DOCSLIB.ORG

Pdf (306.01 K)

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Pdf (306.01 K) REVIEW ARTICLE RECORDS OF PHARMACEUTICAL AND BIOMEDICAL SCIENCES Review article on chemical constituents and biological activity of Thymelaea hirsuta. Ahmed M Badawya, Hashem A Hassaneanb, Amany K. Ibrahimb, Eman S. Habibb, Safwat A. Ahmedb* aDepartment of Pharmacognosy, Faculty of Pharmacy, Sinai University, El-Arish, Egypt, b Department of Pharmacognosy, Faculty of Pharmacy, Suez Canal University, Ismailia, Egypt 41522. Abstract Received on: 07.04. 2019 Thymelaea hirsuta a perennial, evergreen and dioecious shrub, which is native Revised on: 30. 04. 2019 to North Africa. T. hirsuta is a widespread invasive weed and is commonly known as “Methnane”. Along the history, T. hirsuta, family Thymelaeaceae, Accepted on: 10. 04. 2019 has been recognized as an important medicinal plant. Much research has been carried out on the medical applications of Methnane. The choice of the plant was based on the good previous biological study of T. hirsuta plant extract to Correspondence Author: use as anticancer, hepatoprotective and anti-diapetic. Several species of Tel:+ 01092638387 Thymelaeaceae have been the subject of numerous phytochemical studies. Initially, interest may have been due to the marked toxicity of these plants, but E-mail address: the widespread use of some species medicinally has certainly played a part in [email protected] sustaining this interest. Keywords: Thymelaea hirsuta , Chemical constituents, Biological activity 1.Introduction: Near East: Lebanon and Palestine. The choice of the plant was based on the good previous Thymelaea hirsuta a perennial, evergreen and biological study of T. hirsuta plant extract to use dioecious shrub, which is native to North Africa. T. as anticancer, hepatoprotective and anti-diabetic. hirsuta is a widespread invasive weed and is Several species of Thymelaeaceae have been the commonly known as “Methnane”. subject of numerous phytochemical studies. Initially, interest may have been due to the Along the history, T. hirsuta, family marked toxicity of these plants, but the Thymelaeaceae, has been recognized as an widespread use of some species medicinally has important medicinal plant. Much research has been certainly played a part in sustaining this interest carried out on the medical applications of (Beaumont et al., 2001). Several genera such as Methnane. The current systematic classification of Daphne, Thymelaea, Pimelea, Wikstroemia and cannabis is listed in Table 1. Gnidia have been researched upon extensively. Species Profile Geography and Distribution The Daphne genus is of prime importance owing to its richness in a variety of different classes of T. hirsuta grows in the Mediterranean coastal natural products, especially, coumarins (Li et al., plains, the Sinai Peninsula and other Saharo- 2005, Ullah et al., 1999), flavones (Zhang et al., Arabian deserts. Regional: From Morocco to Egypt. 2004, Baba et al., 1995, Liang et al., 2011), Global: The Mediterranean: from Spain to Greece daphnane-type diterpene esters (Li et al., 2006), and Turkey; southern side from Morocco to Egypt. steroids and guianolides (Levyns, 1950). Rec. Pharm. Biomed. Sci. 3(2), 28- 32, 2019 Division Tracheophyta Class Magnoliopsida Superorder Rosanae Order Malvales Family Thymelaeaceae Genus Thymelaea Species hirsuta. Table 1: The systematic classification of Thymelaea hirsuta Fig. (1): A photo of Thymelaea hirsuta 3.1 Anti-inflammatory activities distribution. Azza and Oudghiri reported that the aqueous extract of T. hirsuta had an anti-inflammatory Gnidia genus has similar chemical constitution as activity and inhibited the induction of adjuvant that Daphne. Chemical studies done on some arthritis in male Wistar rats (Azza and Oudghiri, Gnidia species indicated the presence of toxic 2015). diterpene esters of daphnane type, which are the main types of plant orthoesters known and have 3.2 Antioxidant activities remarkable biological activities, such as the tradition use of Gnidia as antineoplastic and Amari conclouded that T. hirsuta extracts are rich cytotoxic (Kupchan et al., 1976, Borris and Cordell, sources of natural antioxidants which appears to 1984, Badawi et al., 1983, Kupchan et al., 1975). be an alternative to synthetic antioxidants(Amari et al., 2014). 