Species Assessment for Common Sanddragon
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Geographical Variation in the Wing Morphology of the Golden-Ringed
Bull. Natl. Mus. Nat. Sci., Ser. A, 38(2), pp. 65–73, May 22, 2012 Geographical Variation in the Wing Morphology of the Golden-ringed Dragonfly Anotogaster sieboldii (Selys, 1854) (Odonata, Cordulegastridae) Detected by Landmark-based Geometric Morphometrics Takuya Kiyoshi1 and Tsutomu Hikida2 1 Department of Zoology, National Museum of Nature and Science, 4–1–1 Amakubo, Tsukuba, Ibaraki, 305–0005 Japan E-mail: [email protected] 2 Department of Zoology, Graduate School of Science, Kyoto University, Kitashirakawaoiwakecho, Sakyo-ku, Kyoto, Kyoto, 606–8502 Japan (Received 18 January 2012; accepted 4 April 2012) Abstract A previous molecular phylogenetic study showed that Anotogaster sieboldii has at least 6 monophyletic groups that are allopatrically distributed in the northern area of Asia (Japanese main islands and Korean Peninsula), Amamioshima, Okinawajima, Yaeyama islands, Taiwan and East China. These groups are difficult to distinguish by qualitative genital morphology; however, canonical variate analysis and linear discriminant analysis of the hind wing shape have been used to clearly distinguish them from each other. In the present study, multiple comparisons of the lengths of the abdomen and hind wing showed that these lengths did not differ significantly among the groups. In particular, the variation in the size range, in the lineage from the northern area, widely overlapped that of the lineages from the other areas. In evaluating the morphological differ- ences, the wing shape was found to be more sensitive than other size variables. Key words : geometric morphometrics, shape, Odonata. Introduction groups (Askew, 2004). Anotogaster sieboldii (Selys, 1854) is a large The family Cordulegastridae of the order Odo- cordulegastrid dragonfly that is distributed in nata consists of 40 species belonging to 5 genera Insular East Asia, Korean Peninsula and East (Tsuda, 2000). -
Using Specimens from the Past to Understand the Living World Through Digitization
Using specimens from the past to understand the living world through digitization Drs. Jessica L. Ware, William Kuhn, Dirk Gassmann and John Abbott Rutgers University, Newark Dragonflies: Order Odonata Suborders Anisoptera (unequal wings): Dragonflies ~3000 species Anisozygoptera ~3 species Zygoptera: Damselflies ~3000 species Perchers, Fliers, Migrators, & Homebodies Dragonfly flight Wing venation affects wing camber, lift, and ultimately flight patterns Dragonfly flight Stiffness varies along length of the wing with vein density and thickness Dragonfly flight Certain wing traits are correlated with specific flight styles Dragonfly collections: invaluable treasures Collection name # spp. #specimens Florida State Collection 2728 150K Ware Lab Collection 373 4K Smithsonian Collection 253 200K M.L. May Collection 300 10K Dragonfly collections: invaluable treasures Harness information in collections Targeted Odonata Wing Digitization (TOWD) project TOWD project scanning protocol TOWD project scanning protocol TOWD project TOWD project TOWD project 10 10 TOWD project More information at: https://willkuhn.github.io/towd/ Aspect ratios: How elongate is the wing compared to its overall area? • High Aspect ratio Low Aspect ratio Long narrow wings, Wing Short broad wings, optimized for: Wing optimized long-distance flight for: Maneuverability, turning High Aspect Low Aspect Ratio Ratio More data on aspect ratios, better interpretations? 2007: Hand measured forewings of 85 specimens, 7 months work More data on apsect ratios, better interpretations? 2007: Hand measured forewings of 85 specimens, 7 months 2019: Odomatic measurements for 206 work specimens, 2-3 minutes of work! Are there differences in aspect ratios? Perchers have significantly lower aspect ratios than fliers. The p-value is .001819. The result is significant at p < .05. -
Dragonflies (Odonata) of the Northwest Territories Status Ranking And
