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Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea
Brigham Young University BYU ScholarsArchive Theses and Dissertations 2018-07-01 Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco Brigham Young University Follow this and additional works at: https://scholarsarchive.byu.edu/etd Part of the Life Sciences Commons BYU ScholarsArchive Citation Pacheco, Yelena Marlese, "Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea" (2018). Theses and Dissertations. 7444. https://scholarsarchive.byu.edu/etd/7444 This Thesis is brought to you for free and open access by BYU ScholarsArchive. It has been accepted for inclusion in Theses and Dissertations by an authorized administrator of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco A thesis submitted to the faculty of Brigham Young University in partial fulfillment of the requirements for the degree of Master of Science Michael F. Whiting, Chair Sven Bradler Seth M. Bybee Steven D. Leavitt Department of Biology Brigham Young University Copyright © 2018 Yelena Marlese Pacheco All Rights Reserved ABSTRACT Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco Department of Biology, BYU Master of Science Phasmatodea exhibit a variety of cryptic ecomorphs associated with various microhabitats. Multiple ecomorphs are present in the stick insect fauna from Papua New Guinea, including the tree lobster, spiny, and long slender forms. While ecomorphs have long been recognized in phasmids, there has yet to be an attempt to objectively define and study the evolution of these ecomorphs. -
Phasmid Studies
ISSN 0966-0011 PHASMID STUDIES. volume 13, numbers 1 & 2. September 2005. Editor: P.E. Bragg. Published by the Phasmid Study Group. Phasmid Studies ISSN 0966-0011 volume 13, numbers I & 2. Contents Phasmids from Sabah Robert Bradburne I A reassessment of some Bornean Lonchodinae and Aschiphasmalidae, with some lectotype designations, new synonyms, and the description of (WO new species P.E. Bragg ................ ........ .. 11 Hap/opus Burmeisler, 1838, replacement name for Aplopus Gray, 1835 (Phasmalodea). Oliver Zornpro ... .. 30 A new species of the genus Baculofraclum. the first record of the genus from Borneo. P.E. Bragg .. ............................. .. 31 Reviews and Abstracts Phasmid Abstracts 38 Cover illustration: Female Parafoxopsis kQrySll!.~ Gilmher, 1932 by r.E. Bragg. Br.dburn, R. (2005) Phasmid Studies, 13(1&2): \-10. - Phasmids from Sabah Robert Bradburne, 26 Royal Avenue, Tonbridge, Kent, TN9 208, UK. Abstract This paper describes a trip (Q six locations in Sabah, Borneo, during October 2003. A 101:11 of around 20 species of stick insects were found al four of these locations, including an undescribed species found at 3300m on Moum Kinabalu. The most commonly encountered species in the lowland forest were Lonchodes spp., Haaniella spp., and Asceles margarilatus. Key words Phasmida, Borneo, Sabah, Sukau, Kinabalu, Danum Valley, Haaniella, Asceles, Prosemoria, Necroscia, Presbistus, Carausius, PhellQcephoms, Dinophasmo. Introduction In October 2003 I travelled to Sabah in North Borneo to spend [WO weeks searching for the wildlife of the region. Our group stayed in six locations, four of which yielded many species of phasmid. The rainy season had started early and therefore it frequently rained all afternoon, and often into the night. -
Belalong Forest: Anarchoides Iyralus (A Male) Was Taken on Bukit Belalong
B A T R E 0 p I L C A A L L R A I By 0 N F Earl of Cranbrook 0 N R David SEdwards E G~ With other contributions by George Argent Charles M Francis Penroose Saleha Martin Barker Freddy Gathome-Hardy Axel Dalberg Poulsen Roger A Beaver David T lanes Klaus Riede Peter F Becker Kamariah Abu Salim Gaden S Robinson Peter S Bellwood Chris J Kofron Sheila M Ross Elizabeth L Bennett Ruth Levy David Roubik Webber E Booth A Q Malik Samhan Nyawa Philip E Bragg C1iveF Mann Martin Sands Dielrich Burkhardl Nick Mawdsley Haji Serudin Tinggal loe K Charles lason, Lord Medway Gavin C Smith Salish C Choy Tom C Mitchell Rowland Snazell Mark