2. Chemical constituents reported from T. hirsuta: 3.3 Antimicrobialactivities 2. 1. Terpenoidal and steroidal compounds Felhi revealed that the essential oil isolated by the hydro-distillation of aerial parts of T. hirsuta Miyamae isolated two new daphnane diterpenoids, exhibited a moderate-to-potent anti-microbial hirsein A and hirsein B from the aerial parts of T. activity against all the microorganisms tested. hirsuta (Miyamae et al., 2009) are listed in Table 2. Gram-positive bacteria were noted to be more sensitive to the oil than gram-negative bacteria 2. 2. Flavonoids and yeasts (Felhi et al., 2017). Yang isolated mono- and biflavonoids as 3.4 Antidiabetic activities genkwanin, genkwanin 5-O-β-D-glucopyranoside, genkwanin 5-O-β-D-primeveroside, tiliroside and El Amrani founded that T. hirsuta possesses both neochamaejasmin B from T. hirsuta (Yang et al., hypoglycaemic and antidiabetic activities in 2014) are listed in Table 2. normoglycaemic and STZ diabetic rats(El Amrani et al., 2009). 2. 3. Coumarins Dahamna concluded that a single administration Yang isolated bi- and tricoumarins (daphnoretin and of aqueous extract of T. hirsute at a dose of 250 triumbelletin) from T. hirsuta (Yang et al., 2014) mg/kg bodyweight caused a very significant are listed in Table 2. decrease in blood glucose level in rat (Dahamna 2. 4. Lignans et al., 2015). Yang isolated lignans (pinoresinol and 3.5 Antimelanogenesis activities syringaresinol) from T. hirsuta (Yang et al., 2014) Miyamae concoluded that T. hirsuta has are listed in Table 2. antimelanogenesis effect (without cytotoxicity) 3. Biological activities reported from T. on B16 murine melanoma cells by hirsuta: downregulating the expressions of the Mitf gene Badawy et.al 30 Table (2): Examples of some compounds isolated from T. hirsuta of family Thymelaeaceae. Structure Name Plant Reference O O Hirsein A T. hirsuta (Miyamae et al., 2009) O H O O H O O HO HO OH HO O O Hirsein B T. hirsuta (Miyamae et al., 2009) O H O O H O O HO HO OH OH H3CO O Genkwanin T. hirsuta (Yang et al., 2014) OH O OH Genkwanin-5-O- RO O primeveroside T. hirsuta (Yang et al., 2014) R1= glc-xyl OR O R =CH 2 3 OH R O O Genkwanin 5-O- βDglucopyranoside T. hirsuta (Yang et al., 2014) R1=glc OR O R2=CH3 OH HO O O O O O Trans-Tiliroside T. hirsuta (Yang et al., 2014) OH OH O HO OH HO OH H HO O O OH H Neochamaejasmin B H T. hirsuta (Yang et al., 2014) OH O O OH H HO Rec. Pharm. Biomed. Sci. 3(2), 28- 32, 2019 H3CO O O O Daphnoretin T. hirsuta (Yang et al., 2014) HO O O O O O O HO O triumbelletin T. hirsuta (Yang et al., 2014) O OH O OH H OCH3 O Pinoresinol T. hirsuta (Yang et al., 2014) H3CO O HO H OH H3CO OCH3 O H Syringaresinol H T. hirsuta (Yang et al., 2014) H3CO O HO OCH3 and the melanogenic enzymes’ genes(Miyamae et 5. Conflict of interest al., 2009). The authors report no declaration of conflict of 3.6 Antifugalactivities interest. Dahamna indicated that extracts of T. hirsuta 6. References possesses antifugal activity on both fungi Akrout, A., Gonzalez, L.A., Eljani, H., Madrid, Microsporum audouinii and Microsporum gypseum P.C., 2011. Antioxidant and antitumor activities (Dahamna et al., 2015). of Artemisia campestris and Thymelaea hirsuta 3.7 Hepatoprotective activities from southern Tunisia. Food and Chemical Toxicology, 49, 342-347. The biochemical observations and the histopathological examination of liver sections Amari, N.O., Bouzouina, M., Berkani, A., indicated that T. hirsute possesses no toxicity Lotmani, B., 2014. Phytochemical screening and effects but hepatoprotective properties against CCl4- antioxidant capacity of the aerial parts of induced hepatic injury in rats(Azza et al., 2012). Thymelaea hirsuta L. Asian Pacific journal of tropical disease, 4, 104-109. 