DRAGONFLIES (ODONATA) OF THE NORTHWEST TERRITORIES STATUS RANKING AND PRELIMINARY ATLAS PAUL M. CATLING University of Ottawa 2003 TABLE OF CONTENTS Abstract ....................................................................3 Acknowledgements ...........................................................3 Methods ....................................................................3 The database .................................................................4 History .....................................................................5 Rejected taxa ................................................................5 Possible additions ............................................................5 Additional field inventory ......................................................7 Collection an Inventory of dragonflies .............................................8 Literature Cited .............................................................10 Appendix Table 1 - checklist ...................................................13 Appendix Table 2 - Atlas and ranking notes .......................................15 2 ABSTRACT: occurrences was provided by Dr. Rex Thirty-five species of Odonata are given Kenner, Dr. Donna Giberson, Dr. Nick status ranks in the Northwest Territories Donnelly and Dr. Robert Cannings (some based on number of occurrences and details provided below). General distributional area within the territory. Nine information on contacts and locations of species are ranked as S2, may be at risk, collections provided by Dr. Cannings -
A Survey of Odonata of the Patoka River National Wildlife Refuge and Management Area
2012. Proceedings of the Indiana Academy of Science 121(1):54–61 A SURVEY OF ODONATA OF THE PATOKA RIVER NATIONAL WILDLIFE REFUGE AND MANAGEMENT AREA Donald L. Batema* and Amanda Bellian: Department of Chemistry, Environmental Studies Program, University of Evansville, 1800 Lincoln Avenue, Evansville, IN 47722 USA Lindsey Landowski: Mingo National Wildlife Refuge, Puxico, MO. 63960 USA ABSTRACT. The Patoka River National Wildlife Refuge and Management Area (hereafter Patoka River Refuge or the Refuge) represents one of the largest intact bottomland hardwood forests in southern Indiana, with meandering oxbows, marshes, ponds, managed moist-soil units, and constructed wetlands that provide diverse and suitable habitat for wildlife. Refuge personnel strive to protect, restore, and manage this bottomland hardwood ecosystem and associated habitats for a variety of wildlife. The Patoka River National Wildlife Refuge Comprehensive Conservation Plan (CCP) lists many species of management priority (McCoy 2008), but Odonata are not included, even though they are known to occur on the Refuge. The absence of Odonata from the CCP is the result of lack of information about this ecologically important group of organisms. Therefore, we conducted a survey, from May to October 2009, to document their presence, with special attention being paid to rare, threatened, and endangered species. A total of 43 dragonfly and damselfly species were collected and identified. No threatened or endangered species were found on the Refuge, but three species were found that are considered imperiled in Indiana based on Nature Serve Ranks (Stein 2002). Additionally, 19 new odonate records were documented for Pike County, Indiana. The results of this survey will be used by Refuge personnel to assist in management decisions and to help establish priorities for the Patoka River Refuge activities and land acquisition goals. -
Swaegers Etal 2014
doi: 10.1111/jeb.12481 Ecological and evolutionary drivers of range size in Coenagrion damselflies J. SWAEGERS*, S. B. JANSSENS†,S.FERREIRA‡§¶,P.C.WATTS¶**, J. MERGEAY††, M. A. MC PEEK‡‡ &R.STOKS* *Laboratory of Aquatic Ecology, Evolution and Conservation, KU Leuven, Leuven, Belgium †Botanic Garden Meise, Meise, Belgium ‡CIBIO/InBIO – Centro de Investigacßao~ em Biodiversidade e Recursos Geneticos da Universidade do Porto, Vairao,~ Portugal §Departamento de Biologia da Faculdade de Ci^encias da Universidade do Porto, Porto, Portugal ¶Institute of Integrative Biology, University of Liverpool, Liverpool, UK **Department of Biology, University of Oulu, Oulu, Finland ††Research Institute for Nature and Forest, Geraardsbergen, Belgium ‡‡Department of Biological Sciences, Dartmouth College, Hanover, NH, USA Keywords: Abstract Bergmann’s rule; Geographic range size is a key ecological and evolutionary characteristic of a dispersal capacity; species, yet the causal basis of variation in range size among species remains latitudinal patterns; largely unresolved. One major reason for this is that several ecological and phylogeny; evolutionary traits may jointly shape species’ differences in range size. We range size; here present an integrated study of the contribution of ecological (dispersal Rapoport’s rule. capacity, body size and latitudinal position) and macroevolutionary (species’ age) traits in shaping variation in species’ range size in Coenagrion damsel- flies. We reconstructed the phylogenetic tree of this genus to account for evolutionary history when assessing the contribution of the ecological traits and to evaluate the role of the macroevolutionary trait (species’ age). The genus invaded the Nearctic twice independently from the Palearctic, yet this was not associated with the evolution of larger range sizes or dispersal capacity. -
