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Investigating Sexual Dimorphism in Ceratopsid Horncores
University of Calgary PRISM: University of Calgary's Digital Repository Graduate Studies The Vault: Electronic Theses and Dissertations 2013-01-25 Investigating Sexual Dimorphism in Ceratopsid Horncores Borkovic, Benjamin Borkovic, B. (2013). Investigating Sexual Dimorphism in Ceratopsid Horncores (Unpublished master's thesis). University of Calgary, Calgary, AB. doi:10.11575/PRISM/26635 http://hdl.handle.net/11023/498 master thesis University of Calgary graduate students retain copyright ownership and moral rights for their thesis. You may use this material in any way that is permitted by the Copyright Act or through licensing that has been assigned to the document. For uses that are not allowable under copyright legislation or licensing, you are required to seek permission. Downloaded from PRISM: https://prism.ucalgary.ca UNIVERSITY OF CALGARY Investigating Sexual Dimorphism in Ceratopsid Horncores by Benjamin Borkovic A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE DEPARTMENT OF BIOLOGICAL SCIENCES CALGARY, ALBERTA JANUARY, 2013 © Benjamin Borkovic 2013 Abstract Evidence for sexual dimorphism was investigated in the horncores of two ceratopsid dinosaurs, Triceratops and Centrosaurus apertus. A review of studies of sexual dimorphism in the vertebrate fossil record revealed methods that were selected for use in ceratopsids. Mountain goats, bison, and pronghorn were selected as exemplar taxa for a proof of principle study that tested the selected methods, and informed and guided the investigation of sexual dimorphism in dinosaurs. Skulls of these exemplar taxa were measured in museum collections, and methods of analysing morphological variation were tested for their ability to demonstrate sexual dimorphism in their horns and horncores. -
Mammal and Plant Localities of the Fort Union, Willwood, and Iktman Formations, Southern Bighorn Basin* Wyoming
Distribution and Stratigraphip Correlation of Upper:UB_ • Ju Paleocene and Lower Eocene Fossil Mammal and Plant Localities of the Fort Union, Willwood, and Iktman Formations, Southern Bighorn Basin* Wyoming U,S. GEOLOGICAL SURVEY PROFESS IONAL PAPER 1540 Cover. A member of the American Museum of Natural History 1896 expedition enter ing the badlands of the Willwood Formation on Dorsey Creek, Wyoming, near what is now U.S. Geological Survey fossil vertebrate locality D1691 (Wardel Reservoir quadran gle). View to the southwest. Photograph by Walter Granger, courtesy of the Department of Library Services, American Museum of Natural History, New York, negative no. 35957. DISTRIBUTION AND STRATIGRAPHIC CORRELATION OF UPPER PALEOCENE AND LOWER EOCENE FOSSIL MAMMAL AND PLANT LOCALITIES OF THE FORT UNION, WILLWOOD, AND TATMAN FORMATIONS, SOUTHERN BIGHORN BASIN, WYOMING Upper part of the Will wood Formation on East Ridge, Middle Fork of Fifteenmile Creek, southern Bighorn Basin, Wyoming. The Kirwin intrusive complex of the Absaroka Range is in the background. View to the west. Distribution and Stratigraphic Correlation of Upper Paleocene and Lower Eocene Fossil Mammal and Plant Localities of the Fort Union, Willwood, and Tatman Formations, Southern Bighorn Basin, Wyoming By Thomas M. Down, Kenneth D. Rose, Elwyn L. Simons, and Scott L. Wing U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1540 UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1994 U.S. DEPARTMENT OF THE INTERIOR BRUCE BABBITT, Secretary U.S. GEOLOGICAL SURVEY Robert M. Hirsch, Acting Director For sale by U.S. Geological Survey, Map Distribution Box 25286, MS 306, Federal Center Denver, CO 80225 Any use of trade, product, or firm names in this publication is for descriptive purposes only and does not imply endorsement by the U.S. -
Defining the Mesozoic