3.8 Cytotoxicity activities Azza, Z., Oudghiri, M., 2015. In vivo anti- Akrout found that all the extracts of T. hirsuta inflammatory and antiarthritic activities of tested (hexane and ethanol–water), except the aqueous extracts from Thymelaea hirsuta. infusion extract, exhibited antitumor activity against Pharmacognosy Res, 7, 213-6. human colon cancer HT-29 cells using MTT test activities (Akrout et al., 2011). Azza, Z., Marnissi, F., Naya, A., Benjelloun, N., Zamyati, S., Amrani, M., Oudghiri, M., 2012. 4. Conclusion Toxicological evaluation of Thymelaea hirsuta and protective effect against CCl 4-induced Thymelae hirsuta are growing all over the world as hepatic injury in rats. International Journal of it used for medicinal purposes. Here we just report a Biological and Chemical Sciences, 6, 379-393. brief review for the chemical constituent and biological activity from the plant. Baba, K., Yoshikawa, M., Taniguchi, M., Badawy et.al 32 Kozawa, M., 1995. Biflavonoids from Daphne Kupchan, S.M., Sweeny, J., Murae, T., Shen, odora. Phytochemistry, 38, 1021-1026. M.S., Bryan, R., 1975. Structure of gnidicoumarin, a novel pentacyclic dicoumarin Badawi, M.M., Handa, S.S., Kinghorn, A.D., from Gnidia lamprantha. Journal of the Cordell, G.A., Farnsworth, N. R., 1983. Plant Chemical Society, Chemical Communications, anticancer agents XXVII: antileukemic and 94-95. cytotoxic constituents of Dirca occidentalis (Thymelaeaceae). Journal of pharmaceutical Levyns, M. 1950. Thymelaeaceae. Flora of the sciences, 72, 1285-1287. Cape Peninsula. Juta. Cape Town. Beaumont, A., Edwards, T., Smith, F., 2001. Leaf Li, P., Xiaofeng, Z., Haifeng, W., Lisheng, D., and bract diversity in Gnidia (Thymelaeaceae): 2006. A new daphnane diterpene from Daphne patterns and taxonomic value. Systematics and tangutica. Geography of Plants, 399-418. Li, S.H., Wu, L.J., Gao, H.Y., Chen, Y.H. & LI, Borris, R.P., Cordell, G.A., 1984. Studies of the Y. 2005. A new dicoumarinoid glycoside from Thymelaeaceae II.
Load more

Recommended publications
  • Plants for a Future Species Database Bibliography
    Plants For A Future Species Database Bibliography Numbers in square brackets are the reference numbers that appear in the database. [K] Ken Fern Notes from observations, tasting etc at Plants For A Future and on field trips. [1] F. Chittendon. RHS Dictionary of Plants plus Supplement. 1956 Oxford University Press 1951 Comprehensive listing of species and how to grow them. Somewhat outdated, it has been replaces in 1992 by a new dictionary (see [200]). [1b] Food Plants International. http://foodplantsinternational.com/plants/ [1c] Natural Resources Conservation Service http://plants.usda.gov [1d] Invasive Species Compendium www.cabi.org [2] Hedrick. U. P. Sturtevant's Edible Plants of the World. Dover Publications 1972 ISBN 0-486-20459-6 Lots of entries, quite a lot of information in most entries and references. [3] Simmons. A. E. Growing Unusual Fruit. David and Charles 1972 ISBN 0-7153-5531-7 A very readable book with information on about 100 species that can be grown in Britain (some in greenhouses) and details on how to grow and use them. [4] Grieve. A Modern Herbal. Penguin 1984 ISBN 0-14-046-440-9 Not so modern (1930's?) but lots of information, mainly temperate plants. [5] Mabey. R. Food for Free. Collins 1974 ISBN 0-00-219060-5 Edible wild plants found in Britain. Fairly comprehensive, very few pictures and rather optimistic on the desirability of some of the plants. [6] Mabey. R. Plants with a Purpose. Fontana 1979 ISBN 0-00-635555-2 Details on some of the useful wild plants of Britain. Poor on pictures but otherwise very good.
