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Conch Structures, Soft-Tissue Imprints and Taphonomy of the Middle Ordovician Cephalopod Tragoceras Falcatum from Estonia
FOSSIL IMPRINT • vol. 75 • 2019 • no. 1 • pp. 70–78 (formerly ACTA MUSEI NATIONALIS PRAGAE, Series B – Historia Naturalis) CONCH STRUCTURES, SOFT-TISSUE IMPRINTS AND TAPHONOMY OF THE MIDDLE ORDOVICIAN CEPHALOPOD TRAGOCERAS FALCATUM FROM ESTONIA ALEXANDER POHLE1,*, CHRISTIAN KLUG1, URSULA TOOM2, BJÖRN KRÖGER3 1 Paläontologisches Institut und Museum, Universität Zürich, Karl-Schmid-Strasse 4, 8006 Zürich, Switzerland; e-mail: [email protected], [email protected]. 2 Department of Geology, Tallinn University of Technology, Ehitajate tee 5, 19086 Tallinn, Estonia; e-mail: [email protected]. 3 Finnish Museum of Natural History, P.O. Box 44 (Jyrängöntie 2), 00014 University of Helsinki, Finland; e-mail: [email protected]. * corresponding author Pohle, A., Klug, C., Toom, U., Kröger, B. (2019): Conch structures, soft-tissue imprints and taphonomy of the Middle Ordovician cephalopod Tragoceras falcatum from Estonia. – Fossil Imprint, 75(1): 70–78, Praha. ISSN 2533-4050 (print), ISSN 2533-4069 (on-line). Abstract: Tragoceras falcatum (SCHLOTHEIM, 1820) is a common, loosely coiled estonioceratid (Tarphycerida, Cephalopoda) occurring in the Kunda Regional Stage (early Darriwilian, Middle Ordovician) of Estonia. Although the species is quite well- known, we document some features for the first time. For example, one specimen from the Harku quarry (Estonia) with a phosphatized replacement shell exhibits growth halts (megastriae) on the body chamber. As they are not preserved in smaller specimens, we suggest that these megastriae formed at the approach of maturity, possibly also reflecting sexual dimorphism and cycles of reproduction (iteroparity?). Additionally, the specimen shows minute soft-tissue imprints (drag bands and pseudosu- tures). These imprints are comparable to patterns in other cephalopods such as ammonoids, bactritids and other nautiloids, but have not yet been reported from Palaeozoic nautiloids. -
Main Morphological Events in the Evolution of Paleozoic Cephalopods I
Stratigraphy and Geological Correlation, Vol. 2, No. 1, 1994, pp. 49 - 55. Translated from Stratigrafiya. Geologicheskaya Korrelyatsiya, Vol. 2, No. 1,1994, pp. 55 - 61. Original Russian Text Copyright © 1994 by Barskov, Bogoslovskaya, Zhuravleva, Kiselev, Kuzina, Leonova, Shimanskii, Yatskov. English Translation Copyright © 1994 by Interperiodica Publishing (Russia). Main Morphological Events in the Evolution of Paleozoic Cephalopods I. S. Barskov*, M. F. Bogoslovskaya*, F. A. Zhuravleva*, G. N. Kiselev**, L. F. Kuzina*, T. B. Leonova*, V. N. Shimanskii*, and S. V. Yatskov* institute of Paleontology, Russian Academy of Sciences, Profsoyuznaya ul. 123, Moscow, 117647 Russia **Department of Paleontology, St. Petersburg State University, 16-ya Liniya 29, St. Petersburg, 199178 Russia Received January 26,1993 Abstract - New morphological features in shell structure, which were the starting points of cephalopod diver sification into taxa of high ranks (subclasses, orders, superfamilies), are considered as phylogenetic events. Main morphological innovations in the evolution of nautiloid cephalopods; the formation of endosiphuncular and cameral deposits, shell coiling, truncation of the phragmocone’s apical end, and contracted aperture, which originated to make the relative shell positioning in the water more efficient. Changes in the lobe line structure and various types of complications in the primary lobes (ventral, umbonal, and lateral) were the most important morphological innovations in convolute ammonoids. The functional significance of these changes remains unclear, but recognition of equally significant changes in primary lobes requires a review of the Paleozoic Ammonoidea taxonomy at the suborder level. This paper is a continuation of a study whose first lation of the buoyancy process and to support in various results have been published (Barskov et al., 1993). -
