The Palm Heart As a New Commercial Crop from Tropical America MICHAELJ

Total Page:16

File Type:pdf, Size:1020Kb

The Palm Heart As a New Commercial Crop from Tropical America MICHAELJ PRINCIPES BALICK: PALM HEART The Palm Heart as a New Commercial Crop from Tropical America MICHAELJ. BAI.ICK 15 Clermonl Road, Wilmington, Delnwnre 19803 A stately species of the genus Euterpe seeded fruits (Fig. 1). Unfortunately, is found throughout the once vast rain these palms contain a sweet tender heart, forests of Costa Rica. 'I'his palm, when and Iwfo~.t,they reach a seed-l~~.odncing mature, reaches the upper limits of the size they are often relle(l to accluire this forest canopy, some 70 feet or more in relatively small amount of edible "hqart the air. Its slender trunk, while only six of palm." In fact, in areas of forest to eight inches thick, supports a splendid much traveled, as soon as these palms crown of fronds, directly under which reach one inch in diameter, they become are produced clusters of small, single- desirable in the eyes of the local inhabi- -2. Bactris gasipues planted close together for harvesting. 1, Two sources of heart of palm, Euterpe to the left, Bnclris gasipaes at tl~eright. PRINCIPES BALICK: PALM HEART 4. The heart removed. More must be peeled to get to the part used for food. tants as.a source of food. The campe- San Vito de Java in the mountains of sinos (rural peasant farmers) decapitate western Costa Rica, is attempting the even these small palms to get at the experimental cropping of a substitute slender heart, which at this stage pro- palm heart, the pejibaye or peach palm vides hardly more than a mouthful of (Racrrk gnsil~nc?~).Several rows of this footl. sl)(:(:ic!s Wc!l.c: l)lilll~c(lill ilIl ilLlc1lll)L lo With the increasing gopulatiori stress clemoristrate thal this palm coulcl serve on Costa Ricn's dwindlin~rain forests. 11s it domesticated slil~stit~itefor tht! tl~ereis tloul,t as Lo ~l~is~)LI~III'S i~lrilily /:'ulc!r.l~c!,ll~c:l.el)y ~.ctluci~~~ 1111: ~rl~clitio~~ul to survive these depredations, as each damage to the forest ecology. These new seedling, upon discovery, is usually palms become large enough to cut for soon destroyed. Palm hearts are con- their hearts in two and one-half to foul. tinually collected for commercial can- years from seed, depending on the cli- ning and for export, thereby continuing mate and location, and they yield up to to reduce further the already low popula- three pounds of edible heart, substan- tions. tially more than the Euterpe palm (Figs. Where does the solution lie? Should 2-4). One additional commercial ad- conservationists press for a prohibition vantage to using the pejibaye palm hear! on the importation of this "delicacy," is that it will not discolor after heinp or is there another ar~swer? (:II~,21s (loc!s ~hcwild pro(111c~.The IIUIII- Lns Cruces 'l'ropical Uo~aliicalCi~rJelr ily of the heart is ex cell en^, having a and Field Station, a unit of the Organi- crisp, nutty flavor. The peach palm is zation for l'rol)ical Stuclies, Inc. rrcal. wc:ll ~IIOWIIill Costa Hica as nri iml)or- DI<ANSFIELD: TERMINAT, FI.OWERING tant food, the fruits being appetizing and given is needed for optimum yield. The nutritious. The trunk contains a sweet peach palm will produce up to 12 off- Terminal Flowering in Daemonorop! sugarcanelike material which has a po- shoots, usually averaging five. These tential for use in the manfacture of can be used for propagation or left to JOHN DRAMFIELD alcoholic beverages. Cattle enjoy graz- mature into a second crou or ratoon. Royul Botanic Gardens, Kew, Richnond, Surrey, Englnrarl ing on the leaves, disregarding the spines One objection sometimes raised to har- in their gusto. vesting this palm is its spiny trunk, For cropping, the plants can be spaced which can be a challenge to the handler. Terminal (hapaxanthic) flowering is hapaxanthic flowering. This is quite close to each other in order to give However, spineless strains of this palm well known in the lepidocaryoid palms; Ijlorrorops calicarpn (Griff. ) Mart. maximum yield. At Las Cruces the are being propagated, which should re- all species of Korthalsia, Plectoconzin, acaulescent forest undergrowth 1 trees were spaced .75 by 2.0 meters, move this difficultv. Plectocomiopsis, Myrialepis, Ancistro- highly characteristic of forest horde while in large-scale plantings at the