A Cook Jonathan Moran Brian M Spooner Ind rane iI Das D Neil Morgan Mary C Slockdale Martin Drake Ingrid de la Motte Peter Wilkie John Dransfield Mark Mulligan Yanilla Willcox Soejatmi Drullsfield Nico Nieser John TWills Alan P Dykes Alben G Orr Yong Nget Yung Peter P Eaton Colin A Pendry Twenty-nine species of Phasmida have been collected in the Belalong forest: Anarchoides Iyralus (a male) was taken on Bukit Belalong . In the daily cycle of changing sounds in the abdomen, and stabbing or pinching Belalong, many noises are produced by movements of the powerfully spined hind insects, their chief purposes to attract po legs. Two species of leaf insects (Phyllium) tential mates and/or advertise territory. recorded from Borneo produce sounds by Orthoptera: Many Orthoptera produce rubbing together the thickened third seg noises, using a great variety of sound ments of their antennae. -
The Phasmid Study Group Newsletter No
The Phasmid Study Group Newsletter No. 122 June 2010 ISSN 0268-3806 Male Brasidas foveolatus with spermatophores. PSG SUMMER MEETING, Saturday, 10th JULY 2010.........................................................................2 News, Information & Updates ......................................................................................................................3 The Committee..........................................................................................................................................3 Diary Dates................................................................................................................................................3 Articles, Reviews & Submissions.................................................................................................................3 Book Review: Big Bugs Life~size...........................................................................................................3 Book Announcement:................................................................................................................................4 Mantids of the Euro-Mediterranean Area .................................................................................................4 Culture survey 2010 ..................................................................................................................................5 Book Announcement: Silent Summer: The State of Wildlife in Britain and Ireland ..............................9 Spermatophores from PSG 301 Brasidas -
Evolution of Flight Morphology in Stick Insects
bioRxiv preprint doi: https://doi.org/10.1101/774067; this version posted October 23, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY-NC-ND 4.0 International license. 1 2 A tale of winglets: evolution of flight morphology in stick insects 3 4 Yu Zeng1,2,†, Conner O’Malley1, Sonal Singhal1,3, Faszly Rahim4,5, 5 Sehoon Park1, Xin Chen6,7, Robert Dudley1,8 6 7 1Department of Integrative Biology, University of California, Berkeley, CA 92870, 8 USA 9 2Schmid College of Science and Technology, Chapman University, Orange, CA 10 92866, USA 11 3 Department of Biology, CSU Dominguez Hills, Carson, CA 90747 USA 12 4Islamic Science Institute, Universiti Sains Islam Malaysia, 71800 Bandar Baru 13 Nilai, Negeri Sembilan, Malaysia 14 5Centre for Insect Systematics, Universiti Kebangsaan Malaysia, 43600 Bangi, 15 Selangor, Malaysia 16 6Department of Biology, The College of Staten Island, The City University of New 17 York, NY 10314, USA 18 7Department of Biology, The Graduate School and University Center, The City 19 University of New York, NY 10016, USA 20 8Smithsonian Tropical Research Institute, Balboa, 21 Republic of Panama 22 23 †Corresponding author: [email protected] 24 25 1 bioRxiv preprint doi: https://doi.org/10.1101/774067; this version posted October 23, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. -
Insect Egg Size and Shape Evolve with Ecology but Not Developmental Rate Samuel H