Ecography ECOG-02578 Pinkert, S., Brandl, R
Ecography ECOG-02578 Pinkert, S., Brandl, R. and Zeuss, D. 2016. Colour lightness of dragonfly assemblages across North America and Europe. – Ecography doi: 10.1111/ecog.02578 Supplementary material Appendix 1 Figures A1–A12, Table A1 and A2 1 Figure A1. Scatterplots between female and male colour lightness of 44 North American (Needham et al. 2000) and 19 European (Askew 1988) dragonfly species. Note that colour lightness of females and males is highly correlated. 2 Figure A2. Correlation of the average colour lightness of European dragonfly species illustrated in both Askew (1988) and Dijkstra and Lewington (2006). Average colour lightness ranges from 0 (absolute black) to 255 (pure white). Note that the extracted colour values of dorsal dragonfly drawings from both sources are highly correlated. 3 Figure A3. Frequency distribution of the average colour lightness of 152 North American and 74 European dragonfly species. Average colour lightness ranges from 0 (absolute black) to 255 (pure white). Rugs at the abscissa indicate the value of each species. Note that colour values are from different sources (North America: Needham et al. 2000, Europe: Askew 1988), and hence absolute values are not directly comparable. 4 Figure A4. Scatterplots of single ordinary least-squares regressions between average colour lightness of 8,127 North American dragonfly assemblages and mean temperature of the warmest quarter. Red dots represent assemblages that were excluded from the analysis because they contained less than five species. Note that those assemblages that were excluded scatter more than those with more than five species (c.f. the coefficients of determination) due to the inherent effect of very low sampling sizes. -
175 Post-Copulatory Behaviour in Calopteryx
International Journal of Odonatology 1 (2): 175-184, 1998. © 1998 Backhuys Publishers. 175 POST-COPULATORY BEHAVIOUR IN CALOPTERYX FEMALES (INSECTA, ODONATA, CALOPTERYGIDAE). Martin Lindeboom Institut fiir systematische Zoologie, Eberhard-Karls-Universitiit Ttibingen, Auf der Morgenstelle 28, 72076 Ttibingen, Germany, e-mail: [email protected] Received 30 May 1998; revised 12 October 1998; accepted 13 October 1998. Key words: Odonata, Calopteryx, post-copulatory behaviour, functional morphology, ovipositor, copulatory organ, microstructures, sperm removal, cryptic female choice. Abstract The post-copulatory behaviour of Calopteryx splendens females was studied under field and laboratory conditions. After termination of copulation females usually perch and bend the abdomen so that its apex touches the ground (post-copulatory posture). The post-cop ulatory posture is a consequence of sperm removal by males. Male and female microstruc tures (spines and scales) interact to move previous sperm from the female sperm storage organs to the outside during copulation stage I, after which moved sperm is located on the ovipositor. After termination of copulation females require an average of 45 seconds to brush off this sperm (N=21). The post-copulatory behaviour of females may also allow males to chase rival males before the females start to oviposit (prevention of dis turbances). The present study shows no evidence of cryptic female choice in C. splendens. Introduction Sperm competition in Odonata has resulted in the evolution of direct sperm removal in Zygoptera and Anisoptera. Sperm removal resulting in last male sperm precedence seems to be widespread in Zygoptera (Waage 1979a, 1984, 1986; Miller & Miller 1981; Fincke 1984; Siva-Jothy & Tsubaki 1989; Cordero & Miller 1992; Siva-Jothy & Hooper 1995). -
A Checklist of North American Odonata
