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The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
Tiago Rodrigues Simões
Diapsid Phylogeny and the Origin and Early Evolution of Squamates by Tiago Rodrigues Simões A thesis submitted in partial fulfillment of the requirements for the degree of Doctor of Philosophy in SYSTEMATICS AND EVOLUTION Department of Biological Sciences University of Alberta © Tiago Rodrigues Simões, 2018 ABSTRACT Squamate reptiles comprise over 10,000 living species and hundreds of fossil species of lizards, snakes and amphisbaenians, with their origins dating back at least as far back as the Middle Jurassic. Despite this enormous diversity and a long evolutionary history, numerous fundamental questions remain to be answered regarding the early evolution and origin of this major clade of tetrapods. Such long-standing issues include identifying the oldest fossil squamate, when exactly did squamates originate, and why morphological and molecular analyses of squamate evolution have strong disagreements on fundamental aspects of the squamate tree of life. Additionally, despite much debate, there is no existing consensus over the composition of the Lepidosauromorpha (the clade that includes squamates and their sister taxon, the Rhynchocephalia), making the squamate origin problem part of a broader and more complex reptile phylogeny issue. In this thesis, I provide a series of taxonomic, phylogenetic, biogeographic and morpho-functional contributions to shed light on these problems. I describe a new taxon that overwhelms previous hypothesis of iguanian biogeography and evolution in Gondwana (Gueragama sulamericana). I re-describe and assess the functional morphology of some of the oldest known articulated lizards in the world (Eichstaettisaurus schroederi and Ardeosaurus digitatellus), providing clues to the ancestry of geckoes, and the early evolution of their scansorial behaviour. -
Stratigraphy, Sedimentary Facies and Diagenesis of the Ecca Group, Karoo Supergroup in the Eastern Cape, South Africa
STRATIGRAPHY, SEDIMENTARY FACIES AND DIAGENESIS OF THE ECCA GROUP, KAROO SUPERGROUP IN THE EASTERN CAPE, SOUTH AFRICA By Nonhlanhla Nyathi Dissertation submitted in fulfilment of the requirements for the degree of Master of Science In Geology FACULTY OF SCIENCE AND AGRICULTURE UNIVERSITY OF FORT HARE SUPERVISOR: PROFESSOR K. LIU CO-SUPERVISOR: PROFESSOR O. GWAVAVA APRIL 2014 DECLARATION I, Nonhlanhla Nyathi, hereby declare that the research described in this dissertation was carried out in the field and under the auspices of the Department of Geology, University of Fort Hare, under the supervision of Professor K. Liu and Prof. O. Gwavava. This dissertation and the accompanying photographs represent original work by the author, and have not been submitted, in whole or in part, to any other university for the purpose of a higher degree. Where reference has been made to the work of others, it has been dully acknowledged in the text. N. NYATHI Date signed: 12/04/2014 Place signed: Alice ACKNOWLEDGEMENTS The author is indebted to Professor K. Liu for his guidance, knowledge on all aspects of the project, constant supervision and help in doing field wok. Professor O. Gwavava is much appreciated for being my co-supervisor and helping me obtain financial support from the Goven Mbeki Research Development Centre. I am deeply grateful for the emotional support and encouragement from my parents and family. I express my profound gratitude to Mr Edwin Mutototwa and Mr Eric Madifor always taking time to read through my dissertation. The Rhodes University Geology laboratory technicians are thanked for their assistance in the making of thin sections. -
The Balfour Formation of the Karoo Basin, South Africa
