Chapter 13: the Flower and Sexual Reproduction
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Inflorescences
Inflorescences - Floral Displays Inflorescences - Floral Displays A shift from widely The vast majority of flowering plants spaced single flowers to an inflorescence required possess flowers in clusters called an condensation of shoots inflorescence. and the loss of the intervening leaves. These clusters facilitate pollination via a prominent visual display and more The simplest efficient pollen uptake and deposition. inflorescence type would thus be indeterminate with the oldest flowers at the base and the younger flowers progressively closer to the apical meristem of the shoot. = a raceme Raceme (Prunus or cherry) One modification of the basic raceme is to make it compound The panicle is essentially compound a series of attached racemes with the oldest racemes at the base and the youngest at the apex of the inflorescence. Panicle (Zigadenus or white camass) Raceme Panicle 1 A second modification of the basic raceme is to lose its pedicels The spike is usually associated with congested reduced flowers and often, but not always, with wind Pedicel loss pollination. wind pollinated Spike (Plantago or plantain) Raceme Spike A third modification of the basic raceme is to lose its internodes The spike is usually associated with congested reduced flowers and often, animal pollinated but not always, with wind Internode loss pollination. Umbel Spike (Combretum - Brent’s plants) Raceme 2 The umbel characterizes specific families (carrot and The umbel is found scattered in many other families ginseng families for example). as well. These families typically show a compound umbel - smaller umbellets on a larger umbel. Umbel Umbel (Cicuta or water hemlock) (Zizia or golden alexander) (Eriogonum or false buckwheat - family Polygonaceae) - Ben’s plants A fourth modification of the basic raceme is for the stem axis to form a head The head or capitulum characterizes specific families - most notably the Compositae or Asteraceae. -
CHESTNUT (CASTANEA Spp.) CULTIVAR EVALUATION for COMMERCIAL CHESTNUT PRODUCTION
CHESTNUT (CASTANEA spp.) CULTIVAR EVALUATION FOR COMMERCIAL CHESTNUT PRODUCTION IN HAMILTON COUNTY, TENNESSEE By Ana Maria Metaxas Approved: James Hill Craddock Jennifer Boyd Professor of Biological Sciences Assistant Professor of Biological and Environmental Sciences (Director of Thesis) (Committee Member) Gregory Reighard Jeffery Elwell Professor of Horticulture Dean, College of Arts and Sciences (Committee Member) A. Jerald Ainsworth Dean of the Graduate School CHESTNUT (CASTANEA spp.) CULTIVAR EVALUATION FOR COMMERCIAL CHESTNUT PRODUCTION IN HAMILTON COUNTY, TENNESSEE by Ana Maria Metaxas A Thesis Submitted to the Faculty of the University of Tennessee at Chattanooga in Partial Fulfillment of the Requirements for the Degree of Master of Science in Environmental Science May 2013 ii ABSTRACT Chestnut cultivars were evaluated for their commercial applicability under the environmental conditions in Hamilton County, TN at 35°13ꞌ 45ꞌꞌ N 85° 00ꞌ 03.97ꞌꞌ W elevation 230 meters. In 2003 and 2004, 534 trees were planted, representing 64 different cultivars, varieties, and species. Twenty trees from each of 20 different cultivars were planted as five-tree plots in a randomized complete block design in four blocks of 100 trees each, amounting to 400 trees. The remaining 44 chestnut cultivars, varieties, and species served as a germplasm collection. These were planted in guard rows surrounding the four blocks in completely randomized, single-tree plots. In the analysis, we investigated our collection predominantly with the aim to: 1) discover the degree of acclimation of grower- recommended cultivars to southeastern Tennessee climatic conditions and 2) ascertain the cultivars’ ability to survive in the area with Cryphonectria parasitica and other chestnut diseases and pests present. -
An Abstract of the Thesis Of
