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A Reassessment of the Phylogenetic Position of Cretaceous Sauropod Dinosaurs from Queensland, Australia
Asociación Paleontológica Argentina. Publicación Especial 7 ISSN 0328-347X VII International Symposium on Mesozoic Terrestrial Ecosystems: 139-144.Buenos Aires, 30-6-2001 A ReasSessmenT of the phylogenetic position of CretaceoUS SaUROPOd dinosaURS from Queensland, Australia Ralph E. MOLNAR1 Abstract. The Cretaceous sauropod material from Queensland, Australia, has been regarded as pertaining to a persistently primitive sauropod lineage (e.g., Coombs and Molnar). The specimens derive from the Toolebuc and Allaru (Albian marine) and Winton (Cenomanian continental) Formations. Recent phyloge- netic analyses carried out by workers in Argentina, the USA and England permit a reassessment of this fragmentary material. As far as can be ascertained from the material, there is no indication from the char- acter states that more than a single taxon is represented. Character states diagnostic of the Titanosauriforrnes, the Titanosauria, the Somphospondyli and the Titanosauridae are present. Thus the Queensland material does not pertain to cetiosaurids but belongs to titanosaurs, extending their range in- to Australia Key words. Sauropods. Austrosaurus. Titanosaurs. Cretaceous. Paleozoogeography. IntroductioN (1998),has made it possible to reassess the phyloge- netic affinities of the Australian Cretaceous sauropod By the 1950's titanosaurs were widely recognized material and address the anomalous absence of ti- both as the latest sauropod group to diversify and as tanosaurs. This paper looks specifically at pre-eminently the sauropods of Gondwanaland. -
Dinosaurs Largest Ornithopod
Leaellynasaura amicagraphica (1989) age: Early Cretaceous REGION: VIC SIZE: 1.5m A small ornithopod, this plant-eater lived in an Australia that was further south and partly within the Antarctic Circle. A well-preserved skull reveals it had a large brain and eyes, which helped it keep watch for predators as s it foraged for plants in the dark of the Antarctic winter. Muttaburrasaurus Dinosaur New research by Dr Matt Herne shows Leaellynasaura (‘lee-allin-ah-sore-ah’) had an extremely long tail. langdoni (1981) of Drs Tom Rich and Patricia Vickers-Rich named Australia the species after their daughter, Leaellyn. AGE: Early Cretaceous REGION: QLD/NSW SIZE: 8–9m Muttaburrasaurus (‘muta-burra-sore-rus’) is our A guide to the dinosaurs largest ornithopod. With one partial skeleton and a second skull from QLD and several teeth from Down Under, which ranged NSW, this powerful herbivore could rear-up on Austrosaurus its back legs to reach high foliage and intimidate predators, though it sometimes moved on four (1933) from ferocious carnivores to mckillopi legs too. It had an unusual bulge on its snout, which may have contained an inflatable air sac. herbivorous behemoths. age: Early Cretaceous REGION: QLD SIZE: 15–20m Discovered in north-central Queensland 80 ILLUSTRATIONS BY LIDA XING years ago, Austrosaurus (‘aus-tro-sore-us’) was our first known Cretaceous sauropod. This long-necked species was able to reach high foliage. Austrosaurus means ‘southern lizard’. Timimus hermani (1994) age: Early Cretaceous REGION: VIC SIZE: 3-5m The femur of Timimus (‘tim-my-mus’) is one of many specimens found at Dinosaur Cove by Drs Tom Rich and Patricia Vickers-Rich. -
Poropat Et Al 2017 Reappraisal Of
Alcheringa For Peer Review Only Reappraisal of Austro saurus mckillopi Longman, 1933 from the Allaru Mudstone of Queensland, Australia’s first named Cretaceous sauropod dinosaur Journal: Alcheringa Manuscript ID TALC-2017-0017.R1 Manuscript Type: Standard Research Article Date Submitted by the Author: n/a Complete List of Authors: Poropat, Stephen; Swinburne University of Technology, Department of Chemistry and Biotechnology; Australian Age of Dinosaurs Natural History Museum Nair, Jay; University of Queensland, Biological Sciences Syme, Caitlin; University of Queensland, Biological Sciences Mannion, Philip