Tur- As experimental cropping and the first phyllum, Oncocalamus, Eleiodoxa, and rivulets in nonswampy lowland and rialba Tropical Agricultural Research few commercial operations have proved Eugeissorm, and apparently all species dipterocarp forest in the Malay Pt and Training Center, 2,200 trees were worthwhile. what is now needed is a of Raphia, have this particular method sula; in such a habitat it forms thicl planted per hectare (2.47 acres), and large-scale growing program promoted of flowering- in which axes develop often in association with Calanzus spaced 3 by 1.5 meters (Activities at by the government and with the coopera- through a vegetative phase and then pass taneus Griff. It also grows in N Turridba l(3) : 4, April-June 1973, Tur- tion of commercial packaging com- into a flowering phase which terminates Sumatra in a small area of forest in rialba, Costa Rica). Conclusions drawn panies. Heart of palm has the potential the -growth of those axes. In Metroxrlon Langkat Nature Reserve which is from the Las Cruces experiment indicate for being an important agricultural all species but one flower hapaxanthi- markable for the large number of pl that a little further spacing than was crop in tropical America. cally; M. amicarum (H. Wendl.) Becc., of Malayan distribution apparently the exception, has pleonanthic flowering, found elsewhere in Sumatra (e.g. ( the inflorescences being produced in the mus castaneus, C. insignis Griff., axils of the leaves until the axis dies of Johannesteijs~nann~innltifrons (Rei carry a hopeful message to those of us injury or old age. That the two ef- f. et Zoll.) H. E. Moore). who have watched an otherwise fine fectively and markedly different methods Dacrnonoro])~~nlir:~r~)a protl ucc:s : specimen palm apparently succumb to of flowering can occur in the same genus podially suckering axes which shock or disease when transplanted. The has stimulated much speculation as to scarcely more than 30 cm. tall and Plroclnix sylvc~risillr~strnted \V;IS I>lnn~ed ~VIIi(,l~ rlow(*ri~~~ l)ro(~*ss is 1 II,~III~Iivv ;III~ I:II.~(! I(::IV(!S (I~I(!II 3 5 111. I;III wi111 I, 111l.ee years ago it1 prime co~~tlilional which advanced. Holttum (1955), Cor- brown sheaths having conspic1 the Polynesian Hotel, Walt Disney ner (1966), Hall6 and Oldeman (1970), oblique combs of pale brown sp World. After a period of some months, and Dransfield (1970) favour the idea The pinnae are regular, fine, and cler signs of trouble were perceived in the that hapaxanthic flowering is basic crowded, and sometimes the leaf 1 older leaves. They had gradually de- whereas Moore (1969), at least in in a short vestigial cirrus. A clump teriorated into the forlorn terminal at Metroxylon, and Moore and Uhl (1973), contain six or more axes of varying a right angle to the trunk. A new bud, regard it as derived. Tomlinson and An axis of D. calicarpa in flower h to our surprise, had in the meantime Moore (1968) regard the whole argu- marvellous appearance; a large k pushed out alongside the original and ment concerning which is primitive, kerry of flowers is found almos gives promise of ultimately assuming a hapaxanthy or pleonanthy, as being re- ground level among the leaf bases. normal aspect. dundant. Like many evolutionary. prob-. flowering axis consists of very crov If nothing else, the message may be lems there are good arguments difficult internodes with short leaves endin, not to be too hasty in consigning Lo the to prove in favour of both views. What- vestigal cirri and undulate, partiall) ~rave~ardthe apparent transldan t casu- ever the real answer mav. be.. the situa- veloped leaflets. These leaves hec alty. tion is certainly an interesting one and smaller ant1 smaller distally. In the it may be of importance to record the of them are the inflorescences n- MORGANEVANS occurrence of hapaxanthy in a genus likewise decrease in size distally I The freak behavior of the palm de- Director, Landscape Architecture hitherto thought to be entirely pleo- the most distal ones are minute ant1 picted in the accompanying photograph WED Enterprises nanthic. tain no flowers. In all, about 25 would not make the subject a candidate 1.4.01 Flower Street Of the 115 species of Daen~onorops, florescences are produced from for a beauty prize. It does, however, Glendale, California 91201 at least one species appears to have axis, not apparently reaching ant1 .
