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Contributions to the Study of the Genus Eriastrum: II

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Contributions to the Study of the Genus Eriastrum: II Brigham Young University Science Bulletin, Biological Series Volume 16 Number 4 Article 1 9-1972 Contributions to the study of the genus Eriastrum: II. Notes concerning the type specimens and descriptions of the species H. Keith Harrison Weber State College, Ogden, Utah Follow this and additional works at: https://scholarsarchive.byu.edu/byuscib Part of the Anatomy Commons, Botany Commons, Physiology Commons, and the Zoology Commons Recommended Citation Harrison, H. Keith (1972) "Contributions to the study of the genus Eriastrum: II. Notes concerning the type specimens and descriptions of the species," Brigham Young University Science Bulletin, Biological Series: Vol. 16 : No. 4 , Article 1. Available at: https://scholarsarchive.byu.edu/byuscib/vol16/iss4/1 This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Brigham Young University Science Bulletin, Biological Series by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. S-M''^(JOOq) MUs. '• Co MP. ZOOL. Brigham Young University Science Bulletin ''ffi 21973 CONTRIBUTIONS TO THE STUDY OF THE GENUS ERIASTRUM II. Notes concerning the type specimens and descriptions of the species by H. Keith Harrison BIOLOGICAL SERIES — VOLUME 16, NUMBER 4 SEPTEMBER 1972 . BRIGHAM YOUNG UNIVERSITY SCIENCE BULLETIN BIOLOGICAL SERIES Editor: Stanley L. Welsli, Department of Botany, Brigham Young University, Provo, Utah Members of the Editorial Board: Vernon J. Tipton, Zoology Ferron L. Anderson, Zoology Joseph R. Murdock, Botany Wilnier W. Tanner, Zoology Ex officio Members: A. Lester Allen, Dean, College of Biological and Agricultural Sciences Ernest L. Olson, Chairman. University Publications The Brigham Young University Science Bulletin, Biological Series, publishes acceptable papers, particularly large manuscripts, on all phases of biology. Separate numbers and back volumes can be purchased from Publication Sales, Brigham Young University, Provo, Utah. All remittances should be made payable to Brigham Young UniversitN. Orders and materials for library exchange should be directed to the Division of Gifts and Exchange, Brigham Young University Library, Provo. Utah 84501 Brigham Young University Science Bulletin CONTRIBUTIONS TO THE STUDY OF THE GENUS ERIASTRUM II. Notes concerning the type specimens and descriptions of the species by H. Keith Harrison BIOLOGICAL SERIES — VOLUME 16, NUMBER 4 SEPTEMBER 1972 TABLE OF CONTENTS INTRODUCTION TAXONOMY ACKNOWLEDGMENTS 25 LITERATURE CITED 26 CONTRIBUTIONS TO THE STUDY OF THE GENUS ERIASTRUM II. Notes concerning the type specimens and descriptions of the species by H. Keith Harrison^ ABSTRACT The species of the genus Eriastnim are cussions with regard to the type specimens or described. Keys to the species and subspecies lectotypes and interspecific relationships are and the geographic ranges are provided. Dis- presented. INTRODUCTION The species of the genus Eriastnim are close- 1945), and as a later work (Harrison 1959) is ly related; and the range of variation found in not generally available, it seems desirable to one species in branching, the lobing of the include the information as to the type speci- leaves, vesture, etc., overlaps that found in other mens and synonomy for all the species, together species to a considerable degree. The species as with the additions and discussions by the author. thev are now generally known were described bv It is understood that the first elements to be Mason ( 1945 ) . At that time, it was noted that found of a new species may represent either the the proportions and size of the corolla and norm or one extreme in the variation of the stamens were important in identification of the species. At a later time, other plants may be plants. Further study (Harrison 1959, 1968) collected which cause the investigator to create introduced new methods, emphasized the im- a name for a subspecies, thereby segregating portance of the relative size and proportions of certain specimens as being closely related to the corolla and androecium, and elucidated the (but still distinct from) the type upon which nature of these characteristics. the species is based. The fonnation of a sub- During the investigation of the species of specific category for the new element then ne- Eriastnim (Harrison 1959), it was found that cessitates, according to the International Code the type specimen for one of the species had of Botanical Nomenclature (Article 25) and not been published and that other problems logic, the establishment of another subspecies to concerning types and lectotypes require some include only those plants most representative of discussion. Inasmuch as nearly three decades the type—one could otherwise make no distinc- have pa.ssed since the first major paper on the tion between the type subspecies and speci- species of Eriastnim was published (Mason mens identified only to the species level. TAXONOMY Eriastnim Wooton and Standlev 22:71, nom. cons. Eriastrtim Wooton and Standley, Contr. U.S. Nat. Herb. 16:160. Hugelia Bentham, Bot. Reg. 19: under pi. 1622, 1913, Gilia and Navarretia of authors, in part. 18.33, not Huegelia Reichenbach, Consp. 