D.; Imperial College London, Earth Science and Engineering Upchurch, Paul; University College London, Earth Sciences, Hocknull, Scott; Queensland Museum, Geosciences Cook, Alex; Queensland Museum, Palaeontology & Geology Tischler, Travis; Australian Age of Dinosaurs Natural History Museum Holland, Timothy; Kronosaurus Korner <i>Austrosaurus</i>, Dinosauria, Sauropoda, Titanosauriformes, Keywords: Australia, Cretaceous, Gondwana URL: http://mc.manuscriptcentral.com/talc E-mail: [email protected] Page 1 of 126 Alcheringa 1 2 3 4 5 6 7 1 8 9 1 Reappraisal of Austrosaurus mckillopi Longman, 1933 from the 10 11 12 2 Allaru Mudstone of Queensland, Australia’s first named 13 14 For Peer Review Only 15 3 Cretaceous sauropod dinosaur 16 17 18 4 19 20 5 STEPHEN F. POROPAT, JAY P. NAIR, CAITLIN E. SYME, PHILIP D. MANNION, 21 22 6 PAUL UPCHURCH, SCOTT A. HOCKNULL, ALEX G. COOK, TRAVIS R. TISCHLER 23 24 7 and TIMOTHY HOLLAND 25 26 27 8 28 29 9 POROPAT , S. F., NAIR , J. P., SYME , C. E., MANNION , P. D., UPCHURCH , P., HOCKNULL , S. A., 30 31 10 COOK , A. G., TISCHLER , T.R. -
71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas. -
The Fragile Legacy of Amphicoelias Fragillimus (Dinosauria: Sauropoda; Morrison Formation – Latest Jurassic)
Volumina Jurassica, 2014, Xii (2): 211–220 DOI: 10.5604/17313708 .1130144 The fragile legacy of Amphicoelias fragillimus (Dinosauria: Sauropoda; Morrison Formation – latest Jurassic) D. Cary WOODRUFF1,2, John R. FOSTER3 Key words: Amphicoelias fragillimus, E.D. Cope, sauropod, gigantism. Abstract. In the summer of 1878, American paleontologist Edward Drinker Cope published the discovery of a sauropod dinosaur that he named Amphicoelias fragillimus. What distinguishes A. fragillimus in the annals of paleontology is the immense magnitude of the skeletal material. The single incomplete dorsal vertebra as reported by Cope was a meter and a half in height, which when fully reconstructed, would make A. fragillimus the largest vertebrate ever. After this initial description Cope never mentioned A. fragillimus in any of his sci- entific works for the remainder of his life. More than four decades after its description, a scientific survey at the American Museum of Natural History dedicated to the sauropods collected by Cope failed to locate the remains or whereabouts of A. fragillimus. For nearly a cen- tury the remains have yet to resurface. The enormous size of the specimen has generally been accepted despite being well beyond the size of even the largest sauropods known from verifiable fossil material (e.g. Argentinosaurus). By deciphering the ontogenetic change of Diplodocoidea vertebrae, the science of gigantism, and Cope’s own mannerisms, we conclude that the reported size of A. fragillimus is most likely an extreme over-estimation. INTRODUCTION saurs pale in comparative size; thus A. fragillimus could be the largest dinosaur, and largest vertebrate in Earth’s history Described by Edward Drinker Cope in 1878, the holo- (the Blue Whale being approximately 29 meters long [Reilly type (and only) specimen of A. -
PRRUCS Paper S4-3 Peter Dodson Oct 2016.Pub
On Fossils and Faith Peter Dodson Program for Research on Religion and Urban Civil Society, University of Pennsylvania, Philadelphia, PA 19104, USA October 14, 2016 I am a geologist, paleontologist, veterinary anat- that time I have spent my entire professional omist, evolutionary biologist, and a lifelong career teaching gross anatomy to veterinary Christian. I am extraordinarily privileged to students at the University of Pennsylvania in teach in a superb research university, and I have Philadelphia, while also supervising undergradu- been blessed with a succession of excellent stu- ate and graduate students in the Department of dents with whom I have traveled the world. I Earth and Environmental Sciences. have been even more greatly blessed with the companionship of my wife of 48 years, Dawn, In the first two decades of my scientific ca- with whom I have two children and three grand- reer, I confined my research to Canada