Cephalopod Reproductive Strategies Derived from Embryonic Shell Size
Biol. Rev. (2017), pp. 000–000. 1 doi: 10.1111/brv.12341 Cephalopod embryonic shells as a tool to reconstruct reproductive strategies in extinct taxa Vladimir Laptikhovsky1,∗, Svetlana Nikolaeva2,3,4 and Mikhail Rogov5 1Fisheries Division, Cefas, Lowestoft, NR33 0HT, U.K. 2Department of Earth Sciences Natural History Museum, London, SW7 5BD, U.K. 3Laboratory of Molluscs Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117997, Russia 4Laboratory of Stratigraphy of Oil and Gas Bearing Reservoirs Kazan Federal University, Kazan, 420000, Russia 5Department of Stratigraphy Geological Institute, Russian Academy of Sciences, Moscow, 119017, Russia ABSTRACT An exhaustive study of existing data on the relationship between egg size and maximum size of embryonic shells in 42 species of extant cephalopods demonstrated that these values are approximately equal regardless of taxonomy and shell morphology. Egg size is also approximately equal to mantle length of hatchlings in 45 cephalopod species with rudimentary shells. Paired data on the size of the initial chamber versus embryonic shell in 235 species of Ammonoidea, 46 Bactritida, 13 Nautilida, 22 Orthocerida, 8 Tarphycerida, 4 Oncocerida, 1 Belemnoidea, 4 Sepiida and 1 Spirulida demonstrated that, although there is a positive relationship between these parameters in some taxa, initial chamber size cannot be used to predict egg size in extinct cephalopods; the size of the embryonic shell may be more appropriate for this task. The evolution of reproductive strategies in cephalopods in the geological past was marked by an increasing significance of small-egged taxa, as is also seen in simultaneously evolving fish taxa. Key words: embryonic shell, initial chamber, hatchling, egg size, Cephalopoda, Ammonoidea, reproductive strategy, Nautilida, Coleoidea. -
JPA Kroeger FINAL DRAFT
1 First record of a non-paleotropical intejocerid cephalopod from 2 Darriwilian (Middle Ordovician) strata of central Spain 3 4 Björn Kröger1 and Juan Carlos Gutiérrez-Marco2 5 6 1 Finnish Museum of Natural History, PO Box 44, Fi-00014 University of Helsinki, Finland, 7 <bjorn. [email protected]> 8 2 Instituto de Geociencias (CSIC, UCM), and Departamento de Geodinámica, Estratigrafía y 9 Paleontología, Facultad CC. Geológicas, José Antonio Novais 12, 28040 Madrid, Spain. 10 <[email protected]> 11 12 Running Header: peri-Gondwana intejocerid 13 14 Abstract.—The order Intejocerida is an enigmatic, short lived cephalopod taxon, known 15 previously only from Early–Middle Ordovician beds of Siberia and the USA. Here, we 16 report a new genus, Cabaneroceras n. gen., and a new species of C. aznari n. sp. from 17 Middle Ordovician strata of central Spain. This finding widens the paleogeographic range of 18 the order toward high-paleolatitudinal areas of peri-Gondwana. A curved conch, 19 characteristic for the new genus, was previously unknown from members of the Intejocerida. 20 21 UUID: http://zoobank.org/21f0a09c-5265-4d29-824b-6b105d36b791 22 23 1 24 Introduction 25 26 Intejocerids are a unique group of cephalopods, previously known only from Siberia (Balashov 27 1960, 1962, 1968) and North America (Flower, 1964, 1968; VanCamp Gil, 1988). The most 28 conspicuous characters of intejocerids are their long septal necks, and their large siphuncles, 29 which are filled with heavy lamellar deposits and which form a coral-like radial pattern in cross 30 section. Like other cephalopods with long septal necks and heavy endosiphuncular deposits, 31 intejocerids were relatively large for their time with conch diameters reaching more than 100 32 mm. -
Sepkoski, J.J. 1992. Compendium of Fossil Marine Animal Families