AN ABSTRACT OF THE THESIS OF Annie M. Chozinski for the degree of Master of Science in Horticulture presented on November 23. 1994. Title: The Evaluation of Cold Hardiness in Corvlus. Abstract approved: Shawn A. Mehlenbacher Anthesis of both staminate and pistillate flowers of Cory/us occurs in midwinter. To insure adequate pollination and nut set, these flowers must attain a sufficient hardiness level to withstand low temperatures. This study estimated cold hardiness of Cory/us cultivars and species using laboratory freezing of shoots without artificial hardening. In December, January and February of 1991-92 and 1992-93, one-year stems were collected 0 0 and frozen at regular intervals from -10 C to -38 C/ and visual browning assessed survival approximately 10 days after freezing. Elongated catkins were clipped prior to freezing. Percent flower bud survival was calculated and plotted against temperature. Linear regression generated an equation relating percent bud survival to temperature. From this equation, estimates of the LT^ (lethal temperature for 50% of the buds) was calculated for catkins, female inflorescences, and vegetative buds. C. avellana L. catkins, on average, were less hardy in both December and January than female inflorescences and vegetative buds. Maximum hardiness was reached in December and nearly all had elongated prior to the February freeze. Cultivars with the most hardy catkins were 'Morell', 'Brixnut', 'Creswell', 'Gem', 'Giresun OSU 54.080', 'Hall's Giant', 'Riccia di Talanico', 'Montebello' and 'Rode Zeller'. Maximum hardiness was observed for both vegetative and pistillate buds in January and was followed by a marked loss of hardiness in February. -
Species Profile for California Live Oak (Quercus Agrifolia)
California Phenology Project: species profile for California Live Oak (Quercus agrifolia) CPP site(s) where this species is monitored: Redwood Regional Park, Roberts Regional Recreation Area What does this species look like? This large evergreen tree has a dark grey, stout, short trunk and wide spreading branches. The leathery leaves are shiny on the upper surface and dull on the lower surface, which is covered with fuzzy hairs. The leaf margins are spiny and holly-like. The individuals are monoecious; each tree bears both male and female flowers but the male flowers produce only anthers and the female flowers produce only pistils. The yellow-green male flowers are clustered in elongated, drooping catkins that are 4-10 cm long, and the female flowers are clustered in reddish green spikes. When monitoring this species, use the USA-NPN broadleaf evergreen (with pollen) trees and shrubs Photo credit: randomtruth (Flickr) datasheet. Species facts! • The CPP four letter code for this species is QUAG. • This oak is very fire resistant. Adaptations to fire include evergreen leaves, thick bark, and the ability to sprout post-fire from the roots, trunk, and upper crown. • Individuals can live up to 250 years. • Susceptible to Sudden Oak Death disease. • Wind pollinated. • Each acorn takes a full year to develop from a pollinated flower. Photo credit: randomtruth (Flickr) Where is this species found? • In valleys, slopes, mixed-evergreen forest, and woodlands at elevations less than 1500 meters. • Endemic to California; found in North Coast Ranges, Central Western California, and SW California. • Occurs on soils ranging from silts and clays to weathered granite. -
Plant Terminology
PLANT TERMINOLOGY Plant terminology for the identification of plants is a necessary evil in order to be more exact, to cut down on lengthy descriptions, and of course to use the more professional texts. I have tried to keep the terminology in the database fairly simple but there is no choice in using many descriptive terms. The following slides deal with the most commonly used terms (more specialized terms are given in family descriptions where needed). Professional texts vary from fairly friendly to down-right difficult in their use of terminology. Do not be dismayed if a plant or plant part does not seem to fit any given term, or that some terms seem to be vague or have more than one definition – that’s life. In addition this subject has deep historical roots and plant terminology has evolved with the science although some authors have not. There are many texts that define and illustrate plant terminology – I use Plant Identification Terminology, An illustrated Glossary by Harris and Harris (see CREDITS) and others. Most plant books have at least some terms defined. To really begin to appreciate the diversity of plants, a good text on plant systematics or Classification is a necessity. PLANT TERMS - Typical Plant - Introduction [V. Max Brown] Plant Shoot System of Plant – stem, leaves and flowers. This is the photosynthetic part of the plant using CO2 (from the air) and light to produce food which is used by the plant and stored in the Root System. The shoot system is also the reproductive part of the plant forming flowers (highly modified leaves); however some plants also have forms of asexual reproduction The stem is composed of Nodes (points of origin for leaves and branches) and Internodes Root System of Plant – supports the plant, stores food and uptakes water and minerals used in the shoot System PLANT TERMS - Typical Perfect Flower [V. -