and the children. These are the three great priorities of United States. My first new discovery was a my life: family, faith and fossils. small horned dinosaur from south central Mon- tana, which in 1986 I named Avaceratops lam- As a child, dinosaurs fascinated me. While mersi. A skeleton of the dinosaur is on display most children grow out of this fascination, I at the Academy of Natural Sciences of Philadel- simply never did. I lived in northern Indiana phia (now the Academy of Natural Sciences of until I was 11. My older brother, Steve, was an Drexel University). This animal is named not amateur naturalist and astronomer. He taught after my wife, but after Ava Cole, the wife of me to love collecting fossils. -
Osteology of the Dorsal Vertebrae of the Giant Titanosaurian Sauropod Dinosaur Dreadnoughtus Schrani from the Late Cretaceous of Argentina
Editors' choice Osteology of the dorsal vertebrae of the giant titanosaurian sauropod dinosaur Dreadnoughtus schrani from the Late Cretaceous of Argentina KRISTYN K. VOEGELE, MATTHEW C. LAMANNA, and KENNETH J. LACOVARA Voegele, K.K., Lamanna, M.C., and Lacovara K.J. 2017. Osteology of the dorsal vertebrae of the giant titanosaurian sauropod dinosaur Dreadnoughtus schrani from the Late Cretaceous of Argentina. Acta Palaeontologica Polonica 62 (4): 667–681. Many titanosaurian dinosaurs are known only from fragmentary remains, making comparisons between taxa difficult because they often lack overlapping skeletal elements. This problem is particularly pronounced for the exceptionally large-bodied members of this sauropod clade. Dreadnoughtus schrani is a well-preserved giant titanosaurian from the Upper Cretaceous (Campanian–Maastrichtian) Cerro Fortaleza Formation of southern Patagonia, Argentina. Numerous skeletal elements are known for Dreadnoughtus, including seven nearly complete dorsal vertebrae and a partial dorsal neural arch that collectively represent most of the dorsal sequence. Here we build on our previous preliminary descrip- tion of these skeletal elements by providing a detailed assessment of their serial positional assignments, as well as comparisons of the dorsal vertebrae of Dreadnoughtus with those of other exceptionally large-bodied titanosaurians. Although the dorsal elements of Dreadnoughtus probably belong to two individuals, they exhibit substantial morpho- logical variation that suggests that there is minimal, if any, positional overlap among them. Dreadnoughtus therefore preserves the second-most complete dorsal vertebral series known for a giant titanosaurian that has been described in detail, behind only that of Futalognkosaurus. The dorsal sequence of Dreadnoughtus provides valuable insight into serial variation along the vertebral column of these enormous sauropods. -
Determining the Largest Known Land Animal: a Critical Comparison of Differing Methods for Restoring the Volume and Mass of Extinct Animals
ANNALS OF CARNEGIE MUSEUM VOL. 85, NUMBER 4, PP. 335–358 31 DECEMBER 2019 DETERMINING THE LARGEST KNOWN LAND ANIMAL: A CRITICAL COMPARISON OF DIFFERING METHODS FOR RESTORING THE VOLUME AND MASS OF EXTINCT ANIMALS GREGORY PAUL 3100 St. Paul Street 604, Baltimore, Maryland, 21218 [email protected] ABSTRACT Recent claims regarding what is and is not the largest known sauropod dinosaur are tested via dimensional comparisons of the most critical metrics of relative size—especially, when possible, the functional lengths of the dorsal vertebral centra and the articulated length of the combined trunk vertebrae—and analog volumetric models based on technical skeletal restorations. The Cretaceous Argentinosaurus massed 65–75 tonnes, and its dorsal vertebrae and dorsal–sacral series are much larger than those of any other described titanosaur. Specimens of Patagotitan indicate a 50–55 tonne titanosaur, and the less complete Notocolossus, Puertasaurus, and ‘Antarctosaurus’ giganteus appear to have occupied a similar size range. Paralititan weighed between 30 and 55 tonnes. The juvenile Dreadnoughtus, as well as Futalognkosaurus and Alamosaurus, were