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Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea
Brigham Young University BYU ScholarsArchive Theses and Dissertations 2018-07-01 Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco Brigham Young University Follow this and additional works at: https://scholarsarchive.byu.edu/etd Part of the Life Sciences Commons BYU ScholarsArchive Citation Pacheco, Yelena Marlese, "Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea" (2018). Theses and Dissertations. 7444. https://scholarsarchive.byu.edu/etd/7444 This Thesis is brought to you for free and open access by BYU ScholarsArchive. It has been accepted for inclusion in Theses and Dissertations by an authorized administrator of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco A thesis submitted to the faculty of Brigham Young University in partial fulfillment of the requirements for the degree of Master of Science Michael F. Whiting, Chair Sven Bradler Seth M. Bybee Steven D. Leavitt Department of Biology Brigham Young University Copyright © 2018 Yelena Marlese Pacheco All Rights Reserved ABSTRACT Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco Department of Biology, BYU Master of Science Phasmatodea exhibit a variety of cryptic ecomorphs associated with various microhabitats. Multiple ecomorphs are present in the stick insect fauna from Papua New Guinea, including the tree lobster, spiny, and long slender forms. While ecomorphs have long been recognized in phasmids, there has yet to be an attempt to objectively define and study the evolution of these ecomorphs. -
Phasmid Studies 1 (June & December 1992)
PSG 118, Aretaon asperrimus (Redtenbacher) Paul Jennings, 14, Grenfell Avenue, Sunnybill, Derby, DE3 7JZ. U.K•• Taxonomic and distribution notes by P .E. Bragg. lliustration of male by E. Newman. Key words Phasmida, Aretaon asperrimus, Breeding, Rearing. Classification This species was originally described as Obrimus asperrimus by Redtenbacher in 1906. In 1938 Rehn & Rehn established the new genus Aretaon (1938: 419), with asperrimus as the type species. I have examined the type specimens of this species and A. muscosus Redtenbacher, which are all in Vienna, and I found that the type specimens of A. asperrimus are all adults while those of A. muscosus are all nymphs. A. muscosus is distinguished by having more prominent spines, particularly on the front tibiae and the top of the abdomen. However, having reared A. asperrimus it is clear that nymphs of this genus are very spiny and these spines in particular are reduced when the insect becomes adult. It is therefore quite likely that A. asperrimus and A. muscosus are the same species. This possibility was considered and rejected by Gunther (1935: 123) but as he had not reared them he would not have known that the spines are reduced when the insects become adult. The species in culture is clearly A. asperrimus, however it has smaller spines than the type specimens so clearly the species is variable. The type specimens of A. muscosus are much more spiny than those in culture, I cannot therefore be certain that A. asperrimus and A. muscosus are the same species. As there is a strong possibility that they are the same species, I am listing the published references and distribution records for both species. -
1 It's All Geek to Me: Translating Names Of
IT’S ALL GEEK TO ME: TRANSLATING NAMES OF INSECTARIUM ARTHROPODS Prof. J. Phineas Michaelson, O.M.P. U.S. Biological and Geological Survey of the Territories Central Post Office, Denver City, Colorado Territory [or Year 2016 c/o Kallima Consultants, Inc., PO Box 33084, Northglenn, CO 80233-0084] ABSTRACT Kids today! Why don’t they know the basics of Greek and Latin? Either they don’t pay attention in class, or in many cases schools just don’t teach these classic languages of science anymore. For those who are Latin and Greek-challenged, noted (fictional) Victorian entomologist and explorer, Prof. J. Phineas Michaelson, will present English translations of the scientific names that have been given to some of the popular common arthropods available for public exhibits. This paper will explore how species get their names, as well as