DEBRA LINDSAY THE MESOZOIC/DEFINING DISCIPLINES: LATE NINETEENTH- CENTURY DEBATES OVER THE JURASSIC–CRETACEOUS BOUNDARY DEBRA LINDSAY Department of History and Politics University of New Brunswick, Saint John, NB, Canada E2L 4L5 [email protected] ABSTRACT The last two decades of the nineteenth century were exciting times in American paleontology, with disputes over Jurassic dinosaurs between Edward Drinker Cope and Othniel Charles Marsh appearing in the press. Less well known is the dispute over defining the Mesozoic that began in 1888 when Marsh invited Lester Frank Ward, a colleague with whom he had been working on the Potomac Formation for the United States Geological Survey, to speak on the plant fossils found there. Initially agreeing with Marsh that the Potomac was a Jurassic formation, work on fossil cycads led Ward to conclude that the Potomac was Lower Cretaceous. As Ward and Marsh grappled with the question of how to determine the age and identity of Mesozoic systems, they joined other paleontologists and Earth Sciences History geologists such as William J. McGee, Albert Charles Seward, and Samuel W. Williston in 2011, Vol. 30, No. 2 a debate that often reflected scientific training and sub-specializations as much as pp. 216–239 stratigraphic principles, becoming caught up in a trans-Atlantic dispute in which their reputations were on the line as they claimed that ‘their’ fossils were key determinants of Mesozoic systems. In the end, Marsh’s reputation as a paleontologist was far better established than that of Ward, who moved on to another career as a sociologist at Brown University, but cycad discoveries from Maryland, Colorado and Wyoming, and fieldwork, trumped laboratory studies—even when performed by a master systematist—as the Potomac Formation proved to be Lower Cretaceous. -
Agate Fossil Beds
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln U.S. National Park Service Publications and Papers National Park Service 1980 Agate Fossil Beds Follow this and additional works at: http://digitalcommons.unl.edu/natlpark "Agate Fossil Beds" (1980). U.S. National Park Service Publications and Papers. 160. http://digitalcommons.unl.edu/natlpark/160 This Article is brought to you for free and open access by the National Park Service at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in U.S. National Park Service Publications and Papers by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. Agate Fossil Beds cap. tfs*Af Clemson Universit A *?* jfcti *JpRPP* - - - . Agate Fossil Beds Agate Fossil Beds National Monument Nebraska Produced by the Division of Publications National Park Service U.S. Department of the Interior Washington, D.C. 1980 — — The National Park Handbook Series National Park Handbooks, compact introductions to the great natural and historic places adminis- tered by the National Park Service, are designed to promote understanding and enjoyment of the parks. Each is intended to be informative reading and a useful guide before, during, and after a park visit. More than 100 titles are in print. This is Handbook 107. You may purchase the handbooks through the mail by writing to Superintendent of Documents, U.S. Government Printing Office, Washington DC 20402. About This Book What was life like in North America 21 million years ago? Agate Fossil Beds provides a glimpse of that time, long before the arrival of man, when now-extinct creatures roamed the land which we know today as Nebraska. -
The Dinosaurs of North America
FEOM THE SIXTEENTH ANNUAL KEPOKT OF THE U. S, GEOLOGICAL SURVEY THE DINOSAURS OF NORTH AMERICA OTHNIEL CHARLES MARSH TALE UNIVERSITY WASHINGTON 1896 ^33/^, I/BRAKt 4 ,\ . THE DINOSAURS OF NORTH AMERICA. BY OTHNIEL CHARLES MARSH. 133 CONTENTS. Pajje. Introduction 143 Part I. —Triassic dinosaurs 146 Theropoda 146 Anchisaurida? 147 Anchisaurus 147 The skull 148 The fore limbs 149 The hind limbs 149 Anchisaurus solus 149 Amniosaurus rTT 150 Eestoration of Anchisaurus 150 Dinosaurian footprints 151 Distribution of Triassic dinosaurs 152 Part II. —Jurassic dinosaurs 152 Theropoda : 153 Hallopus 153 Fore and hind limbs 154 Coelurus _ 155 The vertebra:- 155 The hind limbs 156 Ceratosaurus 156 The skull 157 The brain 159 The lower iaws 159 The vertebra: 159 The scapular arch 160 The pelvic arch 160 The metatarsals 162 Eestoration of Ceratosaurus 163 Allosaurus 163 European Theropoda 163 Sauropoda 164 Atlantosaurus beds 164 Families of Sauropoda 165 Atlantosauridie 166 Atlantosaurus 166 Apatosaurus 166 The sacral cavity 166 The vertebra- 167 Brontosaurus 168 The scapular arch 168 The cervical vertebra.- 169 The dorsal vertebree 169 The sacrum 170 The caudal vertebra- 171 The pelric arch 172 The fore limbs 173 The hind limbs 173 135 136 CONTENTS. Part II. —Jurassic dinosaurs—Continued. Page. Sauropoda—Continued. Atlantosaurida? —Continued. Restoration of Brontosaurus 173 Barnsaurus 174 Diplodoeida? 175 Diplodocus 175 The skull 175 The brain 178 The lower jaws 178 The teeth 179 The vertebra; 180 The sternal bones 180 The pelvic girdle 180 Size and habits 180 Morosaurida? 181 Morosaurus 181 The skull 181 The vertebra? 181 The fore limbs 182 The pelvis 182 The hind limbs 183 Pleuroccelida? : 183 Pleurocoelus 183 The skull 183 The vertebras 183 Distribution of the Sauropoda 185 Comparison with European forms 185 Predentata ». -
