open-access journal ZooKeys 802: 109–120 (2018) A new genus and species of thorny lacewing... 109 doi: 10.3897/zookeys.802.28754 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research A new genus and species of thorny lacewing from Upper Cretaceous Kuji amber, northeastern Japan (Neuroptera, Rhachiberothidae) Hiroshi Nakamine1, Shûhei Yamamoto2 1 Minoh Park Insect Museum, Minoh Park 1-18, Minoh City, Osaka, 562-0002, Japan 2 Integrative Research Center, Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA Corresponding author: Hiroshi Nakamine ([email protected]) Academic editor: S. Winterton | Received 31 July 2018 | Accepted 15 October 2018 | Published 4 December 2018 http://zoobank.org/407331A3-C2B3-4FDF-BA4D-D47ACDDA9BEF Citation: Nakamine H, Yamamoto S (2018) A new genus and species of thorny lacewing from Upper Cretaceous Kuji amber, northeastern Japan (Neuroptera, Rhachiberothidae). ZooKeys 802: 109–120. https://doi.org/10.3897/ zookeys.802.28754 Abstract Kujiberotha teruyukii gen. et sp. n., a remarkable new genus and species of Rhachiberothidae, is described from Upper Cretaceous amber from the Kuji area in northeastern Japan. This discovery represents the first record of this family both from Japan and from East Asia. This fossil taxon has the largest foreleg in the subfamily Paraberothinae found to date and its discovery implies that this group had higher morphologi- cal diversity in the Cretaceous than it does now. This finding also stresses the importance of the insect inclusions in Kuji amber, which have not been well explored in spite of their potential abundance.
    [Show full text]
  • X Revisiok of the Nearctic Hemerobiidae, Berothidae, Sisyridae, Polys'i'oechotidae and Dilaridae (Neuroptera)
    X REVISIOK OF THE NEARCTIC HEMEROBIIDAE, BEROTHIDAE, SISYRIDAE, POLYS'I'OECHOTIDAE AND DILARIDAE (NEUROPTERA) Received February 28, 1940 Presented March 13, 1940 The insects treated in this revision are among and Professor H. 13. Hungerford, University of the most typical of the Seuroptera (Planipennia). Kansas. I am under special obligation to Pro- Khen the family Hemerobiidae was established by fessor R. C. Smith, of Kansas State College, for Leach (1815), it included nearly all of the insects the loan of his extensive private collection of these now comprising the order. Since the beginning of insects, and for the opportunity of seeing many the present century, however, and subsequent to small collections sent to him for identification dur- the publication of Banks' "Revision of the Ne- ing the preparation of this revision. I am deeply ~irctic Hemerobiidae" (1905b), various genera grateful to llr. D. E. Kimmins of the British have been removed from the Hemerobiidae and lluseum of Satural History for placing at my placed in separate families. The groips thus disposal the type specimens in the Lfuseum col- formed (Berothidae, Sisyridae, Polystoechotidae, lection, and for making detailed comparisons with Dilnridae) have little in common with the re- Illclachlan's types, which were received at the stricted family Hemerobiidae; but in order that RIuseum after my visit there in 1938. As on the scope of the present revision be kept identical previous occasions I am indebted to Professor nith that of Banks', these families have also been Banks for many helpful suggestions and criticisms. included. The morphology of the Xeuroptera in general During the course of this revisional study, I and of most of the families occurring in the Ne- have examined somewhat more than eight thou- arctic region has been extensively treated by Kil- sand individuals of the families mentioned.