Auxin Regulation Involved in Gynoecium Morphogenesis of Papaya Flowers
Zhou et al. Horticulture Research (2019) 6:119 Horticulture Research https://doi.org/10.1038/s41438-019-0205-8 www.nature.com/hortres ARTICLE Open Access Auxin regulation involved in gynoecium morphogenesis of papaya flowers Ping Zhou 1,2,MahparaFatima3,XinyiMa1,JuanLiu1 and Ray Ming 1,4 Abstract The morphogenesis of gynoecium is crucial for propagation and productivity of fruit crops. For trioecious papaya (Carica papaya), highly differentiated morphology of gynoecium in flowers of different sex types is controlled by gene networks and influenced by environmental factors, but the regulatory mechanism in gynoecium morphogenesis is unclear. Gynodioecious and dioecious papaya varieties were used for analysis of differentially expressed genes followed by experiments using auxin and an auxin transporter inhibitor. We first compared differential gene expression in functional and rudimentary gynoecium at early stage of their development and detected significant difference in phytohormone modulating and transduction processes, particularly auxin. Enhanced auxin signal transduction in rudimentary gynoecium was observed. To determine the role auxin plays in the papaya gynoecium, auxin transport inhibitor (N-1-Naphthylphthalamic acid, NPA) and synthetic auxin analogs with different concentrations gradient were sprayed to the trunk apex of male and female plants of dioecious papaya. Weakening of auxin transport by 10 mg/L NPA treatment resulted in female fertility restoration in male flowers, while female flowers did not show changes. NPA treatment with higher concentration (30 and 50 mg/L) caused deformed flowers in both male and female plants. We hypothesize that the occurrence of rudimentary gynoecium patterning might associate with auxin homeostasis alteration. Proper auxin concentration and auxin homeostasis might be crucial for functional gynoecium morphogenesis in papaya flowers. -
BIOLOGY ASSIGNMENT CLASS IX UNIT II- FLOWERING PLANT Chap 3- the Flower
BIOLOGY ASSIGNMENT CLASS IX UNIT II- FLOWERING PLANT Chap 3- The Flower Flower is the most attractive,brightly coloured and significant part of plant. It help in sexual reproduction and later forms fruit and seeds. Characterstic of flower : The flower possesss the following characterstic – 1. Flower is the highly modified and specialized shoot meant for Sexual Reproduction. 2.The nodes and internodes are highly condensed to form a flat thalamus or receptacles. 3. Thalamus is quiet short and usually borne at the end of the stalk called Pedicel. 4.The flower arise from the leaf like structure called bract. 5. The floral part of flower are borne on thalamus in the form of four whorls. 6. These four whorl includes from outer side – Calyx ( sepal ), Corolla( petal ) , Androecium (stamen ), Gynoecium ( carpel ). Parts of a Flower : Floweris attached with the stem or its branch with the help of a stalk called Pedicel. If the flower is without pedicel it is called sessile . The pedicel arises from the axil of green leaf like structure called Bract.The other swollen end of the pedicel is Thalamus or torus . On the thalamus usually four whorl of Floral structure are present These four whorl from outside to inside are : i. Calyx iii. Androecium ii. Corolla iv. Gynoecium I. CALYX It is the outermost and the lowermost whorl , green in colour which cover an unopened bud .The single unit of Calyx is called sepal .It may be fussed or free from each other. Additional floral whorl outside the Calyx is called Epicalyx. Function of Calyx : i. -
Adirectionalcline in Mouriri Guianensis (Me Lastom at Ace Ae)