a brief look at some of the naturalists that named them. INTRODUCTION Our education system just isn’t what it used to be. Classic languages such as Latin and Greek are no longer a part of standard curriculum. Unfortunately, this puts modern students of science at somewhat of a disadvantage compared to our predecessors when it comes to scientific names. In the insectarium world, Latin and Greek names are used for the arthropods that we display, but for most young entomologists, these words are just a challenge to pronounce and lack meaning. Working with arthropods, we all know that Entomology is the study of these animals. Sounding similar but totally different, Etymology is the study of the origin of words, and the history of word meaning. -
ZOOLOGY Zoology 110 (2007) 409–429
ARTICLE IN PRESS ZOOLOGY Zoology 110 (2007) 409–429 www.elsevier.de/zool Towards an 18S phylogeny of hexapods: Accounting for group-specific character covariance in optimized mixed nucleotide/doublet models Bernhard Misofa,Ã, Oliver Niehuisa, Inge Bischoffa, Andreas Rickerta, Dirk Erpenbeckb, Arnold Staniczekc aAbteilung fu¨r Entomologie, Zoologisches Forschungsmuseum Alexander Koenig, Adenauerallee 160, D-53113 Bonn, Germany bDepartment of Coelenterata and Porifera (Zoologisch Museum), Institute for Biodiversity and Ecosystem Dynamics, University of Amsterdam, P.O. Box 94766, 1090 GT Amsterdam, The Netherlands cStaatliches Museum fu¨r Naturkunde Stuttgart, Abt. Entomologie, Rosenstein 1, D-70191 Stuttgart, Germany Received 19 May 2007; received in revised form 2 August 2007; accepted 22 August 2007 Abstract The phylogenetic diversification of Hexapoda is still not fully understood. Morphological and molecular analyses have resulted in partly contradicting hypotheses. In molecular analyses, 18S sequences are the most frequently employed, but it appears that 18S sequences do not contain enough phylogenetic signals to resolve basal relationships of hexapod lineages. Until recently, character interdependence in these data has never been treated seriously, though possibly accounting for the occurrence of biased results. However, software packages are readily available which can incorporate information on character interdependence within a Bayesian approach. Accounting for character covariation derived from a hexapod consensus secondary structure model and applying mixed DNA/RNA substitution models, our Bayesian analysis of 321 hexapod sequences yielded a partly robust tree that depicts many hexapod relationships congruent with morphological considerations. It appears that the application of mixed DNA/RNA models removes many of the anomalies seen in previous studies. We focus on basal hexapod relationships for which unambiguous results are missing. -
Stick Insects Fact Sheet
Stick Insects Fact Sheet Female Titan Stick Insect. Image: QM, Jeff Wright. Introduction Biology Stick and leaf insects, scientifically known as phasmids, Females lay eggs one at a time, often with a flick of their are among the largest of all insects in the world. At 26 cm, abdomens to throw the egg some distance. An individual the Titan Stick Insect (Acrophylla titan) is the longest of female drops eggs at a rate of one to several per day and all Australian insects. Phasmids have perfected the art of she can produce between 100 and 1,300 eggs in her life- camouflage. Some resemble sticks and foliage so closely time. They fall to the ground and lie in the leaf litter. they even feature false buds, thorns and ragged leaf-like flanges. Small wonder they are rarely seen except after storms when they are blown out of threes and shrubs. Phasmids are sometimes confused with a different group of insects, the mantids. Also called Praying Mantids, these are predators with large, spiny front legs, held folded ready to strike and grasp prey. In contrast, Phasmids are herbivores (plant-eaters) with simple front legs that are similar in size and structure to their other legs. A variety of insect eggs. (on left). An ant carrying a stick insect egg (on right). Images: QM, Jeff Wright. All stick insects feed on fresh leaves. Some browse on a wide variety of trees and shrubs but others are fussy, eating only a limited range of host plants that are often closely Stick insect eggs are generally oval, and superficially seed- related to each other. -