    [Show full text]
  • APPROVED PLANT LIST Midtown Alliance Tree Well Adoption Program
    APPROVED PLANT LIST Midtown Alliance Tree Well Adoption Program Midtown Alliance launched the Tree Well Adoption program with the primary goal of enriching the experience of Midtown’s workers and residents while encouraging sustainability through the use of low-water, urban tolerant plant species. This list of plants was created to aid individuals and organizations in selecting plant material to plant in their adopted tree wells. This plant list is intended to encourage individual character in the tree wells, rather than restrict creativity in the selection of plants. The plants on the approved list were selected based on the following criteria: • Perennial. All plants listed are perennial, meaning they last for two or more growing seasons. Once established, these plants will require less water to maintain than annuals. • Heat tolerant. Plants in tree wells are exposed to high temperatures caused by vehicles and heat reflected from surrounding buildings, asphalt, and other urban surfaces. They must also be tolerant to high daytime temperatures, typical of Atlanta’s summer months, and cold hardy in the winter months. Atlanta is located in USDA Plant Hardiness Zone 7b/8a. • Water wise. Urban tree wells are surrounded by impervious surfaces and thus, are highly susceptible to periods of drought. Suitable plants must be able to survive periods of low rainfall. • Pollution tolerant. Vehicle exhaust may leave deposits and pollutants on plant foliage, which can kill sensitive plants. • Encourage wildlife. Flowering plants attract insects such as butterflies while others provide food sources for birds and other wildlife. • Grown locally. Many of the plants listed are native to the Atlanta area, and all can be found at local nurseries.
    [Show full text]
  • Thymelaeaceae)
    Origin and diversification of the Australasian genera Pimelea and Thecanthes (Thymelaeaceae) by MOLEBOHENG CYNTHIA MOTS! Thesis submitted in fulfilment of the requirements for the degree PHILOSOPHIAE DOCTOR in BOTANY in the FACULTY OF SCIENCE at the UNIVERSITY OF JOHANNESBURG Supervisor: Dr Michelle van der Bank Co-supervisors: Dr Barbara L. Rye Dr Vincent Savolainen JUNE 2009 AFFIDAVIT: MASTER'S AND DOCTORAL STUDENTS TO WHOM IT MAY CONCERN This serves to confirm that I Moleboheng_Cynthia Motsi Full Name(s) and Surname ID Number 7808020422084 Student number 920108362 enrolled for the Qualification PhD Faculty _Science Herewith declare that my academic work is in line with the Plagiarism Policy of the University of Johannesburg which I am familiar. I further declare that the work presented in the thesis (minor dissertation/dissertation/thesis) is authentic and original unless clearly indicated otherwise and in such instances full reference to the source is acknowledged and I do not pretend to receive any credit for such acknowledged quotations, and that there is no copyright infringement in my work. I declare that no unethical research practices were used or material gained through dishonesty. I understand that plagiarism is a serious offence and that should I contravene the Plagiarism Policy notwithstanding signing this affidavit, I may be found guilty of a serious criminal offence (perjury) that would amongst other consequences compel the UJ to inform all other tertiary institutions of the offence and to issue a corresponding certificate of reprehensible academic conduct to whomever request such a certificate from the institution. Signed at _Johannesburg on this 31 of _July 2009 Signature Print name Moleboheng_Cynthia Motsi STAMP COMMISSIONER OF OATHS Affidavit certified by a Commissioner of Oaths This affidavit cordons with the requirements of the JUSTICES OF THE PEACE AND COMMISSIONERS OF OATHS ACT 16 OF 1963 and the applicable Regulations published in the GG GNR 1258 of 21 July 1972; GN 903 of 10 July 1998; GN 109 of 2 February 2001 as amended.
    [Show full text]
  • Review of Three Dirca Species with Special Emphasis on D. Occidentalis Background
    Review of Three Dirca Species With Special Emphasis on D. Occidentalis Jasper Ridge Biological Preserve Docent Research Report John Kriewall May 29,2001 Table of Contents Summary Page Background 1 Why study dirca? 2 Objectives 2 Conclusions 3 Discussion Literature Search and Network 6 Biogeography 7 Species Comparisons 10 DO Compared to DP 11 DM Compared to DO and DP 13 Habitat Comparisons 14 Toxicity and Chemical Analysis 15 Locations and Quantity of DO at JRBP 16 DO at Other Bay Area Locations 17 Pollinators 21 Propagation by Other Means 22 Seed Dispersal 22 Horticultural Attempts at Propagation 22 Additional Studies Required 23 Research in the Mail 26 Next? 27 N. American Ranges of D. species 1 Figures 1. Locations ofDO at JRPB (GPS) 3a 2. Early Tertiary Paleogeographic Map 8 3A. Mexican~Appalachian Paleogeography-! 9 3B. Mexican-Appalachian Paleogeography-II 9 4A. Stem Photomicrograph 10 4B. Enlarged Stem Photomicrograph 10 5A. DO v DP Flower Comparisons 12 5B DO v DP Leaf Comparisons 12 6 DM Sketches 14 Tables 1 Key: DO and DO 11 2 DP and DO Flower-LeafMetrics 13 3. Key: DO, DP and DM 13 Appendices I Bibliography 8 pages Published Papers 1 Specimen Images (Internet) 2 Descriptions/Overview (Internet) 2 Toxicity, Agricultural (Internet) 3 Horticultural/Landscaping/Nurseries (Internet) 3 Mise (Internet) 4 Citations not Retrieved 4 Pers Communications 4-8 II Network of Contacts 1-6 III Comparison ofDO, DP, and DM 1-7 Docent Research Report John Kriewall 5/29/01 Final Edit 8/01 Review of Three Dirca Species With Special Emphasis on D.