    [Show full text]
  • Describing Undiscovered Insect Diversity: an Introduction to Collected Papers Describing 150 New Taxa
    Zootaxa 3478: 7–10 (2012) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA Copyright © 2012 · Magnolia Press Editorial ISSN 1175-5334 (online edition) urn:lsid:zoobank.org:pub:BE8BE638-A9E4-415D-8ADA-493BBCCE55B7 Describing undiscovered insect diversity: an introduction to collected papers describing 150 new taxa ZHI-QIANG ZHANG New Zealand Arthropod Collection, Landcare Research, 231 Morrin Road, St. Johns, Auckland 1072, New Zealand; E-mail: [email protected] The number of species on earth is estimated to be 8.7 million, with 87% of them to be discovered and described (Mora et al. 2011). The total of 8.7 million seems to be an underestimate, because for Animalia alone, over 1.5 million species have been described (Zhang 2011b). The most successful group, the Insecta, accounts for almost two-thirds of all animals. Zootaxa has been a major force in facilitating the descriptions of undiscovered animal species in the world—publishing about 20% of all new species indexed in Zoological Record each year (Zhang 2011a). In the first week since the publication of the ICZN amendment allowing e-only publication (ICZN 2012), Zootaxa has published 484 new taxa. In this special volume highlighting undiscovered global diversity of insects, Zootaxa publishes 150 new taxa (143 species + 7 genera) of 12 insect orders in 44 papers by authors from different parts of the world (Table 1). TABLE 1. Numbers of new taxa (species-group and genus-group) by insect order (listed following sequence in Zhang 2011c) with references. Insect order Number of new taxa References Species-group Genus-group Ephemeroptera 1 0 Flowers (2012) Orthoptera 2 2 Bolfarini et al.
    [Show full text]
  • (Neuroptera: Berothidae) from Baltic Amber
    Zootaxa 3946 (3): 401–415 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2015 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3946.3.7 http://zoobank.org/urn:lsid:zoobank.org:pub:237D932A-88EB-42D6-B3F3-C97C3BB9ECEF An important new fossil genus of Berothinae (Neuroptera: Berothidae) from Baltic amber VLADIMIR N. MAKARKIN1,3 & MICHAEL OHL2 1Institute of Biology and Soil Sciences, Far East Branch of the Russian Academy of Sciences, Vladivostok, 960022, Russia 2Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse, 43, D-10115 Berlin, Germany. E-mail: [email protected] 3Corresponding author. E-mail: [email protected] Abstract Elektroberotha groehni gen. et sp. nov. (Neuroptera: Berothidae) is described from Baltic amber. The genus is assigned to Berothinae based on female terminalia that have the following characteristics: long hypocaudae on gonocoxite 9; ster- nite 7 that is medially divided into a pair of lateral sclerites; and gonocoxite 8 with a medial process that is very similar to that of some extant berothine genera. The new genus is the first described fossil genus of the subfamily Berothinae. It is noteworthy for the possession of a fully-developed CuP in the hind wing, a vein that is strongly reduced in all other species of the subfamily. The genus Spiroberotha is considered to belong to the Berothinae. Key words: Baltic amber, Eocene, Berothidae, Berothinae Introduction The Berothidae is a small family (approximately 130 extant species, including Rhachiberothinae) that occurs in most warm-temperate to tropical regions of the world, and very rarely in temperate regions (Oswald 2013).
    [Show full text]
  • 1 Universidade Federal Do Ceará Centro De Ciências
    1 UNIVERSIDADE FEDERAL DO CEARÁ CENTRO DE CIÊNCIAS DEPARTAMENTO DE GEOLOGIA PROGRAMA DE PÓS-GRADUAÇÃO EM GEOLOGIA LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL FORTALEZA 2019 2 LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL Tese apresentada ao Programa de Pós- Graduação em Geologia da Universidade Federal do Ceará, como requisito parcial à obtenção do título de doutor em Geologia. Área de concentração: Geologia Sedimentar e Paleontologia. Orientador: Prof. Dr. Geraldo Jorge Barbosa de Moura. Coorientador: Prof. Dr. César Ulisses Vieira Veríssimo. FORTALEZA 2019 3 4 LUÍS CARLOS BASTOS FREITAS DESCRIÇÃO DE NOVOS TAXONS DE INSETOS FÓSSEIS DOS MEMBROS CRATO E ROMUALDO DA FORMAÇÃO SANTANA E COMENTÁRIOS SOBRE A GEODIVERSIDADE DO GEOPARK ARARIPE, BACIA SEDIMENTAR DO ARARIPE, NORDESTE DO BRASIL Tese apresentada ao Programa de Pós- Graduação em Geologia da Universidade Federal do Ceará, como requisito parcial à obtenção do título de doutor em Geologia. Área de concentração: Geologia Sedimentar e Paleontologia. Aprovada em: 18/01/2019. BANCA EXAMINADORA ________________________________________ Prof. Dr. Geraldo Jorge Barbosa de Moura (Orientador) Universidade Federal Rural de Pernambuco (UFRPE) _________________________________________ Prof. Dr. Marcio Mendes Universidade Federal do Ceará (UFC) _________________________________________ Prof. Dr. Marcos Antônio Leite do Nascimento Universidade Federal do Rio Grande do Norte (UFRN) _________________________________________ Prof. Dr Kleberson de Oliveira Porpino Universidade do Estado do Rio Grande do Norte (UERN) ________________________________________ Dra Pâmela Moura Universidade Federal do Ceará (UFC) 5 A Deus.