ARTICLE https://doi.org/10.1038/s41586-019-1302-4 Insect egg size and shape evolve with ecology but not developmental rate Samuel H. Church1,4*, Seth Donoughe1,3,4, Bruno A. S. de Medeiros1 & Cassandra G. Extavour1,2* Over the course of evolution, organism size has diversified markedly. Changes in size are thought to have occurred because of developmental, morphological and/or ecological pressures. To perform phylogenetic tests of the potential effects of these pressures, here we generated a dataset of more than ten thousand descriptions of insect eggs, and combined these with genetic and life-history datasets. We show that, across eight orders of magnitude of variation in egg volume, the relationship between size and shape itself evolves, such that previously predicted global patterns of scaling do not adequately explain the diversity in egg shapes. We show that egg size is not correlated with developmental rate and that, for many insects, egg size is not correlated with adult body size. Instead, we find that the evolution of parasitoidism and aquatic oviposition help to explain the diversification in the size and shape of insect eggs. Our study suggests that where eggs are laid, rather than universal allometric constants, underlies the evolution of insect egg size and shape. Size is a fundamental factor in many biological processes. The size of an 526 families and every currently described extant hexapod order24 organism may affect interactions both with other organisms and with (Fig. 1a and Supplementary Fig. 1). We combined this dataset with the environment1,2, it scales with features of morphology and physi- backbone hexapod phylogenies25,26 that we enriched to include taxa ology3, and larger animals often have higher fitness4. -
The Pregenital Abdominal Musculature in Phasmids and Its Implications for the Basal Phylogeny of Phasmatodea (Insecta: Polyneoptera) Rebecca Klugã, Sven Bradler
ARTICLE IN PRESS Organisms, Diversity & Evolution 6 (2006) 171–184 www.elsevier.de/ode The pregenital abdominal musculature in phasmids and its implications for the basal phylogeny of Phasmatodea (Insecta: Polyneoptera) Rebecca KlugÃ, Sven Bradler Zoologisches Institut und Museum, Georg-August-Universita¨tGo¨ttingen, Berliner Str. 28, 37073 Go¨ttingen, Germany Received 7 June 2005; accepted 25 August 2005 Abstract Recently several conflicting hypotheses concerning the basal phylogenetic relationships within the Phasmatodea (stick and leaf insects) have emerged. In previous studies, musculature of the abdomen proved to be quite informative for identifying basal taxa among Phasmatodea and led to conclusions regarding the basal splitting events within the group. However, this character complex was not studied thoroughly for a representative number of species, and usually muscle innervation was omitted. In the present study the musculature and nerve topography of mid-abdominal segments in both sexes of seven phasmid species are described and compared in detail for the first time including all putative basal taxa, e.g. members of Timema, Agathemera, Phylliinae, Aschiphasmatinae and Heteropteryginae. The ground pattern of the muscle and nerve arrangement of mid-abdominal segments, i.e. of those not modified due to association with the thorax or genitalia, is reconstructed. In Timema, the inner ventral longitudinal muscles are present, whereas they are lost in all remaining Phasmatodea (Euphasmatodea). The ventral longitudinal muscles in the abdomen of Agathemera, which span the whole length of each segment, do not represent the plesiomorphic condition as previously assumed, but might be a result of secondary elongation of the external ventral longitudinal muscles. -
Fam: Bacillidae, Suborden: Areolatae, Orden: Phasmida
