Phylogeny of Early Tertiary Carnivora, with a Description of a New Species of Protictis from the Middle Eocene of Northwestern Wyoming
AMERICAN MUSEUM Nov itates PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK, N.Y. 10024 Number 2725, pp. 1-64, figs. 1-9, tables 1, 2 April 15, 1982 Phylogeny of Early Tertiary Carnivora, With a Description of a New Species of Protictis From the Middle Eocene of Northwestern Wyoming JOHN J. FLYNN1 AND HENRY GALIANO2 ABSTRACT Since Cope erected the Miacidae in 1880, all for the suprageneric taxa Carnivora, Feliformia, early Tertiary carnivorans have generally been Caniformia, Didymictidae (Didymictida) (new), grouped in this family because of their overall Aeluroida (new), Feloidea, and Viverravidae. primitive morphology. A historical review of the Protictis, including the subgenera P. (Protictis), classifications and definitions of the Miacidae is P. (Bryanictis), and P. (Protictoides) (new), is provided. Discovery of a new subgenus of Pro- diagnosed and discussed. Simpsonictis is consid- tictis from the Uintan (mid Eocene) of north- ered a valid genus and is allied with Viverravus western Wyoming led to a reevaluation of the in the Viverravidae (Aeluroida, Feliformia). Pa- phylogeny of the Carnivora. The Miacidae (Mia- laeogale is removed from the Mustelidae (Cani- coidea), as previously constructed, is a paraphy- formia) and is considered a close relative of the letic assemblage of all the early Tertiary Carniv- Viverravidae. Ictidopappus is placed incertae ora. Phylogenetic analysis of the distribution of sedis within the Aeluroida. "Plesiomiacis" and cranial, dental, and postcranial morphology indi- Quercygale, which were previously considered cates that the bipartite division recognized in liv- members of the "Viverravinae," are considered ing Carnivora may be traced to the earliest Ter- members of the Caniformia.
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