Sedimentary Geology 140 (2001) 291±313 www.elsevier.nl/locate/sedgeo Tectonic control on ¯uvial styles: the Balfour Formation of the Karoo Basin, South Africa Octavian Catuneanua,*, Henry N. Elangob aDepartment of Earth and Atmospheric Sciences, University of Alberta, 1-26 Earth Sciences Building, Edmonton, Alta., Canada T6G 2E3 bDepartment of Geology, Rhodes University, Grahamstown 6140, South Africa Received 22 May 2000; accepted 8 December 2000 Abstract The Balfour Formation represents a fully ¯uvial succession of late Late Permian±earliest Triassic age which accumulated in the foredeep of the Karoo Basin during the over®lled phase of the foreland system. The lack of a coeval marine environment within the limits of the preserved Karoo Basin provides an opportunity to study the stratigraphic cyclicity developed during a time when accommodation was solely controlled by tectonics. The Balfour stratigraphy is composed of a succession of six third-order ¯uvial depositional sequences separated by subaerial unconformities. They formed in isolation from eustatic in¯uences, with a timing controlled by orogenic cycles of loading and unloading. Sediment accumulation took place during stages of ¯exural subsidence, whereas the bounding surfaces are related to stages of isostatic uplift. The vertical pro®le of all sequences displays an overall ®ning-upward trend related to the gradual decrease in topographic slope during orogenic loading. At the same time, an upward change in ¯uvial styles can be observed within each sequence, from initial higher to ®nal lower energy systems. The actual ¯uvial styles in each location depend on paleoslope gradients and the position of the stratigraphic section relative to the orogenic front. -
Subject Index
Cambridge University Press 0521811724 - The Evolution and Extinction of the Dinosaurs, Second Edition David E. Fastovsky and David B. Weishampel Index More information Subject index Bold type indicates figures. Absolute age 23–5 Arctic dinosaurs 372–373, 373 Actinopterygii 67 Asteroid impact 425, 426–32 Aguja Formation (Upper Cretaceous, USA) 401 see also Cretaceous–Tertiary boundary; Alxa Desert (China) 297 Iridium; Chicxulub Amarga Canyon (Argentina) 262 Affects of 432–4 American Museum of Natural History (USA) Indicators of 426, 427–9, 428, 429, 430, 431 18n, 19, 253, 259, 292 Atlas Mountains (Morocco) 263 Amnion 77 Auca Mahuevo (Upper Cretaceous, Argentina) Amniota 245, 246 Diagnostic characters 77, 78 Aves, see Theropoda Major groups 78 Azendoh (Morocco) 261 Amphibia 77, 77n, 393 Amur River (Sino-Russian border) 217 Baharije Oasis (Egypt) 291, 292 Anapsida 78, 79–80 Barun Goyot Formation (Upper Cretaceous, Anhui Province (China) 162 Mongolia) 401 Ankylosauria Bernissart (Belgium) 212, 213 Age 133, 396, 397 Biological classification 68 Armament 133, 137, 138, 139 Biostratigraphy 22, 27–8 Behavior 134–7, 138–9 Birds, see Theropoda Clades of 133, 139–42 Body plans 65 Cladogram of, see Cladograms Bone histology 362–7, 363 Derived characters of 140, 139–41, 142 British Museum (Natural History) 234, 258, 336 Distribution 134, 135 Bug Creek (USA) 441 Evolution of 139 Burgess Shale (Middle Cambrian, Canada) 64 Fermentation in 136, 137 Burial 6, 7, 8, 9, 10 History of discovery 142–5 Inferred intelligence 361 Canadian Dinosaur Rush 182, 183, -
Stable Isotopes
Stable isotopes (δ18O and δ13C) give new perspective on the ecology and diet of Endothiodon bathystoma (Therapsida, Dicynodontia) from the late Permian of the South African Karoo Basin Kévin Rey, Michael Day, Romain Amiot, François Fourel, Julie Luyt, Christophe Lécuyer, Bruce Rubidge To cite this version: Kévin Rey, Michael Day, Romain Amiot, François Fourel, Julie Luyt, et al.. Stable isotopes (δ18O and δ13C) give new perspective on the ecology and diet of Endothiodon bathystoma (Therapsida, Di- cynodontia) from the late Permian of the South African Karoo Basin. Palaeogeography, Palaeoclima- tology, Palaeoecology, Elsevier, 2020, 556, pp.109882. 