    [Show full text]
  • Phytochemistry of Dais Cotinifolia L
    Phytochemistry of Dais cotinifolia L. Thesis submitted in fulfilment of the requirements for the degree Master of Science By Busisiwe Buyisile Danca School of Chemistry and Physics University of KwaZulu-Natal Pietermaritzburg Supervisor: Professor Fanie R. van Heerden May 2018 | Page Abstract Many plant-derived secondary metabolites have interesting biological activities and some plant compounds such as artemisinin (antimalarial) and vincristine (anticancer agent) are the active principles of main-stream pharmaceuticals. South Africa has an immense biodiversity and there are in the order of 25 000 indigenous plant species in the country. The phytochemistry of many of these plants has never been investigated. Many bioactive compounds have been isolated from the Thymelaeaceae, such as prostatin (potent anti-HIV agent) and antitumour diterpenoids. Although a large number of Thymelaeaceae species are indigenous to South Africa, only a limited number of species have been subjected to phytochemical investigations. Dais cotinifolia L. is one species on which phytochemical results have not been published and this tree was the subject of this investigation. Four compounds were isolated from D. cotinifolia. Three of these compounds are known metabolites, two furofuran lignans, kobusin and eudesmin, and the flavonoid catechin. A fourth compound is novel norlignan, 2-hydroxy-5-(3-methoxyphenyl)-1-(4- methoxyphenyl)pentan-1-one. The compounds were isolated by applying different chromatographic procedures, such as column chromatography, preparative centrifugal thin-layer chromatography and semi-preparative high-performance liquid chromatography. The structural elucidation of the compounds was based on mass spectrometry and nuclear magnetic spectroscopy. The antioxidant activities of the crude extracts of the different plant parts were determined by the DPPH assay.
    [Show full text]
  • Heterodichogamy.Pdf
    Research Update TRENDS in Ecology & Evolution Vol.16 No.11 November 2001 595 How common is heterodichogamy? Susanne S. Renner The sexual systems of plants usually Heterodichogamy differs from normal (Zingiberales). These figures probably depend on the exact spatial distribution of dichogamy, the temporal separation of underestimate the frequency of the gamete-producing structures. Less well male and female function in flowers, in heterodichogamy. First, the phenomenon known is how the exact timing of male and that it involves two genetic morphs that is discovered only if flower behavior is female function might influence plant occur at a 1:1 ratio. The phenomenon was studied in several individuals and in mating. New papers by Li et al. on a group discovered in walnuts and hazelnuts5,6 natural populations. Differential of tropical gingers describe differential (the latter ending a series of Letters to movements and maturation of petals, maturing of male and female structures, the Editor about hazel flowering that styles, stigmas and stamens become such that half the individuals of a began in Nature in 1870), but has gone invisible in dried herbarium material, population are in the female stage when almost unnoticed7. Indeed, its recent and planted populations deriving from the other half is in the male stage. This discovery in Alpinia was greeted as a vegetatively propagated material no new case of heterodichogamy is unique new mechanism, differing ‘from other longer reflect natural morph ratios. The in involving reciprocal movement of the passive outbreeding devices, such as discovery of heterodichogamy thus styles in the two temporal morphs. dichogamy…and heterostyly in that it depends on field observations.