ADIRECTIONALCLINE IN MOURIRI GUIANENSIS (ME LASTOM AT ACE AE) Thomas Morley ( ;t) Abstract of specialization of the most important variable, the ovary, can be clearly identi Morphological variation in Mouriri guia- fied. The overall pattern of distribution nensis is described and analyzed throughout its range in Brazil and adjacent regions. Featu was briefly described previously (Morley, res that vary are ovary size, locule and ovule 1975, 1976); the present paper is a detai number, shape and smoothness of the leaf blade led report. and petiole length. The largest ovaries with the most ovules occur in west central Amazonia; intermediate sizes and numbers are widespread MATERIAL AND METHODS but reach the coast only between Marajó and Ceará; and the smallest ovaries with the fewest The study was carried out with locules and ovules are coastal or nearcoastal from Delta Amacuro in Venezuela to Marajó. pressed specimens borrowed from many Small ovaries also occur in coastal Alagoas and herbaria, to whose curators I am grateful. at Rio de Janeiro. Ovaries with the fewest locu The most instructive characters are those les and ovules are believed to be the most of the unripened ovary, and therefoie specialized, the result of evolution toward only flowering material was of value in decreased waste of ovules, since the fruits of all members are few-seeded. Leaf characters most cases. It was necessary that speci correlate statistically with ovule numbers. mens have a considerable excess of flo Possible origen of the distribution pattern of wers for the dissections to be made wi the species is compared in terms of present thout harm but fortunately only a few rainfall patterns and in terms of Pleistocene climatic change with associated forest refuges. -
Parts of a Plant Packet - Parts of a Plant Notes - Parts of a Plant Notes Key - Parts of a Plant Labeling Practice
Parts of a Plant Packet - Parts of a Plant Notes - Parts of a Plant Notes Key - Parts of a Plant Labeling Practice Includes Vocabulary: Stigma Stamen Leaf Style Petal Stoma Ovary Receptacle Cuticle Ovule Sepal Shoot System Pistil Xylem Root Hairs Anther Phloem Roots Filament Stem Root System Parts of a Plant Notes 18 14 13 (inside; for food) 15 12 (inside; for water) 16, these are 19 massively out of proportion… 21 17, covering 20 Picture modified from http://www.urbanext.uiuc.edu/gpe/index.html 1. __________- sticky part of the pistil that pollen sticks to 2. __________-long outgrowth of the ovary that collects pollen from the stamens 3. __________- base part of the pistil that holds the ovules 4. __________- unfertilized seed of the plant 5. __________- female part of the flower that contains the stigma, style, ovary and ovules. 6. __________- part of the flower that holds the pollen 7. __________- long thread-like part of the flower that holds the anthers out so insects can get to the pollen. 8. __________- male part of the flower that contains the anther and the filament. 9. __________- colorful part of the flower that protects the flower and attracts insects and other pollinators. 10. __________- stalk that bears the flower parts 11. __________- part that covers the outside of a flower bud to protect the flower before it opens 12. _________- transports water. 13. _________- transports food 14. _________- transport and support for the plant. 15. _________- cells of this perform photosynthesis. 16. _________-holes in the leaf which allow CO2 in and O2 and H2O out. -
Pollination of Cultivated Plants in the Tropics 111 Rrun.-Co Lcfcnow!Cdgmencle
ISSN 1010-1365 0 AGRICULTURAL Pollination of SERVICES cultivated plants BUL IN in the tropics 118 Food and Agriculture Organization of the United Nations FAO 6-lina AGRICULTUTZ4U. ionof SERNES cultivated plans in tetropics Edited by David W. Roubik Smithsonian Tropical Research Institute Balboa, Panama Food and Agriculture Organization of the United Nations F'Ø Rome, 1995 The designations employed and the presentation of material in this publication do not imply the expression of any opinion whatsoever on the part of the Food and Agriculture Organization of the United Nations concerning the legal status of any country, territory, city or area or of its authorities, or concerning the delimitation of its frontiers or boundaries. M-11 ISBN 92-5-103659-4 All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted in any form or by any means, electronic, mechanical, photocopying or otherwise, without the prior permission of the copyright owner. Applications for such permission, with a statement of the purpose and extent of the reproduction, should be addressed to the Director, Publications Division, Food and Agriculture Organization of the United Nations, Viale delle Terme di Caracalla, 00100 Rome, Italy. FAO 1995 PlELi. uion are ted PlauAr David W. Roubilli (edita Footli-anal ISgt-iieulture Organization of the Untled Nations Contributors Marco Accorti Makhdzir Mardan Istituto Sperimentale per la Zoologia Agraria Universiti Pertanian Malaysia Cascine del Ricci° Malaysian Bee Research Development Team 50125 Firenze, Italy 43400 Serdang, Selangor, Malaysia Stephen L. Buchmann John K. S. Mbaya United States Department of Agriculture National Beekeeping Station Carl Hayden Bee Research Center P. -