10.1016/j.palaeo.2020.109882. hal-02991771 HAL Id: hal-02991771 https://hal.archives-ouvertes.fr/hal-02991771 Submitted on 17 Nov 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. Journal Pre-proof Stable isotopes (δ18O and δ13C) give new perspective on the ecology and diet of Endothiodon bathystoma (Therapsida, Dicynodontia) from the late Permian of the South African Karoo Basin Kévin Rey, Michael O. Day, Romain Amiot, François Fourel, Julie Luyt, Christophe Lécuyer, Bruce S. Rubidge PII: S0031-0182(20)30327-8 DOI: https://doi.org/10.1016/j.palaeo.2020.109882 Reference: PALAEO 109882 To appear in: Palaeogeography, Palaeoclimatology, Palaeoecology Received date: 9 April 2020 Revised date: 18 June 2020 Accepted date: 19 June 2020 Please cite this article as: K. -
A Study of the Southwestern Karoo Basin in South Africa Using Magnetic and Gravity Data
A STUDY OF THE SOUTHWESTERN KAROO BASIN IN SOUTH AFRICA USING MAGNETIC AND GRAVITY DATA By ZUSAKHE NXANTSIYA A dissertation submitted in fulfilment of the requirements for the degree of MASTER OF SCIENCE In the Department of Geology Faculty of Science and Agriculture University of Fort Hare, South Africa Supervisor: Prof O Gwavava August 2017 DECLARATION I hereby declare that this dissertation entitled “A study of the southwestern Karoo Basin in South Africa using magnetic and gravity data” is my own work except where proper referencing and acknowledgement has been made. It is being submitted for the degree of Master of Science at the University of Fort Hare, Alice and it has never been submitted before, for the purpose of any degree or examination at any other institution. August 2017 Signature Date Department of Geology Faculty of Science and Agriculture University of Fort Hare, Alice 5700 Eastern Cape, Republic of South Africa i ABSTRACT The early efforts of Booth, Johnson, Rubidge, Catuneanu, de Wit, Chevallier, Stankiewicz, Weckmann and many other scientists in studying the Karoo Supergroup has led to comprehensive documentation of the geology on the main Karoo Basin with regards to understanding the age, sedimentology, sedimentary facies and depositional environments. In spite of these studies, the subsurface structure, variations in thickness of various formations in large parts of the basin, the location and orientation of subsurface dolerite intrusions, and the depth to magnetic and gravity sources remains poorly documented. A geological study with the aid of geophysical techniques, magnetic and gravity, was conducted in the southwestern part of the main Karoo Basin. -
REVISED STRATIGRAPHY of the BEAUFORT GROUP in the SOUTHERN KAROO BASIN by B.R
87 Palaeont. afr., 24 (1981) REVISED STRATIGRAPHY OF THE BEAUFORT GROUP IN THE SOUTHERN KAROO BASIN by B.R. Turner Department of Geology, The University, Newcastle upon Tyne, NEI 7RU, England ABSTRACT The Beaufort Group in the southern Karoo Basin between Graaff-Reinet and Sutherland has been divided into three formations based primarily on the changing ratio of sandstone to mudstone. The former Abrahamskraal Formation is elevated to subgroup status and divided into two new formations, the Lootskloof Formation and the Verlatenkloof Formation, whilst the Teekloof Formation is retained but more precisely defined. The Verlatenkloof Formation includes two members, the Jakhals Valley Member and the Paalhuis Member. The Teekloof Formation includes the Oukloof Member in addition to the previously defined and described Oudeberg Sandstone Member. Stratotypes are erected for the new formations and members in accordance with the recommendations of the South African Committee for Stratigraphy. Subdivision of the formations and their relationship to the established biostratigraphy and facies patterns provides a means of fixing and correlating the most important uranium min eralised units in the succession with greater accuracy. These comprise the Paalhuis Member, the Oukloof Member and the Jakhals Valley Member, although the most important min eralised unit is the Paalhuis Member which contains up to 90 per cent of all known uranium occurrences in the Beaufort West area. CONTENTS Page INTRODUCTION . 87 STRATIGRAPHy...................... .................................................. -