    [Show full text]
  • Towards an Updated Checklist of the Libyan Flora
    Towards an updated checklist of the Libyan flora Article Published Version Creative Commons: Attribution 3.0 (CC-BY) Open access Gawhari, A. M. H., Jury, S. L. and Culham, A. (2018) Towards an updated checklist of the Libyan flora. Phytotaxa, 338 (1). pp. 1-16. ISSN 1179-3155 doi: https://doi.org/10.11646/phytotaxa.338.1.1 Available at http://centaur.reading.ac.uk/76559/ It is advisable to refer to the publisher’s version if you intend to cite from the work. See Guidance on citing . Published version at: http://dx.doi.org/10.11646/phytotaxa.338.1.1 Identification Number/DOI: https://doi.org/10.11646/phytotaxa.338.1.1 <https://doi.org/10.11646/phytotaxa.338.1.1> Publisher: Magnolia Press All outputs in CentAUR are protected by Intellectual Property Rights law, including copyright law. Copyright and IPR is retained by the creators or other copyright holders. Terms and conditions for use of this material are defined in the End User Agreement . www.reading.ac.uk/centaur CentAUR Central Archive at the University of Reading Reading’s research outputs online Phytotaxa 338 (1): 001–016 ISSN 1179-3155 (print edition) http://www.mapress.com/j/pt/ PHYTOTAXA Copyright © 2018 Magnolia Press Article ISSN 1179-3163 (online edition) https://doi.org/10.11646/phytotaxa.338.1.1 Towards an updated checklist of the Libyan flora AHMED M. H. GAWHARI1, 2, STEPHEN L. JURY 2 & ALASTAIR CULHAM 2 1 Botany Department, Cyrenaica Herbarium, Faculty of Sciences, University of Benghazi, Benghazi, Libya E-mail: [email protected] 2 University of Reading Herbarium, The Harborne Building, School of Biological Sciences, University of Reading, Whiteknights, Read- ing, RG6 6AS, U.K.
    [Show full text]
  • These De Doctorat De L'universite Paris-Saclay
    NNT : 2016SACLS250 THESE DE DOCTORAT DE L’UNIVERSITE PARIS-SACLAY, préparée à l’Université Paris-Sud ÉCOLE DOCTORALE N° 567 Sciences du Végétal : du Gène à l’Ecosystème Spécialité de doctorat (Biologie) Par Mlle Nour Abdel Samad Titre de la thèse (CARACTERISATION GENETIQUE DU GENRE IRIS EVOLUANT DANS LA MEDITERRANEE ORIENTALE) Thèse présentée et soutenue à « Beyrouth », le « 21/09/2016 » : Composition du Jury : M., Tohmé, Georges CNRS (Liban) Président Mme, Garnatje, Teresa Institut Botànic de Barcelona (Espagne) Rapporteur M., Bacchetta, Gianluigi Università degli Studi di Cagliari (Italie) Rapporteur Mme, Nadot, Sophie Université Paris-Sud (France) Examinateur Mlle, El Chamy, Laure Université Saint-Joseph (Liban) Examinateur Mme, Siljak-Yakovlev, Sonja Université Paris-Sud (France) Directeur de thèse Mme, Bou Dagher-Kharrat, Magda Université Saint-Joseph (Liban) Co-directeur de thèse UNIVERSITE SAINT-JOSEPH FACULTE DES SCIENCES THESE DE DOCTORAT DISCIPLINE : Sciences de la vie SPÉCIALITÉ : Biologie de la conservation Sujet de la thèse : Caractérisation génétique du genre Iris évoluant dans la Méditerranée Orientale. Présentée par : Nour ABDEL SAMAD Pour obtenir le grade de DOCTEUR ÈS SCIENCES Soutenue le 21/09/2016 Devant le jury composé de : Dr. Georges TOHME Président Dr. Teresa GARNATJE Rapporteur Dr. Gianluigi BACCHETTA Rapporteur Dr. Sophie NADOT Examinateur Dr. Laure EL CHAMY Examinateur Dr. Sonja SILJAK-YAKOVLEV Directeur de thèse Dr. Magda BOU DAGHER KHARRAT Directeur de thèse Titre : Caractérisation Génétique du Genre Iris évoluant dans la Méditerranée Orientale. Mots clés : Iris, Oncocyclus, région Est-Méditerranéenne, relations phylogénétiques, status taxonomique. Résumé : Le genre Iris appartient à la famille des L’approche scientifique est basée sur de nombreux Iridacées, il comprend plus de 280 espèces distribuées outils moléculaires et génétiques tels que : l’analyse de à travers l’hémisphère Nord.