Recommended publications
  • ANATOMICAL PROPERTIES of NINE INDIGENOUS RATTAN SPECIES of JAMBI, INDONESIA Krisdianto*, Jasni and Tutiana Forest Products Research and Development Center, Jl
    Indonesian Journal of Forestry Research Vol. 5, No. 2, October 2018, 147-161 ISSN: 2355-7079/E-ISSN: 2406-8195 ANATOMICAL PROPERTIES OF NINE INDIGENOUS RATTAN SPECIES OF JAMBI, INDONESIA Krisdianto*, -asni and Tutiana Forest Products Research and Development Center, Jl. Gunung Batu 5, Bogor, :est -ava 16610, Indonesia Received: 16 June 2017, Revised: 31 October 2018, Accepted: 31 October 2018 ANATOMICAL 35OPERTIES OF NINE INDIGENOUS RATTAN SPECIES OF -AMBI, INDONESIA. 9arious rattan species grow naturally in -ambi, Indonesia, i.e. opon (Plectocomiopsis geminiflora (Griff.) Beccari), udang (Korthalsia flagelaris Miquel), getah (Daemonorops micracantha (Griff.) Beccari), duduk (D. didymophylla Beccari), tunggal (Calamus laevigatus Martius), sijau (C. tumidus Furtado), buruk ati (C. insignis Griff. var. longispinosus Dransfield), batu (C. zonatus Beccari), and paku (C. exillis Griff.). The rattan species are classified as lesser known species, which its properties are unknown to rattan supplier and consumers. This paper observes the anatomical properties of nine indigeneous rattan species of -ambi. Anatomical observations were conducted from solid, sectioned and macerated samples. Results show that anatomical properties become a diagnostic characteristic for rattan species identification and specific characteristic has been developed for Ney species determination. 9ascular bundles in the outer part of the stem of opon and udang rattans are yellow-capped. Width and length ratio of vascular bundle in the outer part is more than 1, oval shape was found in sijau rattan, while elongated shape vascular bundle with the ratio less than 1 was found in buruk ati. Fiber bundles separated from vessels are found in central ground parencymatous tissue of rattan tunggal. In the peripheral area, fiber bundle forms one or two lines with no specific pattern found in rattan paku, while fiber bundles in one line with alternate pattern found in rattan duduk.
    [Show full text]
  • Rattan Field Guide Change Style-Edit Last New:Layout 1.Qxd
    Contents Page Foreword Acknowledgement 1- Introduction . .1 2- How to use this book . 1 3- Rattan in Cambodia . .1 4- Use . .2 5- Rattan ecology and habitat . 2 6- Rattan characters . 3 6.1 Habit . 4 6.2 Stem/can . .4 6.3 Leaf Sheath . .4 6.4 Leave and leaflet . 6 6.5 Climbing organ . .8 6.6 Inflorescence . .9 6.7 Flower . .10 6.8 Fruit . .11 7- Specimen collection . .12 7.1 Collection method . 12 7.2 Field record . .13 7.3 Maintenance and drying . 13 8- Local names . .14 9- Key Identification to rattan genera . 17 9.1 Calamus L. .18 9.2 Daemonorops Bl. 44 9.3 Korthalsia Bl. 48 9.4 Myrialepis Becc. 52 9.5 Plectocomia Mart. ex Bl. 56 9.6 Plectocomiopsis Becc. 62 Table: Species list of Cambodia Rattan and a summary of abundance and distribution . .15 Glossary . 66 Reference . 67 List of rattan species . .68 Specimen references . .68 FOREWORD Rattan counts as one of the most important non-timber forest products that contribute to livelihoods as source of incomes and food and also to national economy with handicraft and furniture industry. In Cambodia, 18 species have been recorded so far and most of them are daily used by local communities and supplying the rattan industry. Meanwhile, with rattan resources decreasing due to over-harvesting and loss of forest ecosystem there is an urgent need to stop this trend and find ways to conserve this biodiversity that play an important economic role for the country. This manual is one step towards sustainable rattan management as it allows to show/display the diversity of rattan and its contribution.