in the area of 30 tonnes, with the possibility that the last was substantially larger. Entirely analog, skillfully produced, high-anatomical-fidelity skeletal restorations and volumetric models representing a prime-lean condition are approximately as scientifically objective and accurate, as well as more realistic than, analog-digital, crudely-formed convex hull volumetric models, which are based on subjectively and often inconsistently or erroneously mounted skeletons and digitized skeletal reconstructions. The need to ensure that skeletal restorations are as anatomically correct and consistent as the data allow is stressed, which requires that researchers and illustrators be sufficiently skilled in animal and especially dinosaur anatomy, and the proce- dures and standards for achieving the best possible results are detailed. -
1 CLASS 11: ANATOMY & SAUROPODOMORPHS I. BASIC DINOSAUR ANATOMY Anatomical Orientation Anterior
CLASS 11: ANATOMY & SAUROPODOMORPHS I. BASIC DINOSAUR ANATOMY Anatomical orientation Anterior - front Posterior - back Dorsal - top Ventral - bottom Proximal - near Distal - far Medial - towards the middle Lateral - towards the side Major parts of the skeleton you should know Vertebral column: cervical (neck), thoracic (attach to rib), sacral (attach to pelvis), and caudal (tail) vertebrae. Sacral vertebrae = sacrum Forelimb: Scapula, Coracoid, Humerus, Radius, Ulna, Wrist (carpels), Finger bones (metacarpels and phalanges) Hindlimb: Pelvis = Ilium, Pubis, Ischium, Femur, Tibia, Fibula, Ankle (tarsals = Astragalus, Calcaneum), Toe bones (metatarsals and phalanges) Head bony parts: Lower jaw: dentary (with teeth), articular (lower jaw joint), others bones Upper jaw: premaxilla and maxilla (both with teeth), quadrate (upper jaw joint) Skull: skull roof bones (nasals, frontals, parietals, etc.), cheek and eye bones (jugal, lacramal, post-orbital, etc.), palate bones, braincase, occipital condyle (head-neck joint) Head holes: foramen magnum (hole where spinal cord enters braincase), antorbital fenestra, upper and lower temporal fenestra, mandibular fenestra (on lower jaw) Sauropodomorpha All herbivores Shared derived traits Small head (5% body length), long neck with at least 10 vertebrae, elongate peg- like teeth with coarsely serrated crowns, large thumb with claw, long femur, large obturator foramen in pubis Prosauropoda few easily diagnosed shared features whopping big thumb claw that is further twisted, elongate cervical vertebrae (Also: small 5th digit on hind foot, iguana-like teeth) Some used gastroliths Quadrupeds and facultative bipeds Plateosaurus, Anchisaurus, Mussasaurus (representative genera) Known from Late Triassic through Early Jurassic Global distribution (Pangean) These represent the earliest plant-eating dinosaurs, which presumably evolved from bipedal meat eaters. -
Contributions from the Museum of Paleontology, University of Michigan
Contributions from the Museum of Paleontology, University of Michigan VOL. 32, NO. 11, PP. 189–243 APRIL 10, 2017 MOABOSAURUS UTAHENSIS, N. GEN., N. SP., A NEW SAUROPOD FROM THE EARLY CRETACEOUS (APTIAN) OF NORTH AMERICA BY BROOKS B. BRITT1, RODNEY D. SCHEETZ1, MICHAEL F. WHITING2, AND D. RAY WILHITE3 Abstract — The Early Cretaceous was a time of dramatic change for sauropod dinosaurs in North America. Between the Late Jurassic-aged Morrison Formation and overlying Early Cretaceous strata, there was a dramatic decline in sauropod diversity. Here, we describe a new sauropod that adds to the diversity of the Early Cretaceous, from strata that can be no older than the early Aptian, (125 Ma) some 25 million years younger than the Morrison Formation. Moabosaurus utahensis, n. gen., n. sp., is diagnosed in part by the following suite of charac- ters: axially thin ventral basioccipital with posteriorly sweeping basal tubera; low-spined cervical vertebrae with neural spines that range from shallowly notched on anterior cervical vertebrae to shallow, but widely notched on middle and some posterior cervical vertebrae; posterior cervical