MILWAUKEE PUBLIC MUSEUM Contributions . In BIOLOGY and GEOLOGY Number 83 March 1,1992 A Compendium of Fossil Marine Animal Families 2nd edition J. John Sepkoski, Jr. MILWAUKEE PUBLIC MUSEUM Contributions . In BIOLOGY and GEOLOGY Number 83 March 1,1992 A Compendium of Fossil Marine Animal Families 2nd edition J. John Sepkoski, Jr. Department of the Geophysical Sciences University of Chicago Chicago, Illinois 60637 Milwaukee Public Museum Contributions in Biology and Geology Rodney Watkins, Editor (Reviewer for this paper was P.M. Sheehan) This publication is priced at $25.00 and may be obtained by writing to the Museum Gift Shop, Milwaukee Public Museum, 800 West Wells Street, Milwaukee, WI 53233. Orders must also include $3.00 for shipping and handling ($4.00 for foreign destinations) and must be accompanied by money order or check drawn on U.S. bank. Money orders or checks should be made payable to the Milwaukee Public Museum. Wisconsin residents please add 5% sales tax. In addition, a diskette in ASCII format (DOS) containing the data in this publication is priced at $25.00. Diskettes should be ordered from the Geology Section, Milwaukee Public Museum, 800 West Wells Street, Milwaukee, WI 53233. Specify 3Y. inch or 5Y. inch diskette size when ordering. Checks or money orders for diskettes should be made payable to "GeologySection, Milwaukee Public Museum," and fees for shipping and handling included as stated above. Profits support the research effort of the GeologySection. ISBN 0-89326-168-8 ©1992Milwaukee Public Museum Sponsored by Milwaukee County Contents Abstract ....... 1 Introduction.. ... 2 Stratigraphic codes. 8 The Compendium 14 Actinopoda. -
Abstracts and Program. – 9Th International Symposium Cephalopods ‒ Present and Past in Combination with the 5Th
See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/265856753 Abstracts and program. – 9th International Symposium Cephalopods ‒ Present and Past in combination with the 5th... Conference Paper · September 2014 CITATIONS READS 0 319 2 authors: Christian Klug Dirk Fuchs University of Zurich 79 PUBLICATIONS 833 CITATIONS 186 PUBLICATIONS 2,148 CITATIONS SEE PROFILE SEE PROFILE Some of the authors of this publication are also working on these related projects: Exceptionally preserved fossil coleoids View project Paleontological and Ecological Changes during the Devonian and Carboniferous in the Anti-Atlas of Morocco View project All content following this page was uploaded by Christian Klug on 22 September 2014. The user has requested enhancement of the downloaded file. in combination with the 5th International Symposium Coleoid Cephalopods through Time Abstracts and program Edited by Christian Klug (Zürich) & Dirk Fuchs (Sapporo) Paläontologisches Institut und Museum, Universität Zürich Cephalopods ‒ Present and Past 9 & Coleoids through Time 5 Zürich 2014 ____________________________________________________________________________ 2 Cephalopods ‒ Present and Past 9 & Coleoids through Time 5 Zürich 2014 ____________________________________________________________________________ 9th International Symposium Cephalopods ‒ Present and Past in combination with the 5th International Symposium Coleoid Cephalopods through Time Edited by Christian Klug (Zürich) & Dirk Fuchs (Sapporo) Paläontologisches Institut und Museum Universität Zürich, September 2014 3 Cephalopods ‒ Present and Past 9 & Coleoids through Time 5 Zürich 2014 ____________________________________________________________________________ Scientific Committee Prof. Dr. Hugo Bucher (Zürich, Switzerland) Dr. Larisa Doguzhaeva (Moscow, Russia) Dr. Dirk Fuchs (Hokkaido University, Japan) Dr. Christian Klug (Zürich, Switzerland) Dr. Dieter Korn (Berlin, Germany) Dr. Neil Landman (New York, USA) Prof. Pascal Neige (Dijon, France) Dr. -
High-Level Classification of the Nautiloid Cephalopods: a Proposal for the Revision of the Treatise Part K