Insects, Beetles, Bugs and Slugs of Mt Gravatt Conservation Reserve
Insects, beetles, bugs and slugs of Mt Gravatt Conservation Reserve Compiled by: Michael Fox www.megoutlook.org/flora-fauna/ © 2015-20 Creative Commons – free use with attribution to Mt Gravatt Environment Group Ants Dolichoderinae Iridomyrmex sp. Small Meat Ant Attendant “Kropotkin” ants with caterpillar of Imperial Hairstreak butterfly. Ants provide protection in return for sugary fluids secreted by the caterpillar. Note the strong jaws. These ants don’t sting but can give a powerful bite. Kropotkin is a reference to Russian biologist Peter Kropotkin who proposed a concept of evolution based on “mutual aid” helping species from ants to higher mammals survive. 4-Nov-20 Insects Beetles and Bugs - ver 5.9.docx Page 1 of 59 Mt Gravatt Environment Group – www.megoutlook.wordpress.com Insects, beetles, bugs and slugs of Mt Gravatt Conservation Reserve Formicinae Opisthopsis rufithorax Black-headed Strobe Ant Formicinae Camponotus consobrinus Banded Sugar Ant Size 10mm Eggs in rotting log 4-Nov-20 Insects Beetles and Bugs - ver 5.9.docx Page 2 of 59 Mt Gravatt Environment Group – www.megoutlook.wordpress.com Insects, beetles, bugs and slugs of Mt Gravatt Conservation Reserve Formicinae Camponotus nigriceps Black-headed Sugar Ant 4-Nov-20 Insects Beetles and Bugs - ver 5.9.docx Page 3 of 59 Mt Gravatt Environment Group – www.megoutlook.wordpress.com Insects, beetles, bugs and slugs of Mt Gravatt Conservation Reserve Formicinae Polyrhachis ammon Golden-tailed Spiny Ant Large spines at rear of thorax Nest 4-Nov-20 Insects Beetles and Bugs - ver 5.9.docx Page 4 of 59 Mt Gravatt Environment Group – www.megoutlook.wordpress.com Insects, beetles, bugs and slugs of Mt Gravatt Conservation Reserve Formicinae Polyrhachis australis Rattle Ant Black Weaver Ant or Dome-backed Spiny Ant Feeding on sugar secretions produced by Redgum Lerp Psyllid. -
Printed ISSN 0966-0011
Biographies of Phasmatologists – 2. George Robert Gray. P.E. Bragg, 8 The Lane, Awsworth, Nottinghamshire, NG16 2QP, U.K. Abstract George Robert Gray (1808-1872) was an English zoologist and author. His life and phasmid work is outlined. He described half the known species of phasmids at that time and is best known for his work on leaf insects and Australian phasmids. He also produced the first significant catalogue of world species. Key words Phasmida, Phasmatologist, G.R. Gray, Biography. George Robert Gray (1808-1872) George Robert Gray, an English zoologist and author, was born in Chelsea on 8th July 1808. He was born to a family of natural historians; his father, Samuel Gray, was a pharmacologist and botanist; his elder brother, John, went on to become Keeper of Zoology at the British Museum. In 1833 he was one of the founder members of the Entomological Society of London (now the Royal Entomological Society) and was Secretary in the first year. Gray started work at the British Museum in 1831 and went on to became Senior Assistant of the Zoology Department. He contributed the entomological section to an English edition of Cuvier's Animal Kingdom in which he described his first phasmid, Phyllium bioculatum in 1832. He began his museum work by cataloguing insects, and published an Entomology of Australia (1833). In addition to his work on phasmids, Gray described many species of Lepidoptera and his book Descriptions and Figures of some new Lepidopterous Insects chiefly from Nepal (1846) is considered an important work. Entomologists who are not familiar with Gray’s life story may be unaware that he is most famous for his work on birds; he did relatively little work on insects. -
Phasmid Studies