and anterior dorsal neural spines with extremely low, axially thin, laterally wide ridges at the level of the zygapophyses; some cervical ribs with bifid posterior shafts; anterior and posterior caudal vertebrae with strongly procoelous centra, middle caudal vertebrae with mildly procoelous centra, and distal caudal vertebrae with moderately-to-strongly procoelous centra. To determine the phylogenetic position of Moabosaurus we utilized three different datasets and performed four analyses. All results are in agreement that Moabosaurus is a neosauropod. The two most resolved trees indicate it is a macronarian, specifically a basal titanosauriform. -
Subject Index
Cambridge University Press 0521811724 - The Evolution and Extinction of the Dinosaurs, Second Edition David E. Fastovsky and David B. Weishampel Index More information Subject index Bold type indicates figures. Absolute age 23–5 Arctic dinosaurs 372–373, 373 Actinopterygii 67 Asteroid impact 425, 426–32 Aguja Formation (Upper Cretaceous, USA) 401 see also Cretaceous–Tertiary boundary; Alxa Desert (China) 297 Iridium; Chicxulub Amarga Canyon (Argentina) 262 Affects of 432–4 American Museum of Natural History (USA) Indicators of 426, 427–9, 428, 429, 430, 431 18n, 19, 253, 259, 292 Atlas Mountains (Morocco) 263 Amnion 77 Auca Mahuevo (Upper Cretaceous, Argentina) Amniota 245, 246 Diagnostic characters 77, 78 Aves, see Theropoda Major groups 78 Azendoh (Morocco) 261 Amphibia 77, 77n, 393 Amur River (Sino-Russian border) 217 Baharije Oasis (Egypt) 291, 292 Anapsida 78, 79–80 Barun Goyot Formation (Upper Cretaceous, Anhui Province (China) 162 Mongolia) 401 Ankylosauria Bernissart (Belgium) 212, 213 Age 133, 396, 397 Biological classification 68 Armament 133, 137, 138, 139 Biostratigraphy 22, 27–8 Behavior 134–7, 138–9 Birds, see Theropoda Clades of 133, 139–42 Body plans 65 Cladogram of, see Cladograms Bone histology 362–7, 363 Derived characters of 140, 139–41, 142 British Museum (Natural History) 234, 258, 336 Distribution 134, 135 Bug Creek (USA) 441 Evolution of 139 Burgess Shale (Middle Cambrian, Canada) 64 Fermentation in 136, 137 Burial 6, 7, 8, 9, 10 History of discovery 142–5 Inferred intelligence 361 Canadian Dinosaur Rush 182, 183, -
Osteological, Myological, and Biomechanical Investigations of the Sauropod Dinosaur
Osteological, myological, and biomechanical investigations of the sauropod dinosaur Dreadnoughtus schrani and molecular paleontological investigation of the marine crocodile Thoracosaurus neocesariensis A Thesis Submitted to the Faculty of Drexel University by Kristyn K. Voegele in partial fulfillment of the requirements for the degree of Doctor of Philosophy August 2016 © Copyright 2016 Kristyn K. Voegele. All Rights Reserved. ii Dedication To those with dreams and those that help them achieve dreams iii Acknowledgements This project would not be possible without the support, assistance, and guidance of just about everyone I have had interacted with in the last five years. I am grateful for all of your time, effort, patience, and support. First, I would like to thank Drexel University, the College of Arts and Sciences, the Department of Biology, the Academy of Natural Sciences, the Department of Biodiversity, Earth, and Environmental Science, and all the faculty and staff that made my graduate education possible. At Drexel I have been fortunate to have the opportunity to learn from world-class scientists about research, teaching, and everything in between. I appreciate the resources and support provided to me over my graduate career to turn me into a capable scientist. I am also thankful to the organizations that have financially supported my research. Without the financial support from a National Science Foundation Graduate Research Fellowship (DGE Award #1002809) and a Paul Bond Scholarship from the Delaware Valley Paleontological Society much of my research would have been unaffordable. I am thankful to these institutions for helping me expand my area of focus and training in my field of study during my graduate career.