    [Show full text]
  • Investigating the Significance of Rattan Spines from a Small
    Sains Malaysiana 43(7)(2014): 973–976 A View from a Different Angle: Investigating the Significance of Rattan Spines from a Small Mammals’ Visual Point of View Using ImageJ (Pandangan dari Sudut Lain: Menyelidik Signifikan Duri Rotan dari Sudut Pandangan Visual Mamalia Kecil Menggunakan ImageJ) NIK FADZLY*, ASYRAF MANSOR, RAHMAD ZAKARIA & SYED AHMAD EDZHAM ABSTRACT Rattans are one of the most unique and economically important plants for most tropical countries. There is however, a lack of interest in the specific study of the rattan spines. In this paper, we tested a new hypothesis concerning the functional role of rattan spines. We proposed that rattan spines also serve as a visual deterrent against herbivores or seed predators. In our proposed method we used an Imaging software, ImageJ, to measure the spine area of four species of rattan (Calamus insignis, Myrialepis schortechinii, Plectocomiopsis geminiflorus and Calamus caesius) from two different orientations (root to shoot and vice versa). Our results showed that rattan spines were very heterogeneous and highly variable between different species. One common trait that the rattan spines share is that spine area measurements of shoot to root (ShR) are larger than root to shoot (RH) orientation. We propose that the downwards spine angle might be specifically designed to discourage climbing leaf and seed predators. Keywords: Anti-herbivory; ImageJ; rattan; seed predators; small mammals; spines ABSTRAK Rotan adalah tumbuhan yang mempunyai keunikan dan kepentingan daripada segi ekonomi bagi kebanyakan negara tropika. Namun begitu, kurang perhatian diberikan pada kajian duri rotan. Dalam kertas ini, kami menguji hipotesis baru mengenai fungsi peranan duri rotan.
    [Show full text]
  • Pinanga Lepidota (Arecaceae: Arecoideae), a New Record for the Philippines from Palawan Island
    PRIMARY RESEARCH PAPER | Philippine Journal of Systematic Biology DOI 10.26757/pjsb2020c14005 Pinanga lepidota (Arecaceae: Arecoideae), a new record for the Philippines from Palawan Island Edwino S. Fernando1,4,5, Eugene L.R. Logatoc2, Pastor L. Malabrigo Jr.1,4, and Jiro T. Adorador3 Abstract Pinanga lepidota (Arecaceae), previously known only from Borneo, is reported here as a new record for the Philippines from Palawan Island. A key to the identification of similar species of Pinanga in the Philippines is provided, including brief notes on Bornean Arecaceae elements in Palawan. Keywords: Mt Mantalingahan, Palmae, palms, Pinanga Introduction New Guinea (Govaerts et al. 2020). In the Philippines, 20 species were earlier listed by Beccari (1919) and Merrill (1922); Pinanga Blume includes acaulescent or erect, diminutive six species have since been added to this list (Fernando 1988, or robust forest undergrowth palms that occur from sea level up 1994, Adorador et al. 2020). to ca. 2800 m elevation (Dransfield et al. 2008). The genus Our continuing studies on the palms of the Philippine name is the Latinized form of the Malay vernacular name Islands have revealed the presence of Pinanga lepidota Rendle pinang, often applied to the betel nut palm, Areca catechu L., on the lower slopes of Mt Mantalingahan near the southern end and various other species of the genera Areca L., Pinanga, and of Palawan Island, approximately 220 km from Sabah on the Nenga H.Wendl. & Drude (Dransfield et al. 2008). Pinanga northeastern tip of Borneo. There is just one other species of occurs in tropical and subtropical Asia to the northwest Pacific, Pinanga, P.
    [Show full text]
  • 2 HISTORICAL ROLE of PALMS in HUMAN CULTURE Ancient and Traditional Palm Products
    Tropical Palms 13 2 HISTORICAL ROLE OF PALMS IN HUMAN CULTURE Pre-industrial indigenous people of the past as well as of the present have an intimate and direct relationship with the renewable natural resources of their environment. Prior to the Industrial Age, wild and cultivated plants and wild and domesticated animals provided all of the food and most of the material needs of particular groups of people. Looking back to those past times it is apparent that a few plant families played a prominent role as a source of edible and nonedible raw materials. For the entire world, three plant families stand out in terms of their past and present utility to humankind: the grass family (Gramineae), the legume family (Leguminosae) and the palm family (Palmae). If the geographic focus is narrowed to the tropical regions, the importance of the palm family is obvious. The following discussion sets out to provide an overview of the economic importance of palms in earlier times. No single comprehensive study has yet been made of the historical role of palms in human culture, making this effort more difficult. A considerable amount of information on the subject is scattered in the anthropological and sociological literature as part of ethnographic treatments of culture groups throughout the tropics. Moreover, historical uses of products from individual palm species can be found in studies of major economic species such as the coconut or date palms. It should also be noted that in addition to being highly utilitarian, palms have a pivotal role in myth and ritual in certain cultures.