A Checklist of North American Odonata Including English Name, Etymology, Type Locality, and Distribution Dennis R. Paulson and Sidney W. Dunkle 2009 Edition (updated 14 April 2009) A Checklist of North American Odonata Including English Name, Etymology, Type Locality, and Distribution 2009 Edition (updated 14 April 2009) Dennis R. Paulson1 and Sidney W. Dunkle2 Originally published as Occasional Paper No. 56, Slater Museum of Natural History, University of Puget Sound, June 1999; completely revised March 2009. Copyright © 2009 Dennis R. Paulson and Sidney W. Dunkle 2009 edition published by Jim Johnson Cover photo: Tramea carolina (Carolina Saddlebags), Cabin Lake, Aiken Co., South Carolina, 13 May 2008, Dennis Paulson. 1 1724 NE 98 Street, Seattle, WA 98115 2 8030 Lakeside Parkway, Apt. 8208, Tucson, AZ 85730 ABSTRACT The checklist includes all 457 species of North American Odonata considered valid at this time. For each species the original citation, English name, type locality, etymology of both scientific and English names, and approxi- mate distribution are given. Literature citations for original descriptions of all species are given in the appended list of references. INTRODUCTION Before the first edition of this checklist there was no re- Table 1. The families of North American Odonata, cent checklist of North American Odonata. Muttkows- with number of species. ki (1910) and Needham and Heywood (1929) are long out of date. The Zygoptera and Anisoptera were cov- Family Genera Species ered by Westfall and May (2006) and Needham, West- fall, and May (2000), respectively, but some changes Calopterygidae 2 8 in nomenclature have been made subsequently. Davies Lestidae 2 19 and Tobin (1984, 1985) listed the world odonate fauna Coenagrionidae 15 103 but did not include type localities or details of distri- Platystictidae 1 1 bution. -
Argia the News Journal of the Dragonfly Society of the Americas
ISSN 1061-8503 TheA News Journalrgia of the Dragonfly Society of the Americas Volume 22 17 December 2010 Number 4 Published by the Dragonfly Society of the Americas http://www.DragonflySocietyAmericas.org/ ARGIA Vol. 22, No. 4, 17 December 2010 In This Issue .................................................................................................................................................................1 Calendar of Events ......................................................................................................................................................1 Minutes of the 2010 Annual Meeting of the Dragonfly Society of the Americas, by Steve Valley ............................2 2010 Treasurer’s Report, by Jerrell J. Daigle ................................................................................................................2 Enallagma novaehispaniae Calvert (Neotropical Bluet), Another New Species for Arizona, by Rich Bailowitz ......3 Photos Needed ............................................................................................................................................................3 Lestes australis (Southern Spreadwing), New for Arizona, by Rich Bailowitz ...........................................................4 Ischnura barberi (Desert Forktail) Found in Oregon, by Jim Johnson ........................................................................4 Recent Discoveries in Montana, by Nathan S. Kohler ...............................................................................................5 -
Odonata: Aeshnidae): a Study of Traits from Larval Development to Adults
EUROPEAN JOURNAL OF ENTOMOLOGYENTOMOLOGY ISSN (online): 1802-8829 Eur. J. Entomol. 116: 269–280, 2019 http://www.eje.cz doi: 10.14411/eje.2019.031 ORIGINAL ARTICLE Biometry of the large dragonfl y Anax imperator (Odonata: Aeshnidae): A study of traits from larval development to adults MARCEAU MINOT 1, MICKAËL LE GALL2 and AURÉLIE HUSTÉ 1 1 Université de Rouen - ECODIV, Bat Blondel, Place Emile Blondel, Mont-Saint-Aignan 76821, France; e-mails: [email protected], [email protected] 2 IRSTEA, UR RIVERLY, Centre de Lyon – Villeurbanne, 69625 Villeurbanne Cedex, France; e-mail: [email protected] Key words. Odonata, Aeshnidae, Anax imperator, body length, body mass, larval rearing, sexual size dimorphism, traits Abstract. Insect larval development affects adult traits but the biometric relationships are usually poorly understood, including large odonates. In this study, measurements of morphological traits of larvae, exuviae and adults of Anax imperator were recorded. They were used to investigate the effects of early development on adult morphology. Results showed an increase in larval length during the fi nal instar and the length of its exuviae signifi cantly exceeded that of the larva. Length and body mass of teneral adults were strongly related to the length of their exuviae. Adult males were signifi cantly longer than adult females, while both had the same body mass at emergence. Length of teneral adults was negatively related to the date of emergence in both sexes. During maturation, body mass of males only increased slightly whereas that of females increased greatly. Mature specimens were also signifi cantly longer than teneral individuals. -