ISSN 0966-0011 PHASMID STUDIES. volume 13, numbers 1 & 2. September 2005. Editor: P.E. Bragg. Published by the Phasmid Study Group. Phasmid Studies ISSN 0966-0011 volume 13, numbers I & 2. Contents Phasmids from Sabah Robert Bradburne I A reassessment of some Bornean Lonchodinae and Aschiphasmalidae, with some lectotype designations, new synonyms, and the description of (WO new species P.E. Bragg ................ ........ .. 11 Hap/opus Burmeisler, 1838, replacement name for Aplopus Gray, 1835 (Phasmalodea). Oliver Zornpro ... .. 30 A new species of the genus Baculofraclum. the first record of the genus from Borneo. P.E. Bragg .. ............................. .. 31 Reviews and Abstracts Phasmid Abstracts 38 Cover illustration: Female Parafoxopsis kQrySll!.~ Gilmher, 1932 by r.E. Bragg. Br.dburn, R. (2005) Phasmid Studies, 13(1&2): \-10. - Phasmids from Sabah Robert Bradburne, 26 Royal Avenue, Tonbridge, Kent, TN9 208, UK. Abstract This paper describes a trip (Q six locations in Sabah, Borneo, during October 2003. A 101:11 of around 20 species of stick insects were found al four of these locations, including an undescribed species found at 3300m on Moum Kinabalu. The most commonly encountered species in the lowland forest were Lonchodes spp., Haaniella spp., and Asceles margarilatus. Key words Phasmida, Borneo, Sabah, Sukau, Kinabalu, Danum Valley, Haaniella, Asceles, Prosemoria, Necroscia, Presbistus, Carausius, PhellQcephoms, Dinophasmo. Introduction In October 2003 I travelled to Sabah in North Borneo to spend [WO weeks searching for the wildlife of the region. Our group stayed in six locations, four of which yielded many species of phasmid. The rainy season had started early and therefore it frequently rained all afternoon, and often into the night. -
Phasmida (Stick and Leaf Insects)
● Phasmida (Stick and leaf insects) Class Insecta Order Phasmida Number of families 8 Photo: A leaf insect (Phyllium bioculatum) in Japan. (Photo by ©Ron Austing/Photo Researchers, Inc. Reproduced by permission.) Evolution and systematics Anareolatae. The Timematodea has only one family, the The oldest fossil specimens of Phasmida date to the Tri- Timematidae (1 genus, 21 species). These small stick insects assic period—as long ago as 225 million years. Relatively few are not typical phasmids, having the ability to jump, unlike fossil species have been found, and they include doubtful almost all other species in the order. It is questionable whether records. Occasionally a puzzle to entomologists, the Phasmida they are indeed phasmids, and phylogenetic research is not (whose name derives from a Greek word meaning “appari- conclusive. Studies relating to phylogeny are scarce and lim- tion”) comprise stick and leaf insects, generally accepted as ited in scope. The eggs of each phasmid are distinctive and orthopteroid insects. Other alternatives have been proposed, are important in classification of these insects. however. There are about 3,000 species of phasmids, although in this understudied order this number probably includes about 30% as yet unidentified synonyms (repeated descrip- Physical characteristics tions). Numerous species still await formal description. Stick insects range in length from Timema cristinae at 0.46 in (11.6 mm) to Phobaeticus kirbyi at 12.9 in (328 mm), or 21.5 Extant species usually are divided into eight families, in (546 mm) with legs outstretched. Numerous phasmid “gi- though some researchers cite just two, based on a reluctance ants” easily rank as the world’s longest insects. -
Goliath Stick Insect
Husbandry Manual for Eurycnema goliath Tara Bearman Husbandry Manual for Goliath Stick Insect Eurycnema goliath (Gray, 1834) (Insecta: Phasmatidae) Compiled by: Tara Bearman Date of Preparation: 2007 Western Sydney Institute of TAFE, Richmond Course Name and Number: 1068 Certificate III –Captive Animals Lecturer: Graeme Phipps 1 Husbandry Manual for Eurycnema goliath Tara Bearman TABLE OF CONTENTS 1 INTRODUCTION .............................................................................................................................. 6 2 TAXONOMY ....................................................................................................................................... 7 2.1 NOMENCLATURE............................................................................................................................. 7 2.2 SUBSPECIES................................................................................................................................... 7 2.3 RECENT SYNONYMS ....................................................................................................................... 7 2.4 OTHER COMMON NAMES ................................................................................................................ 7 3 NATURAL HISTORY........................................................................................................................ 8 3.1 MORPHOMETRICS........................................................................................................................... 9 3.1.1 Mass and -
