100 Years of Change in the Flora of the Carolinas
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EUPHORBIACEAE 353 Tragia urticifolia Michaux, Nettleleaf Noseburn. Pd (GA, NC, SC, VA), Cp (GA, SC), Mt (SC): dry woodlands and rock outcrops, particularly over mafic or calcareous rocks; common (VA Rare). May-October. Sc. VA west to MO, KS, and CO, south to FL and AZ. [= RAB, F, G, K, W; = T. urticaefolia – S, orthographic variant] Triadica Loureiro 1790 (Chinese Tallow-tree) A genus of 2-3 species, native to tropical and subtropical Asia. The most recent monographers of Sapium and related genera (Kruijt 1996; Esser 2002) place our single naturalized species in the genus Triadica, native to Asia; Sapium (excluding Triadica) is a genus of 21 species restricted to the neotropics. This conclusion is corroborated by molecular phylogenetic analysis (Wurdack, Hoffmann, & Chase (2005). References: Kruijt (1996)=Z; Esser (2002)=Y; Govaerts, Frodin, & Radcliffe-Smith (2000)=X. * Triadica sebifera (Linnaeus) Small, Chinese Tallow-tree, Popcorn Tree. Cp (GA, NC, SC): marsh edges, shell deposits, disturbed areas; uncommon. May-June; August-November, native of e. Asia. With Euphorbia, Chamaesyce, and Cnidoscolus, one of our few Euphorbiaceous genera with milky sap. Triadica has become locally common from Colleton County, SC southward through the tidewater area of GA, and promises to become a serious weed tree (as it is in parts of LA, TX, and FL). [= K, S, X, Y, Z; = Sapium sebiferum (Linnaeus) Roxburgh – RAB, GW] Vernicia Loureiro 1790 (Tung-oil Tree) A genus of 3 species, trees, native of se. Asia. References: Govaerts, Frodin, & Radcliffe-Smith (2000)=Z. * Vernicia fordii (Hemsley) Airy-Shaw, Tung-oil Tree, Tung Tree. Cp (GA, NC): planted for the oil and for ornament, rarely naturalizing; rare, introduced from central and western China. Naturalized in GA, from former plantations; planted and showing a tendency to naturalize in the Coastal Plain of NC (Mount Olive, Wayne Co.). [= K, Z; = Aleurites fordii Hemsley] FABACEAE Lindley 1836 or LEGUMINOSAE A.L. de Jussieu 1789 (Legume Family) A family of about 730 genera and 20,000 species, trees, shrubs, and herbs, cosmopolitan. References: Isely (1990)=SE (throughout the family treatment); Wojciechowski, Lavin, & Sanderson (2004); Wilbur (1963); Isely (1998)=I; Robertson & Lee (1976). 1 Trees, shrubs, or woody vines...........................................................................................................................................Key A 1 Herbs (including herbaceous vines). 2 Leaves 4-many-foliolate. 3 Leaves palmately compound, with 4 or more leaflets .......................................................................................Key B 3 Leaves pinnately or bipinnately compound. 4 Leaves bipinnately compound...................................................................................................................Key C 4 Leaves pinnately compound......................................................................................................................Key D 2 Leaves 1-3-foliolate. 5 Leaves unifoliolate............................................................................................................................................Key E 5 Leaves trifoliolate. 6 Leaves pinnately trifoliolate...................................................................................................................... Key F 6 Leaves palmately trifoliolate.................................................................................................................... Key G Key A – woody legumes (trees, shrubs, or woody vines) 1 Leaves unifoliolate or trifoliolate, or reduced to phyllodial spines. 2 Tree; leaves unifoliolate and > 5 cm wide; [subfamily Caesalpinioideae, tribe Cercideae]..................................... Cercis 2 Shrubs or woody vines (rarely tree in Erythrina); leaves trifoliolate, unifoliolate, or reduced to phyllodial spines (if unifoliolate, < 2 cm wide); [subfamily Papilionoideae]. 3 Woody vine. 4 Calyx 4.5-6 mm long; leaflets unlobed; [tribe Phaseoleae, subtribe Diocleinae] ...................................Dioclea 4 Calyx 10-12 mm long; leaflets generally lobed; [tribe Phaseoleae, subtribe Glycininae] ....................Pueraria 3 Shrub or tree. 5 Shrub or tree with twigs various, but not conspicuously green or flanged; leaves pinnately trifoliolate. 6 Corolla 30-50 mm long, scarlet; legume with several seeds; leaflets lobed; [tribe Phaseoleae, subtribe Erythrininae]............................................................................................................................... Erythrina 6 Corolla 8-15 mm long, purplish, pink, or white; legume 1-seeded; leaflets not lobed; [tribe Desmodieae, subtribe Lespedezinae] ................................................................................................................Lespedeza FABACEAE 354 5 Shrub with angled or flanged green twigs; leaves palmately trifoliolate, unifoliolate, or reduced to spine- tipped phyllodes; flowers bright yellow; [introduced, usually of roadsides or as remnants of cultivation]; [tribe Genisteae, subtribe Genistinae]. 