Biodiversity Journal, 2020, 11 (2): 435–437 https://doi.org/10.31396/Biodiv.Jour.2020.11.2.435.437

Rediscovery of Volvulella volvulaeformis (G. Seguenza, 1879) in the Italian Pliocene ( Rhizoridae)

Mauro M. Brunetti

Calle Navas 106, 14511 Navas del Selpillar, Spain

ABSTRACT A fossil gastropod for the Italian Pliocene is here reported. Following bibliographical research, it is assigned to a taxon previously described and never cited: Cylichna volvulaeformis G. Seguenza, 1880, which, according to its morphological characteristics, is assigned to the genus Volvulella Newton, 1891 (Gastropoda Rhizoridae).

KEY WORDS Gastropoda; Rhizoridae; Pliocene.

Received 12.01.2020; accepted 19.04.2020; published online 29.05.2020

INTRODUCTION RESULTS

Malacological research carried in the Italian Systematics Pliocene led to the discovery of some specimens of a Rhizoridae Dell, 1952, previously described as Classis GASTROPODA Cuvier, 1795 Volvulella sp. in Brunetti & Cresti, 2019. The sub- Ordo P. Fischer, 1883 sequent examination of numerous literature led to Superfamilia BULLOIDEA Gray, 1827 the determination of the specimens examined as Familia RHIZORIDAE Dell, 1952 Cylichna volvulaeformis G. Seguenza, 1880, de- Genus Volvulella Newton, 1891 scribed by the author for the Pliocene of Calabria, type species: acuminata Bruguiere, 1792 which is now assigned to the genus Volvulella New- ton, 1891. Volvulella volvulaeformis (G. Seguenza, 1879) (Figs. 1, 2)

MATERIAL AND METHODS Cylichna volvulaeformis G. Seguenza, 1880: p. 252, tav. 16, fig. 11. Volvulella sp. Brunetti & Cresti, 2019: p. 116, fig. The examined material was collected manually in 520. the ground, on the surface. The following abbrevi- ORIGINAL DIAGNOSIS. G. Seguenza, 1880: “17. ations are used in the text: H = total height of the *C. volvulaeformis n. sp. Tav. XVI. fig. 11. Testa shell; CCM = Cresti Massimo collection (Florence, ovata, antice rotundata, postice subacuta: col- Italy). For suprageneric and generic attributions, umella subarcuata, fìsuram umbìlicalem con- the WoRMS was followed (http://www.marine- stituens: os strìctum, arcuatum, antice elargatum, species.org/). postice gradatim attenuatum”. 436 MAURO M. BRUNETTI

Figures 1, 2. Volvulella volvulaeformis (G. Seguenza, 1879). Fig. 1: original figure from Seguenza (1879), H = 5 mm. Fig. 2: Orciano Pisano (Pisa), Piacenzian, H = 3 mm (CCM). Figure 3. Volvulella acuminata (Bruguière, 1792) Orciano Pisano (Pisa), Piacenzian, H = 4 mm (CCM).

DESCRIPTION. Shell with maximum dimensions more evident at the ends, absent in the center of the <5 mm. Oval shape, with thin shell, convex sides, shell. pointed apex. The shell is formed entirely from the last whorl. Aperture rear wide, narrows to two REMARKS. I think it is interesting to propose the thirds of the whorl, with thin, sharp lip. Rounded author’s observations on this taxon. This species is front end, narrow navel, partially closed. Columel- very distinctive for its shape, which does not pre- lar callus slightly extended, there is a hint of plica sent any parting to the posterior region, and which inside. Sculpture composed of thin spiral grooves instead protrudes and is almost acute. It appears to Rediscovery of Volvulella volvulaeformis (G. Seguenza, 1879) in the Italian Pliocene (Gastropoda Rhizoridae) 437

be carved from the mouth which continues to be neotype against this possibility if not in possession narrower and narrower until to the extreme part. of topotypical material, absent in this case. This character is found in the genus Volvula, and therefore this species is to be considered as a link between it and the Cylichna (G. Seguenza, 1879: ACKNOWLEDGEMENTS 253). This species is easily recognizable for the char- Special thanks to my friend Massimo Cresti (San acteristic oval shape at the base with the pointed Casciano V.P., Florence, Italy) for the material made apex. The most similar species is Volvulella acumi- available. nata (Bruguière, 1792) (Fig. 3) which differs from V. volvulaeformis (G. Seguenza, 1879) for the shape of the narrowest shell, the thinnest and most pointed REFERENCES apex, the different shape of the lip. The species had recently been iconographed as Volvulella sp. by Bertolaso L. & Palazzi S., 2000. Note sulla raccolta Brunetti & Cresti (2019). Seguenza di molluschi plio-pleistocenici della The material examined consists of 6 specimens provincia di Messina conservata presso il Museo di Geologia e paleontologia dell'Università di Firenze. from the Piacenziano of Orciano Pisano (Pisa). An- Bollettino Malacologico, 35: 3-44. other specimen, unfortunately destroyed, was found Brunetti M.M. & Cresti M., 2018. I fossili di Orciano by the writer in the middle Pliocene of the Rio Pisano, Atlante Iconografico. Edizioni Danaus, Crevalese (Piacenza). The specimen shown by G. Palermo, 228 pp. Seguenza (1879) (Fig. 1) had been found in the lo- La Perna R. & D’Abramo M., 2010. Una collezione di cality “Astian” of Gallina (Reggio Calabria) from G. Seguenza conservata presso l’Istituto Tecnico which most of the species established by G. Commerciale “O.G. Costa” di Lecce. Bollettino Seguenza came, unfortunately destroyed due to the Malacologico, 46: 29-35. Seguenza G., 1879. Le formazioni terziarie della provin- Messina earthquake of 1901. The locality “Astian” cia di Reggio (Calabria). Memorie della Reale Ac- of Gallina includes a chronostratigraphic interval cademia dei Lincei, Classe di Scienze Fisiche, that varies from late Pliocene to the early Pleistocene Matematiche e Naturalistiche, 3, 6: 1-446. (La Perna & d’Abramo 2010). We agree with Berto- WoRMS, 2020. WoRMS, World Register Marine laso & Palazzi (2000) whether or not to establish a Species: www.marinespecies.org .