Phasmid Studies ISSN 0966-0011 Volume 9, Numbers 1 & 2
Phasmid Studies ISSN 0966-0011 volume 9, numbers 1 & 2. Contents Species Report PSG. 122, Anisomorpha monstrosa Hebard Paul A. Hoskisson . 1 Cigarrophasma, a new genus of stick-insect (Phasmatidae) from Australia Paul D. Brock & Jack Hasenpusch . 0 •••••• 0 ••• 0 ••••••• 4 A review of the genus Medaura Stal, 1875 (Phasmatidae: Phasmatinae), including the description of a new species from Bangladesh Paul Do Brock & Nicolas Cliquennois 11 First records and discovery of two new species of Anisomorpha Gray (Phasmida: Pseudophasmatidae) in Haiti and Dominican Republic Daniel E. Perez-Gelabert 0 .. .. 0 • • • • • • 0 • • • • 0 • • 0 • 0 • • 0 0 • • • 27 Species report on Pharnacia biceps Redtenbacher, PSG 203 Wim Potvin 0 ••• 28 How Anisomorpha got its stripes? Paul Hoskisson . 33 Reviews and Abstracts Book Reviews . 35 Phasmid Abstracts 38 Cover illustr ation : Orthonecroscia pulcherrima Kirby, drawing by PoE. Bragg. Species Report PSG. 122, Anisomorpha monstrosa Hebard Paul A. Hoskisson, School of Biomolecular Sciences, Liverpool John Moores University, Byrom Street, Liverpool, 13 3AF, UK. With illustrations by P.E. Bragg. Abstract This report summarises the care and breeding of Anisomorpha monstrosa Hebard, the largest species in the genus. Behaviour and defence mechanism are also discussed along with descriptions of the eggs, nymphs, and adults. Key words Phasmida, Anisomorpha monstrosa, Pseudophasmatinae, Rearing, Distribution, Defence. Taxonomy Anisomorpha monstrosa belongs to the sub-family Pseudophasmatinae. It was described in 1932 by Hebard (1932: 214) and is the largest species in the genus. The type specimen is a female collected from Merida, in Yucatan, Mexico. Culture History The original culture of this species was collected in Belize, approximately 150km north of Belize City by Jan Meerman in 1993 or 1994 (D'Hulster, personal communication). -
The Phasmid Study Group Is Invited to Exhibit at Some Show S
/ ,0 ,0 ,0 («i |0 ,« (V ,» ,\\\\\\ t\\* ,» |«. ,* !«. ,S ,\\\\\\\\» ,S ,\\1 f% t«. t0 ,\ ,\\ »K f* f* rSSfe ^ £ * I' > f The Phasmid ... Study Group .V CHAIRMAN: Judith Marshall. - Dept. of Entomology, The Natural History Museum, Cromwell Road, London SW7 5BD. (Tel: 020 7942 5610; FAX 020 7942 5229) E-mail: [email protected]. TREASURER/MEMBERSHIP: Paul Brock. "Papillon", 40 Thorndike Road, Slough, Berks. SL2 1SR. (Tel: 01753 579447) E-mail: Not currently available on e-mail i' SECRETARY: Phil Bragg. 8 The Lane, Awsworth, Nottinghamshire, NG16 2QP. (Tel: 0115 9305010). Pictures as featured inside this Newsletter!! DECEMBER 2002 NEWSLETTER No 93 ISSN 0268-3806 i' Newsletter 93.1 „ —. yr» k<t to-M> k«* »"«* p-M> =«• + ~ + »•* »''* ^"^ K"» ».-*»> p4* #."** ^« t\0 ,V (l ,» f\X t* «S |0 (|» |\» |«> ««. ,1 ,1 »V ,1 |% |S t\N »% «% |1 |» |» |0 «% |1 f H f1 |* f\» f<l |» |0 |0 f\0 «% |* ([I Diary Dates 2003 January 18th; Saturday, 11.30 am F*SG AGM zyxwvutsrqponmlkjihgfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA& Winter Meeting Spencer Gallery, Natural History Museum, Cromwell Road, London. [Note the change of room, re refurbishment]. February 9th; Sunday Bugs & Beasties Show Selby, North Yorkshire (Contact Steve, www.bugsnstuff.co.uk). NB Steve is looking for some Phasmid Exhibitors — can you help? April 6th; Sunday, 10.30 am Spring Entomological Show (formerly Kettering Show) Kettering Leisure Village (Contact Jack Harris, 01455 444792). [It's like a mini AES Exhib, Ed.] British Tarantula Society Exhibition May 2003 — more details in next Newsletter Nature Matters Event June 2003 — Has anyone any details on this? F*SG Summor Meeting July 2003 - more details after the AGM.