    [Show full text]
  • Illustrated Flora of East Texas Illustrated Flora of East Texas
    ILLUSTRATED FLORA OF EAST TEXAS ILLUSTRATED FLORA OF EAST TEXAS IS PUBLISHED WITH THE SUPPORT OF: MAJOR BENEFACTORS: DAVID GIBSON AND WILL CRENSHAW DISCOVERY FUND U.S. FISH AND WILDLIFE FOUNDATION (NATIONAL PARK SERVICE, USDA FOREST SERVICE) TEXAS PARKS AND WILDLIFE DEPARTMENT SCOTT AND STUART GENTLING BENEFACTORS: NEW DOROTHEA L. LEONHARDT FOUNDATION (ANDREA C. HARKINS) TEMPLE-INLAND FOUNDATION SUMMERLEE FOUNDATION AMON G. CARTER FOUNDATION ROBERT J. O’KENNON PEG & BEN KEITH DORA & GORDON SYLVESTER DAVID & SUE NIVENS NATIVE PLANT SOCIETY OF TEXAS DAVID & MARGARET BAMBERGER GORDON MAY & KAREN WILLIAMSON JACOB & TERESE HERSHEY FOUNDATION INSTITUTIONAL SUPPORT: AUSTIN COLLEGE BOTANICAL RESEARCH INSTITUTE OF TEXAS SID RICHARDSON CAREER DEVELOPMENT FUND OF AUSTIN COLLEGE II OTHER CONTRIBUTORS: ALLDREDGE, LINDA & JACK HOLLEMAN, W.B. PETRUS, ELAINE J. BATTERBAE, SUSAN ROBERTS HOLT, JEAN & DUNCAN PRITCHETT, MARY H. BECK, NELL HUBER, MARY MAUD PRICE, DIANE BECKELMAN, SARA HUDSON, JIM & YONIE PRUESS, WARREN W. BENDER, LYNNE HULTMARK, GORDON & SARAH ROACH, ELIZABETH M. & ALLEN BIBB, NATHAN & BETTIE HUSTON, MELIA ROEBUCK, RICK & VICKI BOSWORTH, TONY JACOBS, BONNIE & LOUIS ROGNLIE, GLORIA & ERIC BOTTONE, LAURA BURKS JAMES, ROI & DEANNA ROUSH, LUCY BROWN, LARRY E. JEFFORDS, RUSSELL M. ROWE, BRIAN BRUSER, III, MR. & MRS. HENRY JOHN, SUE & PHIL ROZELL, JIMMY BURT, HELEN W. JONES, MARY LOU SANDLIN, MIKE CAMPBELL, KATHERINE & CHARLES KAHLE, GAIL SANDLIN, MR. & MRS. WILLIAM CARR, WILLIAM R. KARGES, JOANN SATTERWHITE, BEN CLARY, KAREN KEITH, ELIZABETH & ERIC SCHOENFELD, CARL COCHRAN, JOYCE LANEY, ELEANOR W. SCHULTZE, BETTY DAHLBERG, WALTER G. LAUGHLIN, DR. JAMES E. SCHULZE, PETER & HELEN DALLAS CHAPTER-NPSOT LECHE, BEVERLY SENNHAUSER, KELLY S. DAMEWOOD, LOGAN & ELEANOR LEWIS, PATRICIA SERLING, STEVEN DAMUTH, STEVEN LIGGIO, JOE SHANNON, LEILA HOUSEMAN DAVIS, ELLEN D.
    [Show full text]
  • Daphne Mezereum L. Mezereon
    Daphne mezereum L. Mezereon Starting references Family Thymelaeaceae IUCN category (2001) Vulnerable Habit Deciduous shrub. Habitat Calcareous woodland, often on steep, sometimes rocky slopes with little ground cover, but rarely in deep shade; also in chalk-pits, and in wet, species-rich fens. Reasons for decline Habitat loss and uprooting. Distribution in wild Country Locality & Vice County Sites Population (10km2 occurences) (plants) Wales Carmarthenshire 2 England Westmorland 1 W Lancashire 1 MW Yorkshire 3 Derbyshire 4 W Norfolk 1 E Gloucestershire 1 Oxfordshire 2 Berkshire 3 Buckinghamshire 1 Surrey 2 N Hampshire 2 W Sussex 1 Dorset 1 Ex situ Collections Gardens close to the region of distribution of the species 1 Holehird Gardens 2 Sizergh Castle (NT) 3 Sheffield Botanical Gardens 4 National Botanic Garden of Wales 5 Swansea Botanic Garden,17 6 Hidcote Manor (NT) 7 Batsford Arboretum 8 Abbotsbury Subtropical Garden 9 University of Oxford Botanic Garden 10 The Harris Garden 11 Windsor Gardens 12 Cliveden (NT) 13 High Beeches Gardens 14 Nymans Garden (NT) 15 Borde Hill Garden 16 RBG Kew 17 RHS Wisley 18 Cambridge University Botanic Garden Gardens with specialisation on genus Daphne Ness Botanic Garden Potential to grow the species in ex situ Collections In the wild, reproduces by seed and is self-fertile. From Plants For A Future • Propagation Seed - best sown in a greenhouse as soon as it is ripe with the pot sealed in a polythene bag to hold in the moisture. Remove this bag as soon as germination takes place. The seed usually germinates better if it is harvested 'green' (when it has fully developed but before it dries on the plant) and sown immediately.