    [Show full text]
  • Systematics and Evolution of the Rattan Genus Korthalsia Bl
    SYSTEMATICS AND EVOLUTION OF THE RATTAN GENUS KORTHALSIA BL. (ARECACEAE) WITH SPECIAL REFERENCE TO DOMATIA A thesis submitted by Salwa Shahimi For the Degree of Doctor of Philosophy School of Biological Sciences University of Reading February 2018 i Declaration I can confirm that is my own work and the use of all material from other sources have been properly and fully acknowledged. Salwa Shahimi Reading, February 2018 ii ABSTRACT Korthalsia is a genus of palms endemic to Malesian region and known for the several species that have close associations with ants. In this study, 101 new sequences were generated to add 18 Korthalsia species from Malaysia, Singapore, Myanmar and Vietnam to an existing but unpublished data set for calamoid palms. Three nuclear (prk, rpb2, and ITS) and three chloroplast (rps16, trnD-trnT and ndhF) markers were sampled and Bayesian Inference and Maximum Likelihood methods of tree reconstruction used. The new phylogeny of the calamoids was largely congruent with the published studies, though the taxon sampling was more thorough. Each of the three tribes of the Calamoideae appeared to be monophyletic. The Eugeissoneae was consistently resolved as sister to Calameae and Lepidocaryeae, and better resolved, better supported topologies below the tribal level were identified. Korthalsia is monophyletic, and novel hypotheses of species level relationships in Korthalsia were put forward. These hypotheses of species level relationships in Korthalsia served as a framework for the better understanding of the evolution of ocrea. The morphological and developmental study of ocrea in genus Korthalsia included detailed study using Light and Scanning Electron Microscopy for seven samples of 28 species of Korthalsia, in order to provide understanding of ocrea morphological traits.
    [Show full text]
  • Patterns of Distribution of Malesian Vascular Plants
    Malesian plant distributions 243 Patterns of distribution of Malesian vascular plants W J Baker1, M J E Coode, J Dransfield, S Dransfield, M M Harley, P Hoffmann and R J Johns The Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey, TW9 3AE, UK 1Department of Botany, Plant Science Laboratories, University of Reading, Whiteknights, Reading, Berkshire, RG6 6AS, UK Key words: biogeography, phytogeography, palynology, SE Asia, Malesia, Palmae, Gramineae, Euphorbiaceae, Elaeocarpaceae, Antidesma, Elaeocarpus, Nypa, Spinizonocolpites Abstract analytical phase Biogeographical work con- cerned with the analytical phase has appeared A miscellaneous selection of Malesian plant distributions is increasingly in the systematic literature and it is presented, including examples from the Palmae, here that modern methods are most evident Gramineae, Euphorbiaceae, Elaeocarpaceae, and various fern genera Hypotheses of the tectonic evolution of the Previously, most classifications have been based area may be required to explain many of the observed pat- on intuition and overall similarity which, though terns that are described Two major distribution types are they may stand the test of time, are nevertheless identified repeatedly, the first displaying a strongly Sundaic subjective Despite the introduction of statistical bias and the second focusing on E Malesia Patterns involv- techniques which aimed to make similarity- ing New Guinea are complex as they tend to include a vari- able combination of other islands such as Sulawesi, Maluku, based or phenetic
    [Show full text]
  • Palmtraits 1.0, a Species-Level Functional Trait Database of Palms Worldwide
    www.nature.com/scientificdata OPEN PalmTraits 1.0, a species-level Data Descriptor functional trait database of palms worldwide Received: 3 June 2019 W. Daniel Kissling 1, Henrik Balslev2, William J. Baker 3, John Dransfeld3, Bastian Göldel2, Accepted: 9 August 2019 Jun Ying Lim1, Renske E. Onstein4 & Jens-Christian Svenning2,5 Published: xx xx xxxx Plant traits are critical to plant form and function —including growth, survival and reproduction— and therefore shape fundamental aspects of population and ecosystem dynamics as well as ecosystem services. Here, we present a global species-level compilation of key functional traits for palms (Arecaceae), a plant family with keystone importance in tropical and subtropical ecosystems. We derived measurements of essential functional traits for all (>2500) palm species from key sources such as monographs, books, other scientifc publications, as well as herbarium collections. This includes traits related to growth form, stems, armature, leaves and fruits. Although many species are still lacking trait information, the standardized and global coverage of the data set will be important for supporting future studies in tropical ecology, rainforest evolution, paleoecology, biogeography, macroecology, macroevolution, global change biology and conservation. Potential uses are comparative eco- evolutionary studies, ecological research on community dynamics, plant-animal interactions and ecosystem functioning, studies on plant-based ecosystem services, as well as conservation science concerned with the loss and restoration of functional diversity in a changing world. Background & Summary Most ecosystems are composed of a large number of species with diferent characteristics. Tese characteristics (i.e. traits) refect morphological, reproductive, physiological, phenological, or behavioural measurements of spe- cies that are usually collected to study intraspecifc trait variation (i.e.