Swiss Journal of Palaeontology (2019) 138:65–85 https://doi.org/10.1007/s13358-019-00186-4 (0123456789().,-volV)(0123456789().,- volV) REGULAR RESEARCH ARTICLE High-level classification of the nautiloid cephalopods: a proposal for the revision of the Treatise Part K 1 2 Andy H. King • David H. Evans Received: 4 November 2018 / Accepted: 13 February 2019 / Published online: 14 March 2019 Ó Akademie der Naturwissenschaften Schweiz (SCNAT) 2019 Abstract High-level classification of the nautiloid cephalopods has been largely neglected since the publication of the Russian and American treatises in the early 1960s. Although there is broad general agreement amongst specialists regarding the status of nautiloid orders, there is no real consensus or consistent approach regarding higher ranks and an array of superorders utilising various morphological features has been proposed. With work now commencing on the revision of the Treatise Part K, there is an urgent need for a methodical and standardised approach to the high-level classification of the nautiloids. The scheme proposed here utilizes the form of muscle attachment scars as a diagnostic feature at subclass level; other features (including siphuncular structures and cameral deposits) are employed at ordinal level. We recognise five sub- classes of nautiloid cephalopods (Plectronoceratia, Multiceratia, Tarphyceratia nov., Orthoceratia, Nautilia) and 18 orders including the Order Rioceratida nov. which contains the new family Bactroceratidae. This scheme has the advantage of relative simplicity (it avoids the use of superorders) and presents a balanced approach which reflects the considerable morphological diversity and phylogenetic longevity of the nautiloids in comparison with the ammonoid and coleoid cephalopods. -
The Hydrostatics and Hydrodynamics of Prominent Heteromorph Ammonoid Morphotypes and the Functional Morphology of Ammonitic Septa
Wright State University CORE Scholar Browse all Theses and Dissertations Theses and Dissertations 2020 The Hydrostatics and Hydrodynamics of Prominent Heteromorph Ammonoid Morphotypes and the Functional Morphology of Ammonitic Septa David Joseph Peterman Wright State University Follow this and additional works at: https://corescholar.libraries.wright.edu/etd_all Part of the Environmental Sciences Commons Repository Citation Peterman, David Joseph, "The Hydrostatics and Hydrodynamics of Prominent Heteromorph Ammonoid Morphotypes and the Functional Morphology of Ammonitic Septa" (2020). Browse all Theses and Dissertations. 2306. https://corescholar.libraries.wright.edu/etd_all/2306 This Dissertation is brought to you for free and open access by the Theses and Dissertations at CORE Scholar. It has been accepted for inclusion in Browse all Theses and Dissertations by an authorized administrator of CORE Scholar. For more information, please contact [email protected]. THE HYDROSTATICS AND HYDRODYNAMICS OF PROMINENT HETEROMORPH AMMONOID MORPHOTYPES AND THE FUNCTIONAL MORPHOLOGY OF AMMONITIC SEPTA A dissertation submitted in partial fulfillment of the requirements for the degree of Doctor of Philosophy By DAVID JOSEPH PETERMAN M.S., Wright State University, 2016 B.S., Wright State University, 2014 2020 Wright State University WRIGHT STATE UNIVERSITY GRADUATE SCHOOL April 17th, 2020 I HEREBY RECOMMEND THAT THE DISSERTATION PREPARED UNDER MY SUPERVISION BY David Joseph Peterman ENTITLED The hydrostatics and hydrodynamics of prominent heteromorph ammonoid morphotypes and the functional morphology of ammonitic septa BE ACCEPTED IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF Doctor of Philosophy Committee on Final Examination Christopher Barton, PhD Dissertation Director Charles Ciampaglio, PhD Don Cipollini, PhD Director, Environmental Sciences PhD program Margaret Yacobucci, PhD Barry Milligan, PhD Interim Dean of the Graduate School Sarah Tebbens, PhD Stephen Jacquemin, PhD ABSTRACT Peterman, David Joseph. -