144. 1828. Wehvitschia Reichenbach, Handb. 194. Erect, or spreading and erect, annuals or 1837, not Welwitschia Hooker, Card. Chron. perennials, virgately, racemosely, corymbosely, 'WcbiT State College, Ogden, Utah - 2 Bricham Young UNivEHsiTi- Science Bulletin diffusely, or paniculately branched; herbage verform, regular or irregular; stamens ecjually subglabrate to densely floccose; leaves linear, en- inserted at the base of the throat, on the throat tire or pinnately lobed; bracteate floral heads, or just below the sinuses, equal, subequal, or few or numerous, few- to many-flowered, rarely distinctly unequal; anthers sagittate, versatile; solitary, lightly floccose to densely so with capsule ellipsoid, three-loculed, few-seeded; arachnoid wool; calyx lobes unequal, joined seeds mucilaginous. along the lower two-thirds of their length by a Type species: E. filifolium (Nutt.) Woot. hyaline margin; corollas blue, yellow or white, & Standi. sometimes cream, rarely pink, funnelform to sal- Range: Western North America. Key to the Species A. Plants perennial, woody, branching from the base; stamens inserted at the sinus 1. E. densifolium AA. Plants annual, herbaceous, branching from the base or above; stamens inserted at the sinus or below. B. Corolla 15.0 to 23.0 mm long. C. Stamens inserted at the sinus; corolla salverform, or very narrowly funnelform — 2. E. phdriflonim CC. Stamens inserted at the base of the throat; corolla funnelform. D. Corolla bilabiate, or irregular to obscurely so; stamens distinctlv unequal, fila- ments often oriented toward lower lip, and anthers turned toward upper lip, lavender blue; desert areas 6. E. erernicuni DD. Corolla regular, or very nearly so; stamens usually equal, filaments not oriented toward one side of corolla, bright blue; Monterey County 3. E. virgatiim BB. Corolla 5.0 to 15.0 mm long. C. Stamens 5.0 to 10.0 mm long, nearly equalling or exceeding tips of the corolla lobes; anthers (restored)" 2.0 to 3.0 mm long. D. Corolla bright golden yellow, 8.0 mm long, regular to very slightly irregular; Monterey, San Luis Obispo counties 4. E. luteitm DD. Corolla bright sapphire blue, ca. 1.0 to 1.5 cm long, distinctlv irregular to ob- scurely so; southern California to Baja, 1000 ft to 9000 ft 5. E. sapphirinum CC. Stamens less than 5.0 mm long, not equalling or exceeding tips of corolla lobes; an- thers (restored) not more than 2.0 mm long. D. Stamens equal; corolla lobes 1.0 mm broad, or less; corolla narrowly funnelform; throat expanding slightly or not at all. E. Filaments 2.5 to 3.0 mm long; floral heads distinctly "brushy" with tips of bracts and calyx lobes conspicuous; ovary cylindric, three-times longer than wide 7. E. filifolium EE. Filaments 1.5 mm or less; tips of bracts and calyx lobes not conspicuously prominent; ovary 1 to 1.5 times longer than wide. F. Corolla 7.0 to 11.0 mm long; plants 15.0 to 35.0 cm high. G. Leaves mostly 1.0 to 1.5 cm long and entire, or to 3.0 cm and with one pair of lateral lobes; foliage often remaining somewhat soft; throat expanding slightly 8. E. sparsiflorum GO. Leaves mostly 1.5 to 2.5 cm long with one pair of lateral lobes, less commonly entire; foliage becoming rigid and brittle; throat not ex- panding. H. Corolla lobes 3.0 to 3.5 mm long and 1.0 mm bro;id . 11. E. hrandegeae 'The addition of a drop of 10 percent liquid detergent restores dried .intliors tn the size .it the time of dehiscence. Biological Series, Vol. 16, No. 4 The Genus Eriastrum 3 HH. Corolla lobes 2.5 mm long and 1.5 mm broad 12 E. traijci FF. Corolla 5.0 to 8.0 mm long; plants to 15.0 cm high. G. Leaves, when present, 2.0 to 4.5 cm long, with one to three pair of lateral lobes to 1.0 cm or longer; floral heads many-flowered, very densely floccose, often found under Adenostoma chaparral or on ob- sidian rubble 13. E. ahramsii GG. Leaves filiform, 0.5 to 2.5 cm long, entire or with one pair of lateral lobes 2.0 to 5.0 mm long; floral heads few-flowered, somewhat floc- cose, occurring in dry stream beds and on alkaline flats _ 14. E. booveri DD. Stamens unequal, corolla lobes 1.25 to 2.25 mm broad, corolla funnelform. E. Plants robust to 30.0 cm high, racemose, corymbose, or branched from the base; herbage floccose, often densely so; leaves 1.5 to 3.0 cm long, usually with one or two pair of lateral lobes; floral heads several-flowered 9. E. tuilcoxii EE. Plants to 15.0 cm high, diffusely branched from the base; stem and leaves slender; herbage lightly floccose; leaves 1.0 to 2.5 cm, usually with one pair of lateral lobes; floral heads few-flowered .
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