    [Show full text]
  • From the Middle Jurassic of Inner Mongolia, China
    Zootaxa 2897: 51–56 (2011) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2011 · Magnolia Press ISSN 1175-5334 (online edition) A new lacewing (Insecta: Neuroptera: Grammolingiidae) from the Middle Jurassic of Inner Mongolia, China YUSHUANG LIU1, 2, CHAOFAN SHI 2& DONG REN2 1Paleontological Institute, Shenyang Normal University, Shenyang 110034, China. E-mail: [email protected] 2College of Life Sciences, Capital Normal University, 105 Xisanhuanbeilu, Haidian District; Beijing 100048; China. E-mail: [email protected] Abstract A new species of the family Grammolingiidae (Neuroptera) (Leptolingia imminuta sp. nov.) is described from Daohugou village (Middle Jurassic), Inner Mongolia, China. In this new species, MA forks at the same level as the separation of Rs2 from Rs, close to the middle of forewing, this structure of MA is peculiar in Grammolingiidae and is different from that of all other known species. Moreover, this new species is the smallest species known in the family Grammolingiidae (30 mm wing span). Key words: Leptolingia, Jiulongshan Formation, new species, Daohugou Introduction Grammolingiidae Ren, 2002 is a small family of Jurassic Neuroptera found in Central and East Asia. Three genera and nine species were described previously in this family. Among them, eight species were found in the Middle Jurassic deposits of Daohugou Village, Inner Mongolia, China (Ren 2002, Shi et al. 2011); only one species (Lep- tolingia shartegica Khramov, 2010) has been found in the Late Jurassic deposits of Sharteg, Mongolia (Khramov 2010). Leptolingia was erected by Ren (2002) who described two species; Shi et al. (2011) revised this genus, iden- tifying definitive generic characters and added a new species, L.
    [Show full text]
  • Neuroptera, Berothidae) from Australia 1 Doi: 10.3897/Zookeys.64.403 RESEARCH ARTICLE Launched to Accelerate Biodiversity Research
    A peer-reviewed open-access journal ZooKeys 64: 1–8A (2010) new species of Stenobiella Tillyard (Neuroptera, Berothidae) from Australia 1 doi: 10.3897/zookeys.64.403 RESEARCH ARTICLE www.pensoftonline.net/zookeys Launched to accelerate biodiversity research A new species of Stenobiella Tillyard (Neuroptera, Berothidae) from Australia Shaun L. Winterton California State Collection of Arthropods, California Department of Food & Agriculture, 3294 Meadowview Rd. Sacramento, CA, USA 95832-1148 urn:lsid:zoobank.org:author:37F5AC48-EC3A-47ED-902B-2BD1467CCA72 Corresponding author: Shaun L. Winterton ( [email protected] ) Academic editor: A. Contreras-Ramos | Received 2 February 2010 | Accepted 25 August 2010 | Published 22 October 2010 urn:lsid:zoobank.org:pub:2DC68952-8896-4E71-A4FE-15A3D25D9D47 Citation: Winterton SL (2010) A new species of Stenobiella Tillyard (Neuroptera, Berothidae) from Australia. ZooKeys 64 : 1 – 8 . doi: 10.3897/zookeys.64.403 Abstract Stenobiella variola sp. n., a new species of beaded lacewing (Neuroptera: Berothidae), is described and fi gured from south-eastern Australia. A preliminary key to Stenobiella species is presented. Keywords Berothidae, Neuroptera, lacewing Introduction Beaded lacewings (Berothidae) are a small family of Neuroptera comprising approxi- mately 100 species occurring throughout most biogeographical regions. Members of the family are recognised by elongation of the pronotum, female usually with hypocau- dae and substantial cubital veins in both wings. Th e larvae are associated with termites, and undergo a degree of hypermetamorphosis during development (Brushwein, 1987). Four subfamilies of Berothidae are recognised: Rhachiberothinae, Cyrenoberothi- nae, Berothinae and Nosybinae (Aspöck, 1986; MacLeod and Adams, 1967; New, Copyright Shaun L. Winterton. This is an open access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