Biostratigraphic Correiation in Tiie Karoo
Research Letter Biostratigraphic correiation and Eunotosaurus Page 1 of 4 Biostratigraphic correiation in tiie Karoo: The case AUTHORS: of the Middie Permian parareptiie Eunotosaurus Mike Dayi Bruce Rubidge^ The richness of fossil tetrapods from the Beaufort Group of South Africa has enabled biostratigraphic John Almond^ subdivision of this Permo-Triassic succession, with global applicability. Despite being the thickest of the seven Sifeiani Jirah^ biozones recognised, attempts at further subdivision of the Middie Permian TafJinocephalus Assemblage AFFILIATIONS: Zone (Abrahamskraal Formation) have not been successful, largely because the exact stratigraphie ranges 'Bernard Price Institute of of fossil taxa are unknown. This gap in knowledge has limited stratigraphie eorrelation of the Abrahamskraal Paiaeontoiogy, University of the Formation and hindered understanding of Middle Permian Karoo basin development. Currently, the lowermost Witwatersrand, Johannesburg, South Africa Beaufort Group is split between an eastern and a western stratigraphie scheme and, beeause of poor outerop ^Natura Viva, Cape Town, and the reiative paueity of fossiis in the east, stratigraphie eorrelation between the two areas has been South Africa uneertain. Reeent fossil diseoveries of the parareptile Eunotosaurus africafius in the Eastern Cape and Free State provinees have extended its known geographic range in ttie east. An additional speeimen from the lower CORRESPONOENCE TO: Middleton Formation in the Eastern Cape has, forthe first time, enabied the biostratigraphie eorrelation of this Mike Day unit with the Poortjie Member of the Teekloof Formation in the west. These finds eonfirm the diaehroneity of the boundary between the marine Eeea Group and the terrestrial Beaufort Group. EMAIL: [email protected]. acza Introduction POSTAL AOORESS: The taxonomic affinity of the enigmatic Permian parareptiie Eunotosaurus africanus, easily recognised in the BPi Palaeontology, fieid by its distinctive broad ribs (Figure 1), has iong intrigued paiaeontoiogists. -
1 the Proposed Relocation of Dwellers at Ingula Pumped Storage
The Proposed Relocation of Dwellers at Ingula Pumped Storage Scheme Phumelela and Maluti A - Phofung Local Municipalities, Thabo Mofutsanyana District Municipality, Free State Province Farm: Wilge Rivier 319 Palaeontological Impact Assessment: Phase 1: Field Study Tsimba Archaeological Footprints 24 Lawson Mansions, 74 Loveday Street, Johannesburg, 2000 Tel: 061 912 5118 2020/11/30 Ref: Pending Regisaurus (ESI) (H. Fourie) B. Executive summary Outline of the development project: Tsimba Archaeological Footprints (Pty) Ltd has been appointed by Myezo Environmental Management Services to undertake a Palaeontological Impact Assessment (PIA), Desktop Study 1 of the suitability of The Proposed Relocation of Dwellers at Ingula Pumped Storage Scheme, Phumelela and Maluti A - Phofung Local Municipalities, Thabo Mofutsanyana District Municipality, Free State Province on the Farm: Wilge Rivier 319. The applicant, Eskom Holdings SOC Limited (Eskom) intends to embark on a relocation programme for six families at Ingula Pumped Storage Scheme, within the Ingula Nature Reserve. It was decided to relocate the remaining six families to the Farm Wilge Rivier 319. The Project includes one locality Option (see map): Option 1: An area outlined in red located about 23 km north east of Van Reenen, approximately 10 km north of the Ingula Pumped Storage Scheme, and about 42 km north-east-east of Harrismith. The approximate size of the site is X hectares. The National Heritage Resources Act (Act No. 25 of 1999) (NHRA) requires that all heritage resources, that is, all places or objects of aesthetic, architectural, historical, scientific, social, spiritual, linguistic or technological value or significance are protected. The Republic of South Africa (RSA) has a remarkably rich fossil record that stretches back in time for some 3.5 billion years and must be protected for its scientific value. -