    [Show full text]
  • Flora Mediterranea 26
    FLORA MEDITERRANEA 26 Published under the auspices of OPTIMA by the Herbarium Mediterraneum Panormitanum Palermo – 2016 FLORA MEDITERRANEA Edited on behalf of the International Foundation pro Herbario Mediterraneo by Francesco M. Raimondo, Werner Greuter & Gianniantonio Domina Editorial board G. Domina (Palermo), F. Garbari (Pisa), W. Greuter (Berlin), S. L. Jury (Reading), G. Kamari (Patras), P. Mazzola (Palermo), S. Pignatti (Roma), F. M. Raimondo (Palermo), C. Salmeri (Palermo), B. Valdés (Sevilla), G. Venturella (Palermo). Advisory Committee P. V. Arrigoni (Firenze) P. Küpfer (Neuchatel) H. M. Burdet (Genève) J. Mathez (Montpellier) A. Carapezza (Palermo) G. Moggi (Firenze) C. D. K. Cook (Zurich) E. Nardi (Firenze) R. Courtecuisse (Lille) P. L. Nimis (Trieste) V. Demoulin (Liège) D. Phitos (Patras) F. Ehrendorfer (Wien) L. Poldini (Trieste) M. Erben (Munchen) R. M. Ros Espín (Murcia) G. Giaccone (Catania) A. Strid (Copenhagen) V. H. Heywood (Reading) B. Zimmer (Berlin) Editorial Office Editorial assistance: A. M. Mannino Editorial secretariat: V. Spadaro & P. Campisi Layout & Tecnical editing: E. Di Gristina & F. La Sorte Design: V. Magro & L. C. Raimondo Redazione di "Flora Mediterranea" Herbarium Mediterraneum Panormitanum, Università di Palermo Via Lincoln, 2 I-90133 Palermo, Italy [email protected] Printed by Luxograph s.r.l., Piazza Bartolomeo da Messina, 2/E - Palermo Registration at Tribunale di Palermo, no. 27 of 12 July 1991 ISSN: 1120-4052 printed, 2240-4538 online DOI: 10.7320/FlMedit26.001 Copyright © by International Foundation pro Herbario Mediterraneo, Palermo Contents V. Hugonnot & L. Chavoutier: A modern record of one of the rarest European mosses, Ptychomitrium incurvum (Ptychomitriaceae), in Eastern Pyrenees, France . 5 P. Chène, M.
    [Show full text]
  • Samenkatalog Graz 2016.Pdf
    SAMENTAUSCHVERZEICHNIS Index Seminum Seed list Catalogue de graines des Botanischen Gartens der Karl-Franzens-Universität Graz Ernte / Harvest / Récolte 2016 Herausgegeben von Christian BERG, Kurt MARQUART & Jonathan WILFLING ebgconsortiumindexseminum2012 Institut für Pflanzenwissenschaften, Januar 2017 Botanical Garden, Institute of Plant Sciences, Karl- Franzens-Universität Graz 2 Botanischer Garten Institut für Pflanzenwissenschaften Karl-Franzens-Universität Graz Holteigasse 6 A - 8010 Graz, Austria Fax: ++43-316-380-9883 Email- und Telefonkontakt: [email protected], Tel.: ++43-316-380-5651 [email protected], Tel.: ++43-316-380-5747 Webseite: http://garten.uni-graz.at/ Zitiervorschlag : BERG, C., MARQUART, K. & Wilfling, J. (2017): Samentauschverzeichnis – Index Seminum – des Botanischen Gartens der Karl-Franzens-Universität Graz, Samenernte 2016. – 54 S., Karl-Franzens-Universität Graz. Personalstand des Botanischen Gartens Graz: Institutsleiter: Ao. Univ.-Prof. Mag. Dr. Helmut MAYRHOFER Wissenschaftlicher Gartenleiter: Dr. Christian BERG Gartenverwalter: Jonathan WILFLING, B. Sc. Gärtnermeister: Friedrich STEFFAN GärtnerInnen: Doris ADAM-LACKNER Viola BONGERS Magarete HIDEN Franz HÖDL Kurt MARQUART Franz STIEBER Ulrike STRAUSSBERGER Monika GABER Gartenarbeiter: Philip FRIEDL René MICHALSKI Oliver KROPIWNICKI Gärtnerlehrlinge: Gabriel Buchmann (1. Lehrjahr) Bahram EMAMI (3. Lehrjahr) Mario MARX (3. Lehrjahr) 3 Inhaltsverzeichnis / Contents / Table des matières Abkürzungen / List of abbreviations / Abréviations
    [Show full text]