    [Show full text]
  • 00011694.Pdf
    6th International Congress on Fossil Insects, Arthropods and Amber Byblos, April 2013 ----------------------------------------------------------------------------------------------------------------------------------- Current State of Knowledge of the Mesozoic Neuroptera of China Qiang YANG1, Yongjie WANG1, Chaofan SHI1, Yuanyuan PENG1, Vladimir N. MAKARKIN1,2, Dong REN1 1- College of Life Sciences, Capital Normal University, Beijing, 100048, China. E-mail: [email protected] 2- Institute of Biology and Soil Sciences, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia. E-mail: [email protected] The order Neuroptera is known in the fossil record since the Early Permian (about 280 to 260 Ma) and it thrives ever since. The neuropterans occur relatively rare in the pre-Mesozoic localities. The order was most diverse in the Mesozoic. Most of the extinct and extant groups have been present at this time. Many families, genera and species of the Chinese neuropteran fossils have been published since the order was first reported by Ping Chi in 1928. Up to now, 20 families have been recorded from the Mesozoic of China, and 58 genera and 97 species have been described from this era. We mainly study the fossils materials from two localities of northeast China: the Middle Jurassic Daohugou (Jiulongshan Formation) in Inner Mongolia, and the Early Eocene Huangbanjigou (Yixian Formation) in Liaoning Province. The Daohugou beds contain a diverse insect fauna, with 19 insect orders being represented. We have examined approximately 4000 Neuroptera specimens which are housed in the Capital Normal University. Hitherto, only 58 species (29 genera) belonging to 11 families were described. Osmylidae are most abundant and diverse among neuropterans in the assemblage.
    [Show full text]
  • Юрские Сетчатокрылые (Insecta: Neuroptera) Центральной Азии
    РОССИЙСКАЯ АКАДЕМИЯ НАУК ПАЛЕОНТОЛОГИЧЕСКИЙ ИНСТИТУТ им. А.А. Борисяка на правах рукописи Храмов Александр Валерьевич ЮРСКИЕ СЕТЧАТОКРЫЛЫЕ (INSECTA: NEUROPTERA) ЦЕНТРАЛЬНОЙ АЗИИ 25.00.02 Палеонтология и стратиграфия Диссертация на соискание ученой степени кандидата биологических наук Научный руководитель: доктор биологических наук Пономаренко Александр Георгиевич Москва - 2014 Оглавление ВВЕДЕНИЕ............................................................................................................................. стр. 4 Глава 1. История изучения юрских Neuroptera................................................................стр.7 Глава 2. Отряд Neuroptera..................................................................................................стр. 11 2.1. Система и биология современных Neuroptera....................................................... стр. 11 2.2. Строение крыльев и номенклатура жилкования Neuroptera............................. стр. 14 2.3. Палеонтологическая летопись Neuroptera.............................................................. стр. 17 Глава 3. Материалы и методы.......................................................................................... стр. 31 3.1. Коллекции юрских Neuroptera и их обработка...................................................... стр. 31 3.2. Описание местонахождений юрских Neuroptera Центральной Азии................ стр. 32 Глава 4. Обзор фаун юрских Neuroptera Центральной Азии..................................... стр. 42 4.1. Согюты (Киргизия).....................................................................................................
    [Show full text]