Lakamie Basin, Wyoming
DEPARTMENT OF THE INTERIOR UNITED STATES GEOLOGICAL SURVEY GEORGE OTIS SMITH, DIRECTOR BULLETIN 364 GEOLOGY AND MINERAL RESOURCES OF THE LAKAMIE BASIN, WYOMING A PRELIMINARY REPORT BY N. H. DARTON AND C. E. SIEBENTHAL WASHINGTON GOVERNMENT PRINTING OFFICE 1909 CONTENTS. Page. Introduction............................................................. 7 Geography ............................................................... 8 Configuration........................................................ 8 Drainage ............................................................ 9 Climate ............................................................. 9 Temperature...................................................... 9 Precipitation..................................................... 10 Geology ................................................................. 11 Stratigraphy.......................................................... 11 General relations........................../....................... .11 Carboniferous system............................................. 13 Casper formation......................... .................... 13. General character........................................ 13 Thickness ............................................... 13 Local features............................................ 14 Erosion and weathering of limestone slopes ................ 18 Paleontology and age..................................... 19 Correlation .............................................. 20 Forelle limestone............................................ -
Body-Size Evolution in the Dinosauria
8 Body-Size Evolution in the Dinosauria Matthew T. Carrano Introduction The evolution of body size and its influence on organismal biology have received scientific attention since the earliest decades of evolutionary study (e.g., Cope, 1887, 1896; Thompson, 1917). Both paleontologists and neontologists have attempted to determine correlations between body size and numerous aspects of life history, with the ultimate goal of docu- menting both the predictive and causal connections involved (LaBarbera, 1986, 1989). These studies have generated an appreciation for the thor- oughgoing interrelationships between body size and nearly every sig- nificant facet of organismal biology, including metabolism (Lindstedt & Calder, 1981; Schmidt-Nielsen, 1984; McNab, 1989), population ecology (Damuth, 1981; Juanes, 1986; Gittleman & Purvis, 1998), locomotion (Mc- Mahon, 1975; Biewener, 1989; Alexander, 1996), and reproduction (Alex- ander, 1996). An enduring focus of these studies has been Cope’s Rule, the notion that body size tends to increase over time within lineages (Kurtén, 1953; Stanley, 1973; Polly, 1998). Such an observation has been made regarding many different clades but has been examined specifically in only a few (MacFadden, 1986; Arnold et al., 1995; Jablonski, 1996, 1997; Trammer & Kaim, 1997, 1999; Alroy, 1998). The discordant results of such analyses have underscored two points: (1) Cope’s Rule does not apply universally to all groups; and (2) even when present, size increases in different clades may reflect very different underlying processes. Thus, the question, “does Cope’s Rule exist?” is better parsed into two questions: “to which groups does Cope’s Rule apply?” and “what process is responsible for it in each?” Several recent works (McShea, 1994, 2000; Jablonski, 1997; Alroy, 1998, 2000a, 2000b) have begun to address these more specific questions, attempting to quantify patterns of body-size evolution in a phylogenetic (rather than strictly temporal) context, as well as developing methods for interpreting the resultant patterns. -