Project Name
BUFFALO RIVER ESTUARY SITUATION ASSESSMENT REPORT Prepared for: Department of Environmental Affairs: Oceans Nelson Mandela Metropolitan University and Coasts Prepared by: EOH Coastal & Environmental Services EAST LONDON 25 Tecoma Street, Berea East London, 5214 043 726 7809 Also in Grahamstown, Port Elizabeth, Cape Town and Maputo www.cesnet.co.za | www.eoh.co.za March 2016 EOH Coastal & Environmental Services i Buffalo River Estuary Draft Situation Assessment Report – March 2016 THE PROJECT TEAM Dr Alan Carter – Team Leader, stakeholder engagement, legislation and policy and report review Alan is an Executive at EOH CES, holds a PhD in Marine Biology and is a certified Public Accountant, with extensive training and experience in both financial accounting and environmental science disciplines with international accounting firms in South Africa and the USA. He has 15 years’ experience in environmental management and has specialist skills in sanitation, coastal environments and industrial waste. Alan is registered as a Professional Natural Scientist under the South African Council for Natural Scientific Professions (SACNASP) and is a Certified Environmental Assessment Practitioner (CEAPSA). Amy Hunter – Project Manager, stakeholder engagement and data gathering Amy is an Environmental Consultant at EOH-CES. Amy holds a B.Sc. in Biochemistry and Zoology as well as a B.Sc. Honours in Zoology, both from the University of Johannesburg. Her honours project investigated the role of a tenebrionid beetle in the ecology of Bakwena Cave, Pretoria. Her M.Sc project, through Stellenbosch University, was a study on the effects of probiotics on the physiological and biochemical development of hatchery raised dusky kob (Argyrosomus japonicus) larvae. -
Palaeontological Impact Assessment for the Proposed Construction of Three Residential Townships in Kroonstad, Free State Province
Palaeontological Impact Assessment for the proposed construction of three residential townships in Kroonstad, Free State Province Desktop Study For Archaeological and Heritage Services Africa (Pty) Ltd 07 July 2019 Prof Marion Bamford Palaeobotanist P Bag 652, WITS 2050 Johannesburg, South Africa [email protected] Expertise of Specialist The Palaeontologist Consultant is: Prof Marion Bamford Qualifications: PhD (Wits Univ, 1990); FRSSAf, ASSAf Experience: 30 years research; 22 years PIA studies Declaration of Independence This report has been compiled by Professor Marion Bamford, of the University of the Witwatersrand, sub-contracted by Archaeological and Heritage Services Africa (Pty) Ltd, South Africa. The views expressed in this report are entirely those of the author and no other interest was displayed during the decision making process for the Project. Specialist: Prof Marion Bamford Signature: 1 Executive Summary A palaeontological Impact Assessment was requested for the proposed construction of three areas of residential housing, the Maokeng Housing Development, Kroonstad. To comply with the South African Heritage Resources Agency (SAHRA) in terms of Section 38(8) of the National Heritage Resources Act, 1999 (Act No. 25 of 1999) (NHRA), a desktop Palaeontological Impact Assessment (PIA) was completed for the proposed project. The proposed sites lie on the sandstones and mudstones of the late Permian, Adelaide Subgroup, Beaufort Group, Karoo Supergroup. Although fossils have not been reported from this site there is a small chance that typical vertebrates of the Pristerognathus, Tropidostoma, Cistecephalus and Dicynodon Assemblage Zones could occur, as well as typical (but very infrequent) late Glossopteris flora plants, could occur in the sediments just below the surface.