    [Show full text]
  • Plant Resources of South-East Asia Is a Multivolume Handbook That Aims
    Plant Resources of South-East Asia is a multivolume handbook that aims to summarize knowledge about useful plants for workers in education, re­ search, extension and industry. The following institutions are responsible for the coordination ofth e Prosea Programme and the Handbook: - Forest Research Institute of Malaysia (FRIM), Karung Berkunci 201, Jalan FRI Kepong, 52109 Kuala Lumpur, Malaysia - Indonesian Institute of Sciences (LIPI), Widya Graha, Jalan Gatot Subroto 10, Jakarta 12710, Indonesia - Institute of Ecology and Biological Resources (IEBR), Nghia Do, Tu Liem, Hanoi, Vietnam - Papua New Guinea University of Technology (UNITECH), Private Mail Bag, Lae, Papua New Guinea - Philippine Council for Agriculture, Forestry and Natural Resources Re­ search &Developmen t (PCARRD), Los Banos, Laguna, the Philippines - Thailand Institute of Scientific and Technological Research (TISTR), 196 Phahonyothin Road, Bang Khen, Bangkok 10900, Thailand - Wageningen Agricultural University (WAU), Costerweg 50, 6701 BH Wage- ningen, the Netherlands In addition to the financial support of the above-mentioned coordinating insti­ tutes, this book has been made possible through the general financial support to Prosea of: - the Finnish International Development Agency (FINNIDA) - the Netherlands Ministry ofAgriculture , Nature Management and Fisheries - the Netherlands Ministry of Foreign Affairs, Directorate-General for Inter­ national Cooperation (DGIS) - 'Yayasan Sarana Wanajaya', Ministry of Forestry, Indonesia This work was carried out with the aid of a specific grant from : - the International Development Research Centre (IDRC), Ottawa, Canada 3z/s}$i Plant Resources ofSouth-Eas t Asia No6 Rattans J. Dransfield and N. Manokaran (Editors) Droevendaalsesteeg 3a Postbus 241 6700 AE Wageningen T r Pudoc Scientific Publishers, Wageningen 1993 VW\ ~) f Vr Y DR JOHN DRANSFIELD is a tropical botanist who gained his first degree at the University of Cambridge.
    [Show full text]
  • Stem Anatomy of Climbing Palms in Relation to Long-Distance Water Transport P
    Aliso: A Journal of Systematic and Evolutionary Botany Volume 22 | Issue 1 Article 22 2006 Stem Anatomy of Climbing Palms in Relation to Long-distance Water Transport P. Barry Tomlinson Harvard University; National Tropical Botanical Garden Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons Recommended Citation Tomlinson, P. Barry (2006) "Stem Anatomy of Climbing Palms in Relation to Long-distance Water Transport," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 22: Iss. 1, Article 22. Available at: http://scholarship.claremont.edu/aliso/vol22/iss1/22 Aliso 22, pp. 265-277 © 2006, Rancho Santa Ana Botanic Garden STEM ANATOMY OF CLIMBING PALMS IN RELATION TO LONG-DISTANCE WATER TRANSPORT P. BARRY TOMLINSON Harvard Forest, Harvard University, Petersham, Massachusetts 01366, USA and National Tropical Botanical Garden, 3530 Papalina Road, Kalaheo, Hawaii 96741, USA ([email protected]) ABSTRACT Palms lack secondary growth so their primary vascular system is long-lived and must be minimally vulnerable to dysfunction. For water movement, the axial xylem must be well defended against cav­ itation. Climbing palms can be very long and represent a maximum solution to transport problems. How is this demonstrated in their anatomy? This article contrasts stem vascular anatomy in a cane­ like "tree palm" (Rhapis excelsa) with that in the American climbing palm Desmoncus and the Old World rattan genus Calamus. Rhapis, representing the basic classical palm vasculature, has a contin­ uously integrated vascular system determined by branching of the axial (stem) system to produce leaf traces, bridges, and continuing axial bundles. Axial transport is favored over appendicular structures because leaves are irrigated solely by narrower protoxylem tracheids.