Lesothosaurus, "Fabrosaurids," and the Early Evolution of Ornithischia Author(S): Paul C
Lesothosaurus, "Fabrosaurids," and the Early Evolution of Ornithischia Author(s): Paul C. Sereno Source: Journal of Vertebrate Paleontology, Vol. 11, No. 2 (Jun. 20, 1991), pp. 168-197 Published by: Taylor & Francis, Ltd. on behalf of The Society of Vertebrate Paleontology Stable URL: https://www.jstor.org/stable/4523370 Accessed: 28-01-2019 13:42 UTC JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact [email protected]. Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at https://about.jstor.org/terms The Society of Vertebrate Paleontology, Taylor & Francis, Ltd. are collaborating with JSTOR to digitize, preserve and extend access to Journal of Vertebrate Paleontology This content downloaded from 69.80.65.67 on Mon, 28 Jan 2019 13:42:40 UTC All use subject to https://about.jstor.org/terms Journal of Vertebrate Paleontology 11(2): 168-197, June 1991 © 1991 by the Society of Vertebrate Paleontology LESOTHOSAURUS, "FABROSAURIDS," AND THE EARLY EVOLUTION OF ORNITHISCHIA PAUL C. SERENO Department of Organismal Biology and Anatomy, University of Chicago, 1025 E. 57th Street, Chicago, Illinois 60637 ABSTRACT -New materials of Lesothosaurus diagnosticus permit a detailed understanding of one of the earliest and most primitive ornithischians. Skull proportions and sutural relations can be discerned from several articulated and disarticulated skulls. The snout is proportionately long with a vascularized, horn-covered tip. -
Dinictis ROCK ROCK UNIT COLUMN PERIOD EPOCH AGES MILLIONS of YEARS AGO Common Name: Holocene Oahe .01 Saber-Tooth Cat
North Dakota Stratigraphy Dinictis ROCK ROCK UNIT COLUMN PERIOD EPOCH AGES MILLIONS OF YEARS AGO Common Name: Holocene Oahe .01 Saber-tooth cat Coleharbor Pleistocene QUATERNARY Classification: 1.8 Pliocene Unnamed 5 Miocene Class: Mammalia 25 Arikaree Order: Carnivora Family: Felidae Brule Oligocene 38 South Heart Skull of the saber-toothed cat, Dinictis. The black arrows point Chadron Chalky Buttes to tooth puncture marks in the skull indicating that another animal Camels Butte Eocene Golden killed this cat. The saber teeth were not found. This skull is on 55 Valley Bear Den exhibit at the North Dakota Heritage Center, Bismarck. Brule Sentinel Formation. Stark County. Width 174 mm. North Dakota State Butte TERTIARY Fossil Collection ND1.1. Description: Bullion Dinictis was one of the early saber-toothed cats. Its body was Paleocene Creek sleek, 3 to 4 feet long, and resembled the modern-day lynx. They lived in North Dakota about 30 million years ago and were much Slope smaller than the huge saber-toothed cat called Smilodon that lived Cannonball during the Ice Age, a few thousand years ago. Like other saber- Ludlow 65 toothed cats, Dinictis upper canine teeth were elongated, serrated, Hell Creek and curved. They were extremely efficient predators, and the teeth were used in a stabbing action into the skin and flesh of the throat Fox Hills and lower neck of its prey. These teeth were also used to slice into muscle after the initial stab of the teeth killed the victim. Their lower jaws were also modified to open to a wide gape and they ACEOUS Pierre possessed very strong neck muscles. -
Othniel Charles Marsh
NATIONAL ACADEMY OF SCIENCES OF THE UNITED STATES OF AMERICA BIOGRAPHICAL MEMOIRS VOLUME XX-FIRST MEMOIR BIOGRAPHICAL hlEhlOIR OE OTHNIEL CHARLES MARSH BY CHARLES SCHUCHEKT PRESENTED TO THE -4C.4DEhlY AT THE ASSU.\L JIEETIPU'G, 1938 OTHNIEL CHARLES MARSH BY CHARLES SCHUCIIERT Othniel Charles Marsh, for ,twelve years president of the National r\caderny of Sciences, was born to Caleb Marsh and Mary Gaines Peabody on October 29, 1831, in Lockport, New York, and died in New Haven, Connecticut, on March 18, 1899. One of the three founders of the science of vertebrate paleon- tology in America, his career furnishes an outstanding exatnple of the indomitable spirit that drives men on to a determined goal. His motto might well have been. "\l.That 1 have, I hold." He asked no quarter, and gave none. :It home around a camp fire or in an army tent, formal as a presiding officer or in society, at times austere and autocratic, at others a raconteur of note, he left a lasting impression on his chosen 