Aquatic and Terrestrial Vegetation Influence
AQUATIC AND TERRESTRIAL VEGETATION INFLUENCE LACUSTRINE DRAGONFLY (ORDER ODONATA) ASSEMBLAGES AT MULTIPLE LIFE STAGES By Alysa J. Remsburg A dissertation submitted in partial fulfillment of the requirements for the degree of Doctor of Philosophy (Zoology) at the UNIVERSITY OF WISCONSIN – MADISON 2007 i ACKNOWLEDGMENTS Reflecting on the contributions of my colleagues and friends during my graduate studies gives me a strong sense of gratitude for the community of support that I have enjoyed. The people who surround and support me deserve more thanks than I can describe here. Friends and family have supported my graduate studies by generously accommodating my tight schedule and warmly offering encouragement throughout the process. Monica Turner guided my graduate studies in numerous ways. It was her trust in my abilities and willingness to learn about a new study organism that first made this research possible. She encouraged me to pursue the research questions that most interested and inspired me, although it meant charting territory that was new to both of us. Monica served as the ideal mentor for me by requiring clear communication, modeling an efficient and balanced work ethic, providing critical reviews, and listening compassionately. This research benefited from the expertise and generosity of outstanding Wisconsin ecologists. Members of my graduate research committee, Steve Carpenter, Claudio Gratton, Tony Ives, Bobbi Peckarsky, and Joy Zedler, all offered useful suggestions and critiques on experimental design, pressing research questions, and the manuscripts. Cecile Ane provided additional statistical advice and smiles. Bill Smith, Bob DuBois, and Robert Bohanan answered (or reassured me that I should try to answer) many questions about field methods, Odonata biology, and species identification. -
Simultaneous Control of Head and Thoracic Temperature by the Green Darner Dragonfly Anax Junius (Odonata: Aeshnidae)
The Journal of Experimental Biology 198, 2373–2384 (1995) 2373 Printed in Great Britain © The Company of Biologists Limited 1995 SIMULTANEOUS CONTROL OF HEAD AND THORACIC TEMPERATURE BY THE GREEN DARNER DRAGONFLY ANAX JUNIUS (ODONATA: AESHNIDAE) MICHAEL L. MAY Department of Entomology, Cook College, New Jersey Agricultural Experiment Station, Rutgers University, New Brunswick, NJ 08903, USA Accepted 24 July 1995 Summary Anax junius is a large dragonfly that regulates thoracic during unrestrained flight in the field, Th is regulated temperature (Tth) during flight. This species, like several actively by increasing hemolymph circulation from the other intermittently endothermic insects, achieves control warm thorax at low Ta. Concurrent measurements of of Tth at least in part by increasing circulation of abdominal temperature (Tab) confirm that the abdomen is hemolymph to the abdomen at high air temperature (Ta), used as a ‘thermal window’ at Ta>30 ˚C but apparently not thus facilitating heat loss from the thorax. In this paper, I at lower Ta; thus, some additional mechanism(s) must exist demonstrate that heat transfer to the head is also under for regulation of Tth at low Ta. active control, very probably owing to temperature- sensitive alteration of hemolymph circulation. As a result, head temperature (Th) is strikingly elevated above Ta Key words: Anax junius, Anisoptera, body temperature, circulatory during endothermic warm-up and flight. Furthermore, control, dragonfly, green darner, heat exchange, thermoregulation. Introduction Numerous insects regulate Tth (most recently and The primary aim of this study is to investigate the sources comprehensively reviewed by Heinrich, 1993), among them of variation of Th, its mechanism of control and its responses the subject of this paper Anax junius (Drury) (Heinrich and to environmental temperature and internal variables in A.