7 Leaves all reduced to phyllodial spines; flowers axillary; calyx 10-15 mm long .................................Ulex 7 Leaves with normal lamina, either unifoliolate or trifoliolate; flowers in terminal racemes; calyx 3-6 mm long. 8 Leaves trifoliolate lower on the stem, often unifoliolate above; corolla 15-22 mm long..........Cytisus 8 Leaves unifoliolate throughout; corolla 10-14 mm long....................................................... [Genista] 1 Leaves pinnate. 9 Woody vines; [subfamily Papilionoideae]. 10 Leaves even-pinnate; legume 3.5-4 cm long; seeds shiny scarlet and black; [tribe Abreae]...........................[Abrus] 10 Leaves odd-pinnate; legume 4-15 cm long; seeds brown; [tribe Milletieae]..................................................Wisteria 9 Trees or shrubs. 11 Leaves 2×-paripinnate; [subfamily Mimosoideae] 12 Stamens connate at the base; inflorescence pink, 2.5-5 cm in diameter; [tribe Ingeae] ...........................Albizia 12 Stamens free; inflorescence orange or yellowish-white, 1.0-2.2 cm in diameter. 13 Inflorescence yellowish-white, 1.8-2.2 cm in diameter; stamens 10; [tribe Mimoseae] .............. Leucaena 13 Inflorescence orange, 1.0-1.3 cm in diameter; stamens many; [tribe Acacieae]...........................Vachellia 11 Leaves otherwise. 14 [subfamily Papilionoideae]. 15 Leaves glandular-punctate; corolla of only 1 petal; inflorescence a spike; shrubs; [tribe Amorpheae] ....... .....................................................................................................................................................Amorpha 15 Leaves not glandular-punctate; corolla of 5 petals; inflorescence a raceme or panicle; trees or shrubs. 16 Leaflets alternate on the rachis; leaflets 4-15 (-20) cm long; [tribe Sophoreae]..................Cladrastis 16 Leaflets opposite on the rachis, leaflets (1-) 2-5 (-6) cm long. 17 Leaflets with persistent linear stipels; native and cultivated, collectively widespread in our area; [tribe Robinieae] .................................................................................................... Robinia 17 Leaflets lacking stipels; cultivated, perhaps not established; [tribe Sophoreae].......................... ..........................................................................................................................[Styphnolobium] 14 [subfamily Caesalpinioideae]. 18 Leaves all 2-pinnate, or a mixture of 2-pinnate and 1-pinnate on the same plant; shrub or tree; [tribe Caesalpineae]. 19 Leaves a mixture of 1-pinnate and 2-pinnate.........................................................................Gleditsia 19 Leaves all 2-pinnate. 20 Leaves petiolate; leaflets 20-70 mm long ..............................................................Gymnocladus 20 Leaves subsessile (the pinnae simulating 1-pinnate leaves); leaflets 1-5 mm long...................... ................................................................................................................................. Parkinsonia 18 Leaves all 1-pinnate (or appearing so in Parkinsonia); herb, shrub or tree. 21 Shrub with prominent glands on the leafstalk; [tribe Cassieae] ................................................ Senna 21 Tree or shrub (if a shrub, then lacking prominent glands on the leafstalk); [tribe Caesalpineae]. 22 Leaflets 13-45 mm long; tree; leaves 1-pinnate.............................................................Gleditsia 22 Leaflets 1-5 mm long; shrub; leaves actually 2-pinnate, but subsessile, the pinnae simulating 1-pinnate leaves....................................................................................................... Parkinsonia Key B – herbaceous legumes with palmate leaves with 4 or more leaflets Lupinus, Orbexilum, Pediomelum, Psoralidium, Zornia Key C – herbaceous legumes with bipinnate leaves 1 Petiole with 1-several glands; stems ascending to erect; flowers greenish-white ...................................................Desmanthus 1 Petiole without glands; stems prostrate to weakly arching; flowers pink-purple, yellow, or greenish-yellow. 2 Flowers pink-purple; legume ribbed, the ribs with prickles ................................................................................... Mimosa 2 Flowers yellow to greenish-yellow; legume not ribbed or prickly.....................................................................[Neptunia] Key D – pinnate plus Aeschynomene, Apios, Arachis, Astragalus, Clitoria,