111-anch of science. Summarizing his work statistically, it may be said that he- tween 1861 and 1899 he published about 300 papers, reports, and books. Of new genera he described 225, and of new species, 496; of new families 64, of su1)orders 8, of orders 19, and of subclasses I. Of his work on vertebrate fossils in general, Osbom says that he "carried out the most intensive field esl)loration known to science ant1 pul~lishetl a large num1)er of 1)reliminary papcrs, which fairly revol~~tionizedour knowledge." ANCESTRY AKD TIZ:\INISG John hlarsh of Salem, the first of his name recorded as emigrating from England to America, is believctl to have reached In the preparation of this memorial, the writer has been aided greatly by the excellent skctches of Professor Marsh writtcn by George Bird Grinnell, Charles E. -
Aplodontid, Sciurid, Castorid, Zapodid and Geomyoid Rodents of the Rodent Hill Locality, Cypress Hills Formation, Southwest Saskatchewan
APLODONTID, SCIURID, CASTORID, ZAPODID AND GEOMYOID RODENTS OF THE RODENT HILL LOCALITY, CYPRESS HILLS FORMATION, SOUTHWEST SASKATCHEWAN A Thesis Submitted to the College of Graduate Studies and Research in Partial Fulfillment of the Requirements for the Degree of Master of Science in the Department of Geological Sciences University of Saskatchewan Saskatoon By Sean D. Bell © Copyright Sean D. Bell, December 2004. All rights reserved. PERMISSION TO USE In presenting this thesis in partial fulfilment of the requirements for a Master’s degree from the University of Saskatchewan, I agree that the libraries of the University of Saskatchewan may make it freely available for inspection. I further agree that permission for copying of this thesis in any manner, in whole or in part, for scholarly purposes may be granted by the professors who supervised my thesis work or, in their absence, by the Head of the Department of Geological Sciences or the Dean of the College of Graduate Studies and Research. It is understood that any copying or publication or use of this thesis or parts thereof for financial gain shall not be allowed without my written permission. It is also understood that due recognition shall be given to me and to the University of Saskatchewan in any scholarly use which may be made of any material in my thesis. Requests for permission to copy or to make other use of material in this thesis in whole or part should be addressed to: Head of the Department of Geological Sciences 114 Science Place University of Saskatchewan Saskatoon, Saskatchewan S7N 5E2 i ABSTRACT The Rodent Hill Locality is a fossil-bearing site that is part of the Cypress Hills Formation, and is located roughly 15 km northwest of the town of Eastend, Saskatchewan. -
Article XXXVIII. -NOTE on SOME WORM (?) BUR- ROWS in ROCKS of the CHEMUNG GROUP of NEW YORK
Article XXXVIII. -NOTE ON SOME WORM (?) BUR- ROWS IN ROCKS OF THE CHEMUNG GROUP OF NEW YORK. By R. P. WHITFIELD. Arenicolites chemungensis, sp. nov. PLATE XIV, FIGS. I AND 2. While working up the fossils of the Potsdam sandstone for the Wisconsin Report, in I876 and I877, there came into my hands a number of specimens representing the so-called Scolithus, which I described as Arenicolites woodi in Volume IV of Prof. T. C. Chamberlain's Report of I882. One block of that series showed the original surface of the mud-covered sandstone with the burrows of the worm (?) which made the perforations, together with the little hillocks surrounding the outlet of the burrows, just as the animal built them up by its castings during life; proving pretty conclusively that it must have been a marine worm-like animal which caused the perforations. Among the geological specimens of the Chemung Group in the Museum, from near Bath, Steuben Co., New York, I find an example so nearly resembling that figured in the Wis- consin Report above referred to, that there can be no ques- tion as to the similarity of its origin. On this Chemung specimen the hillocks are somewhat larger and the funnels more distinct, being generally 5 or 6 mm. in diameter, and the walls surrounding them about 2 mm. thick, while some of the hillocks are much larger and higher and appear to have collapsed from the semifluidity of the sand, closing up the top of the burrow so as to show a mere slit in its place.