Fásmidos espinosos. La Familia Heteropterygidae ( orden: Phasmatodea, suborden: Areolatae, Zompro 2005) Por Sergi Romeu 1- Introducción: En esta familia Heteropterygidae encontramos los insectos más peculiares que podemos imaginarnos, llenos de espinas por todo el cuerpo y con un camuflaje de formas y colores típico del hábitat de sotobosque de las selvas húmedas. Hojas secas, líquenes, musgos, cortezas, pequeñas ramas, brotes, astillas...toman vida al intentar leerlos en este artículo. Principalmente estamos hablando de especies de distribución Asiática presentes en Malaysia, Sumatra, Borneo y muchas otras islas de Indonesia. 2- Clasificación: Durante los últimos años, varios autores han estudiado la sistemática del orden phasmatodea. Principalmente se trata de revisiones teóricas, basadas en descripciones de los ejemplares tipo depositados en los museos de todo el mundo. Paul Brock trata el grupo que nos interesa dentro la familia Bacillidae, como una sub-familia llamada Heteropteryginae, dividiéndola a su vez en cuatro tribus: Datamini, Anisacanthini, Obrimini y Heteropterygini. La mayoría de especies de esta familia Bacillidae no tienen alas, exceptuando algunas especies con rudimentos alares o alas reducidas dentro de nuestra sub-familia Heteropteryginae. Desde la familia Bacillidae, la clave taxonómica para llegar a la sub-familia Heteropteryginae es según P. Brock (1999): - 1) Antena mas larga que el fémur delantero. Alados o sin alas, pero nunca presentes en África y Europa........................................................................................................................................................2 -
©Zoologische Staatssammlung München;Download: Http
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Spixiana, Zeitschrift für Zoologie Jahr/Year: 1994 Band/Volume: 017 Autor(en)/Author(s): Carlberg Ulf Artikel/Article: Bibliography of Phasmida (Insecta). VII. 1985-1989 179- 191 ©Zoologische Staatssammlung München;download: http://www.biodiversitylibrary.org/; www.biologiezentrum.at SPIXIANA ©Zoologische Staatssammlung München;download: http://www.biodiversitylibrary.org/; www.biologiezentrum.at Allred, M. L., Stark, B. P. & Lentz, D. L. 1986. Egg capsule morphology of Anisomorpha buprestoides (Phasmatodea: Pseudophasmatidae). - Ent. News 97: 169-174 Baccetti, B. 1985. Evolution of the sperm cell. In: Metz, C. B. & Monroy, A. (Eds.), Biology of Fertilization, vol. 2, pp. 3-58. New York (Academic Press) - - 1987a. Spermatozoa and stick insect phylogeny. - In: Mazzini & Scali (Eds.) 1987: 177-123 - - (Ed.) 1987b. Evolutionary Biology of Orthopteroid Insects. Chichester (EUis Horwood), 1-612 pp. - - 1987c. Spermatozoa and phylogeny in orthopteroid insects. - In: Baccetti (Ed.) 1987c: 12-112 Bart, A. 1988. Proximal leg regeneration in Cmmisius morosus: growth, intercalation and proximaliza- tion. - Development 102: 71-84 Bässler, U. 1985. Proprioceptive control of stick insect Walking. - In: Gewecke & Wendler (Eds.) 1985: 43-48 - - 1986a. On the definition of central pattern generator and its sensory control. - Biol. Cybern. 54: 65-69 - - 1986b. Afferent control of Walking movements in the stick insect C/;n/af/fna impigra. 1. Decerebrated - 345-349 animals on a treadband. J. Comp Physiol. A 158: - - - 1986c. Ibid. 11. Reflex reversal and the release of the swing phase in the restrained foreleg. J. Comp. Physiol. A 158: 351-362 - - 1987a. Timing and shaping influences on the motor Output for Walking in stick insects. -
As Pests of Agriculture and Forestry, with a Generalised Theory of Phasmid Outbreaks Edward Baker*
Baker Agric & Food Secur (2015) 4:22 DOI 10.1186/s40066-015-0040-6 REVIEW Open Access The worldwide status of phasmids (Insecta: Phasmida) as pests of agriculture and forestry, with a generalised theory of phasmid outbreaks Edward Baker* Abstract Stick insects have been reported as significant phytophagous pests of agricultural and timber crops since the 1880s in North America, China, Australia and Pacific Islands. Much of the early literature comes from practical journals for farmers, and even twentieth Century reports can be problematic to locate. Unlike the plaguing Orthoptera, there has been no synthesis of the pest status of this enigmatic order of insects. This paper provides a literature synthesis of those species known to cause infestation or that are known to damage plants of economic importance; summarises historical and modern techniques for infestation management; and lists known organisms with potential for use as biological control agents. A generalised theory of outbreaks