Evolution of the Suctorial Proboscis in Pollen Wasps (Masarinae, Vespidae)
Arthropod Structure & Development 31 (2002) 103–120 www.elsevier.com/locate/asd Evolution of the suctorial proboscis in pollen wasps (Masarinae, Vespidae) Harald W. Krenna,*, Volker Maussb, John Planta aInstitut fu¨r Zoologie, Universita¨t Wien, Althanstraße 14, A-1090, Vienna, Austria bStaatliches Museum fu¨r Naturkunde, Abt. Entomologie, Rosenstein 1, D-70191 Stuttgart, Germany Received 7 May 2002; accepted 17 July 2002 Abstract The morphology and functional anatomy of the mouthparts of pollen wasps (Masarinae, Hymenoptera) are examined by dissection, light microscopy and scanning electron microscopy, supplemented by field observations of flower visiting behavior. This paper focuses on the evolution of the long suctorial proboscis in pollen wasps, which is formed by the glossa, in context with nectar feeding from narrow and deep corolla of flowers. Morphological innovations are described for flower visiting insects, in particular for Masarinae, that are crucial for the production of a long proboscis such as the formation of a closed, air-tight food tube, specializations in the apical intake region, modification of the basal articulation of the glossa, and novel means of retraction, extension and storage of the elongated parts. A cladistic analysis provides a framework to reconstruct the general pathways of proboscis evolution in pollen wasps. The elongation of the proboscis in context with nectar and pollen feeding is discussed for aculeate Hymenoptera. q 2002 Elsevier Science Ltd. All rights reserved. Keywords: Mouthparts; Flower visiting; Functional anatomy; Morphological innovation; Evolution; Cladistics; Hymenoptera 1. Introduction Some have very long proboscides; however, in contrast to bees, the proboscis is formed only by the glossa and, in Evolution of elongate suctorial mouthparts have some species, it is looped back into the prementum when in occurred separately in several lineages of Hymenoptera in repose (Bradley, 1922; Schremmer, 1961; Richards, 1962; association with uptake of floral nectar. -
Functional Analyses of Genetic Pathways Controlling Petal Specification in Poppy Sinéad Drea1, Lena C
RESEARCH ARTICLE 4157 Development 134, 4157-4166 (2007) doi:10.1242/dev.013136 Functional analyses of genetic pathways controlling petal specification in poppy Sinéad Drea1, Lena C. Hileman1,*, Gemma de Martino1 and Vivian F. Irish1,2,† MADS-box genes are crucial regulators of floral development, yet how their functions have evolved to control different aspects of floral patterning is unclear. To understand the extent to which MADS-box gene functions are conserved or have diversified in different angiosperm lineages, we have exploited the capability for functional analyses in a new model system, Papaver somniferum (opium poppy). P. somniferum is a member of the order Ranunculales, and so represents a clade that is evolutionarily distant from those containing traditional model systems such as Arabidopsis, Petunia, maize or rice. We have identified and characterized the roles of several candidate MADS-box genes in petal specification in poppy. In Arabidopsis, the APETALA3 (AP3) MADS-box gene is required for both petal and stamen identity specification. By contrast, we show that the AP3 lineage has undergone gene duplication and subfunctionalization in poppy, with one gene copy required for petal development and the other responsible for stamen development. These differences in gene function are due to differences both in expression patterns and co- factor interactions. Furthermore, the genetic hierarchy controlling petal development in poppy has diverged as compared with that of Arabidopsis. As these are the first functional analyses of AP3 genes in this evolutionarily divergent clade, our results provide new information on the similarities and differences in petal developmental programs across angiosperms. Based on these observations, we discuss a model for how the petal developmental program has evolved.