    [Show full text]
  • Projek Etnobotani Kinabalu
    PEOPLE AND PLANTS WORKING9 PAPER - FEBRUARY 2002 This series of working papers is intended to provide information and to generate fruitful discussion Projek Etnobotani on key issues Kinabalu in the sustainable The making of a Dusun Ethnoflora and equitable use (Sabah, Malaysia) of plant resources. G.J. Martin, A. Lee Agama, J.H. Beaman and J. Nais Please send comments on this paper and suggestions for future issues to People and Plants Initiative, Division of Ecological Sciences, UNESCO, 7 Place de Fontenoy, 75352 Paris CEDEX 07 SP, France. The designations employed and the presentation of material throughout this publication do not imply the expression of any opinion whatsoever on the part of UNESCO concerning the legal status of any country, territory, city, or area of its authorities, or concerning the delimitation of its frontiers or boundaries. The opinions expressed in this paper are entirely those of the authors and do not commit any Organization. Authors’ addresses: Gary J. Martin John H. Beaman The Global Diversity Foundation The Herbarium B.P. 262 Marrakesh-Medina Royal Botanic Gardens Kew Morocco Richmond, Surrey TW9 3AB <[email protected]> United Kingdom <[email protected]> Agnes Lee Agama WWF Malaysia Jamili Nais P.O. Box 14393 Sabah Parks 88850 Kota Kinabalu P.O. Box 10626 Sabah 88806 Kota Kinabalu Malaysia Malaysia <[email protected]> <[email protected]> Photos: Gary Martin, Michael Doolittle, Robert Höft Published in 2002 by the United Nations Educational, Scientific and Cultural Organization 7, place de Fontenoy, 75352 Paris Cedex 07 SP, FRANCE Printed by UNESCO on chlorine-free recycled paper Edited by Martin Walters Design: Ivette Fabbri Layout: Martina Höft © UNESCO / G.
    [Show full text]
  • Insects on Palms
    Insects on Palms i Insects on Palms F.W. Howard, D. Moore, R.M. Giblin-Davis and R.G. Abad CABI Publishing CABI Publishing is a division of CAB International CABI Publishing CABI Publishing CAB International 10 E 40th Street Wallingford Suite 3203 Oxon OX10 8DE New York, NY 10016 UK USA Tel: +44 (0)1491 832111 Tel: +1 (212) 481 7018 Fax: +44 (0)1491 833508 Fax: +1 (212) 686 7993 Email: [email protected] Email: [email protected] Web site: www.cabi.org © CAB International 2001. All rights reserved. No part of this publication may be repro- duced in any form or by any means, electronically, mechanically, by photocopying, recording or otherwise, without the prior permission of the copyright owners. A catalogue record for this book is available from the British Library, London, UK. Library of Congress Cataloging-in-Publication Data Insects on palms / by Forrest W. Howard … [et al.]. p. cm. Includes bibliographical references and index. ISBN 0-85199-326-5 (alk. paper) 1. Palms--Diseases and pests. 2. Insect pests. 3. Insect pests--Control. I. Howard, F. W. SB608.P22 I57 2001 634.9’74--dc21 00-057965 ISBN 0 85199 326 5 Typeset by Columns Design Ltd, Reading Printed and bound in the UK by Biddles Ltd, Guildford and King’s Lynn Contents List of Boxes vii Authors and Contributors viii Acknowledgements x Preface xiii 1 The Animal Class Insecta and the Plant Family Palmae 1 Forrest W. Howard 2 Defoliators of Palms 33 Lepidoptera 34 Forrest W. Howard and Reynaldo G. Abad Coleoptera 81 Forrest W.
    [Show full text]