The Hydrostatics and Hydrodynamics of Prominent Heteromorph Ammonoid Morphotypes and the Functional Morphology of Ammonitic Septa
THE HYDROSTATICS AND HYDRODYNAMICS OF PROMINENT HETEROMORPH AMMONOID MORPHOTYPES AND THE FUNCTIONAL MORPHOLOGY OF AMMONITIC SEPTA A dissertation submitted in partial fulfillment of the requirements for the degree of Doctor of Philosophy By DAVID JOSEPH PETERMAN M.S., Wright State University, 2016 B.S., Wright State University, 2014 2020 Wright State University WRIGHT STATE UNIVERSITY GRADUATE SCHOOL April 17th, 2020 I HEREBY RECOMMEND THAT THE DISSERTATION PREPARED UNDER MY SUPERVISION BY David Joseph Peterman ENTITLED The hydrostatics and hydrodynamics of prominent heteromorph ammonoid morphotypes and the functional morphology of ammonitic septa BE ACCEPTED IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF Doctor of Philosophy Committee on Final Examination Christopher Barton, PhD Dissertation Director Charles Ciampaglio, PhD Don Cipollini, PhD Director, Environmental Sciences PhD program Margaret Yacobucci, PhD Barry Milligan, PhD Interim Dean of the Graduate School Sarah Tebbens, PhD Stephen Jacquemin, PhD ABSTRACT Peterman, David Joseph. PhD. Environmental Sciences PhD Program, Wright State University, 2020. The hydrostatics and hydrodynamics of prominent heteromorph ammonoid morphotypes and the functional morphology of ammonitic septa. Ammonoid cephalopods have chambered shells that regulated buoyancy. The morphology of their shells strongly influenced the physical properties acting on these animals during life. Heteromorph ammonoids, which undergo changes in coiling throughout ontogeny, are the focus of this dissertation. The biomechanics of these cephalopods are investigated in a framework involving functional morphology, paleoecology, and possible modes of life. Constructional constraints were investigated for the marginally-corrugated septal walls within the chambered ammonoid shell. These constraints governed the positive relationship between septal complexity and terminal size. Furthermore, increased septal complexity facilitated liquid retention via surface tension. -
Nautiloid Cephalopods – a Review of Their Use and Potential in Biostratigraphy
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Denisia Jahr/Year: 2014 Band/Volume: 0032 Autor(en)/Author(s): Evans David H., King Andy H., Histon Kathleen, Cichowolski Marcela Artikel/Article: Nautiloid cephalopods - a review of their use and potential in biostratigraphy 7-22 © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at Nautiloid cephalopods – a review of their use and potential in biostratigraphy D.H. EVANS, A.H. KING, K. HISTON & M. CICHOWOLSKI Abstract: In terms of their use as biostratigraphical tools, nautiloid cephalopods are the poor relations of ammonoids. Neverthe- less, in certain situations, they may provide useful biostratigraphical data, particularly where other biostratigraphically valuable taxa are not present; or in certain situations demonstrate a resolution as great as, or greater than ammonoids, trilobites, grapto- lites or conodonts. Nautiloid cephalopods are of especial value in palaeobiogeographical studies, but their use for this purpose may be hampered by the poor understanding of the stratigraphical ranges of individual taxa. The biostratigraphical value of nautiloid cephalopods is demonstrated here through a number of case studies of Ordovician taxa, combined with a review of their biostrati- graphical use in Palaeozoic and Mesozoic successions. These both demonstrate the potential of this group and indicate great scope for further research. Keywords: Nautiloids, cephalopods, biostratigraphy, Palaeozoic, Mesozoic Introduction taxonomically without investigation of the internal morphology of the phragmocone due to the similarities The widespread stratigraphical and geographical of the external features, thus entailing the preparation occurrence of nautiloid cephalopods, particularly, but of polished or thin sections for study, requiring more ef- not exclusively during the Lower Palaeozoic is docu- fort, and possibly resulting in more equivocal data than mented in many substantial monographic works (e.g. -
