A peer-reviewed open-access journal ZooKeys 315: 17–54 Western (2013) Hemisphere Zuphiini: descriptions of Coarazuphium whiteheadi... 17 doi: 10.3897/zookeys.315.5293 RESEARCH ARTICLE www.zookeys.org Launched to accelerate biodiversity research Western Hemisphere Zuphiini: descriptions of Coarazuphium whiteheadi, new species, and Zuphioides, new genus, and classification of the genera (Coleoptera, Carabidae) George E. Ball1,†, Danny Shpeley1,‡ 1 Department of Biological Sciences, University of Alberta, Edmonton, Alberta T5G 2E9, Canada † urn:lsid:zoobank.org:author:51D69BDE-0573-4366-9120-561BA15289C1 ‡ urn:lsid:zoobank.org:author:406FFF55-7AE7-40B8-9D10-42D24A7A22E6 Corresponding author: Danny Shpeley (
[email protected]) Academic editor: T. Assmann | Received 11 April 2013 | Accepted 14 June 2013 | Published 4 July 2013 urn:lsid:zoobank.org:pub:B5525448-9DBF-41E2-957B-71A6CB1872CA Citation: Ball GE, Shpeley D (2013) Western Hemisphere Zuphiini: descriptions of Coarazuphium whiteheadi, new species, and Zuphioides, new genus, and classification of the genera (Coleoptera, Carabidae). ZooKeys 315: 17–54. doi: 10.3897/zookeys.315.5293 Abstract Based on small samples (exemplars) analyzed with morphological methods, including detailed descrip- tions and illustrations, this study treats primarily the Zuphium genus-group in the Western Hemisphere, which comprises two precinctive genera: Coarazuphium Gnaspini, Vanin & Godoy, 1998 (type species Parazuphium tessai Godoy & Vanin, 1990) and Zuphioides gen. n. (type species Zuphium mexicanum Chaudoir, 1863). The genus Coarazuphium includes six troglobitic species from Brazilian caves, and one probably hypogaeic (troglophilic) species from the mountains of Oaxaca, in Mexico (C. whiteheadi, sp. n., type locality, ridge top, in western Oaxaca, Mexico, at 2164 m, 35 km north of San Pedro Juchatengo, 16.462N, 97.010W).