is presented and suggestions for future research efforts are made. Keywords: Pests, Infestation, Agriculture, Forestry Background a significant factor in the scale of phasmid outbreaks— in most species, females lay several hundred eggs [6]. In “The unexampled multiplication and destructive- addition, their wasteful eating habits [7] and their often ness of this insect at Esperance farm is but one of the rapid growth [8] means they consume a large quantity many illustrations of the fact, long since patent to all of vegetation [9]. Considerable efforts have been put close students of economic entomology, that species into controlling the three species of Australian phasmid normally harmless may suddenly become very inju- known to cause periodic infestation [10]. -
Egg Database Bibliography
Egg Database Bibliography Samuel H. Church1;∗;y, Seth Donoughe1;2;∗, Bruno A. S. de Medeiros1, Cassandra G. Extavour1;3;y Note: This document is a list of the 1,756 published sources that were used to generate the assembled dataset of insect egg traits. ‘Diss.’ indicates a PhD dissertation, whereas ‘MA thesis’ indicates a Master’s thesis. For more information on the dataset, please see Church et al. 2018: “A database of egg size and shape from more than 6,700 insect species” (preprint). It describes the criteria that were used to include sources, definitions of each trait, and details on the procedure that was used to collect data from each source. References Abbassy, M. M., N. Helmy, M. Osman, S. E. Cope, and S. M. Presley. “Embryogenesis of the sand fly Phlebotomus pa- patasi (Diptera: Psychodidae): cell cleavage, blastoderm formation, and gastrulation”. Annals of the Entomological Society of America 88.6 (1995): 809–814. Abdel-Razak, S. I., S. M. Beshr, A. K. Mourad, and K. S. Moursi. “Ultrastructure of egg shell of four different Coccoidea species in Egypt”. Communications in Agricultural and Applied Biological Sciences 73.3 (2007): 521– 527. Abdurahiman, U. C. and K. J. Joseph. “Observations on the biology and behaviour of Ceratosolen marchali Mayr (Agaonidae, Chalcidoidea, Hymenoptera)”. Entomon 1.2 (1976): 115–122. Abraham, Y. J., D. Moore, and G. Godwin. “Rearing and aspects of biology of Cephalonomia stephanoderis and Prorops nasuta (Hymenoptera: Bethylidae) parasitoids of the coffee berry borer, Hypothenemus hampei (Coleoptera: Scolytidae)”. Bulletin of Entomological Research 80.2 (1990): 121–128. Adamski, D., J. -
Phasmid Studies, 2(1&2)
ISSN 0966-0011 PHASMID STUDIES. volume 2, numbers 1 & 2. June & December 1993. Editor: P.E. Bragg. Published by the Phasmid Study Group. Phasmid Studies ISSN 0966-0011 volume 2, numbers 1 & 2. Contents The World of Stick and Leaf-Insects in Books, some general remarks Paul D . Brock . A very pretty phasmid : Parectatosoma hystrix J. Roget . 7 Looking at Baculum eggs John Sellick . .. 10 Keeping and breeding Haani ella species successfully Ian Abercrombie . 14 List of stick and leaf-insect (Phasmatod ea = Phasmida) type material in the Natural History Museum, published since Kirby's 1904 Cat alogue Paul D. Brock . .. 17 A survey of the distribution of the unarmed stick insect Acanthoxyla inermis in Port Gaverne and Port Isaac, North Cornwall in 1992 Malcolm Lee. .. 25 The Phasmid Database: changes to version 1 P.E . Bragg 33 Reviews and Abstracts Phasmid Abstracts . .. 35 PSG 121 , Phenac ephorus spinulosus (Haus1eithner) P .E . Bragg . .. 41 Pharnacia serratipes (Gray) Frank Hennemann . ... 45 Phena cocephalus coronatus Werner P.E. Bragg . 51 The leaf-piercing eggs of Asceles John Sellick . 54 Defensive and flying behaviour in Sipyloidea sp. (PSG 103) R .P. Bradburne 56 A new Libethra from Ecuador Wim Potvin . 59 Some notes on Din ophasma gutti gera (Westwood) from Borneo P.E. Bragg . 62 Revi ews and Abstracts Pha smid Abstracts . 66 Publications not ed . 67 Cover illustration: Female Phenacephorus spinulosus (Hausleithner) by P.E . Bragg. The World of Stick and Leaf-Insects in Books, some general remarks. Paul D. Brock, "Papillon , 40, Thorndike Road , Slough, SU ISR. UK. Key words Stick and Leaf Insects, Books.