8 International Symposium Cephalopods
8th International Symposium Cephalopods – Present and Past August 30 – September 3, 2010 Abstracts Volume University of Burgundy & CNRS Dijon - France http://www.u-bourgogne.fr/cephalopods/ Honorary Committee Sigurd von Boletzky, DR CNRS - Banyuls-sur-Mer - France Raymond Enay, Prof. University of Lyon Lyon - France Didier Marchand, University of Burgundy Dijon - France Jacques Thierry, Prof. University of Burgundy Dijon - France Scientific committee Giambattista Bello, ARION, Mola di Bari - Italy Vyacheslav Bizikov, Russian Federal Research Institute of Marine Fisheries and Oceanography, Moscou - Russia Christian Klug, Universitaet Zuerich, Zurich - Switzerland Neil. H. Landman, American Museum of Natural History, New York - USA Pascal Neige, University of Burgundy, Dijon - France Isabelle Rouget, University Pierre & Marie Curie, Paris - France Kazushige Tanabe, University of Tokyo, Tokyo - Japan Margareth Yacobuci, Bowling Green State University, Bowling Green - USA Organizing committee Pascal Neige, University of Burgundy – France Isabelle Rouget, University Pierre and Marie Curie – France Alex Bauer, CNRS, University of Burgundy – France Arnaud Brayard, CNRS, University of Burgundy – France Guillaume Dera, University of Burgundy – France Jean-Louis Dommergues, CNRS, University of Burgundy – France Emmanuel Fara, University of Burgundy – France Myette Guiomar, Rserve Gologique de Haute-Provence Digne-les Bains, France Clotilde Hardy, University of Burgundy – France Isabella Kruta, MNHN – France Rémi Laffont, CNRS, University of Burgundy -
Morphological Characters, Systematic Position and Age of Fossil Genera Pertaining to Cephalopods
EXPERIMENT 3 MORPHOLOGICAL CHARACTERS, SYSTEMATIC POSITION AND AGE OF FOSSIL GENERA PERTAINING TO CEPHALOPODS Outline of Experiment_______________________________________________________ 3.1 Introduction 3.5 Common Genera of Ammonoids Expected Learning Skills 3.6 Laboratory Exercises 3.2 Requirements 3.7 Results and Discussion 3.3 Cephalopods: Systematics, General 3.8 References Morphology and Age Range 3.9 Further/Suggested Readings 3.4 Common Genera of Nautiloids 3.1 INTRODUCTION In the previous experiment, you have made line drawings and described morphological characters along with palaeoecology and stratigraphic range of various genera of bivalves (pelecypods). Every genus of bivalves displays a set of distinctive morphological characters, on which basis they can be differentiated from each other. You know that bivalves show a wide range of adaptation, from marine to freshwater and from infaunal burrowers, epifaunal to active swimmers. Therefore, marine bivalves possess different morphological characters as compared with those of non-marine forms. In this experiment, you will work with some common genera of cephalopod molluscs. They are exclusively marine animals and are more active, quick movers, intelligent carnivorous predators with varied shell morphology. They occupy the same ecological niche as fishes. The Class Cephalopoda consists of three subclasses: Nautiloidea, Ammonoidea and Coleoidea. Among which Nautiloidea and extinct Ammonoidea are commonly found as fossils. The third subclass, Coleoidea, comprises BGYCL………………………………………………………………………………………………….….............….…...-138 Stratigraphy and Palaeontology: Laboratory primarily soft bodied animals. Like fishes, cephalopods are equipped with highly developed eyes and sensory organs. They are adapted to fast swimming and have developed more efficient mechanisms to locate their prey, detect predators and to escape from them.