<<

University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln

Transactions of the Nebraska Academy of Sciences and Affiliated Societies Nebraska Academy of Sciences

1981

Sedimentology of Norden Bridge and Egelhoff Fossil Quarries () of North-Central Nebraska

Carl F. Wellstead McGill University

Follow this and additional works at: https://digitalcommons.unl.edu/tnas

Part of the Life Sciences Commons

Wellstead, Carl F., "Sedimentology of Norden Bridge and Egelhoff Fossil Quarries (Miocene) of North- Central Nebraska" (1981). Transactions of the Nebraska Academy of Sciences and Affiliated Societies. 269. https://digitalcommons.unl.edu/tnas/269

This Article is brought to you for free and open access by the Nebraska Academy of Sciences at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Transactions of the Nebraska Academy of Sciences and Affiliated Societiesy b an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. 1981. Transactions a/the Nebraska Academy a/Sciences, IX:67-85.

SEDIMENTOLOGY OF NORDEN BRIDGE AND EGELHOFF

FOSSIL QUARRIES (MIOCENE) OF NORTH-CENTRAL NEBRASKA

Carl F. Wellstead Redpath Museum McGill University 859 Sherbrooke Street West Montreal, Quebec H3A 2K6 Canada

Measured sections demonstrate the positions of the Norden Bridge 2.5 km northwest of Norden Bridge Quarry. Fossil vertebrates an, Egelhoff fossil quarries in the local stratigraphy and show Egelhoff from these quarries have been the subject of a series of papers QL ,rry to be topographically higher. The sections do not resolve the during the last 20 years (Smith, 1962; Klingener, 1968; reL live stratigraphic positions of the quarries. Descriptions of sedi­ me'lts at the two quarries demonstrate that coarser sediments exist at Undsay, 1972; Rich and Rasmussen, 1973; Storer, 1973; Nc,,jen Bridge Quarry. However, these coarse sediments, as well as the Holman, 1976). In addition to the principal descriptive and fOHl remains of large vertebrates, are limited to two particular beds at taxonomic purposes of these papers, they demonstrate that N(, jen Bridge Quarry, while three other beds comprise sediments the fossil vertebrate remains from the quarries are, in most eit"~r finer or statistically indistinguishable from those at Egelhoff cases, water-transported, isolated bones. Qu" rry. The association of large clasts and large bones suggests hy­ dr.)' lic sorting of these sedimentary particles and supports the sugges­ tio;. that the difference in fossil faunas between the two quarries is Remarkably for two such well publicized localities, no depositional. Evaluation of cross-strata sets at both quarries indicates stratigraphic section has been published marking their posi­ that local paleocurrents do not reflect the easterly regional dip of local tions in the regional stratigraphy. Holman (1973) and Chantell Tefl iary strata. (1971) reported that Egelhoff Quarry is topographically 7.7 m (25 ft) higher than Norden Bridge Quarry. Holman (1973) t t t suggested additionally that the quarries are temporally equiva­ lent. Similarities in their faunas (cf Tables I and II) support INTRODUCTION the temporal equivalency of the two sites; however, fossils of larger such as perissodactyls and artiodactyls Leidy (1858) published the first report of fossil verte­ are much less common at Egelhoff Quarry and proboscideans bra,~s from the fluvial Tertiary rocks in the vicinity of Valen­ are entirely absent. Experimental work in taphonomy by tine Nebraska. Since then, discussions of these rocks have Voorhies (1969), Dodson (1973), and Korth (1979) suggests con inued, including considerable debate over the propriety that the difference in the size ranges of fossil vertebrates at of lcal stratigraphiC names, particularly use of the term the two quarries may be a depositional phenomenon, reflect­ "V~ ~ntine" regarding what is now agreed to be the Valentine ing a difference in competency between the currents deposit­ For. Ilation and its fossil faunas. These debates are now re­ ing the bones and sediments at the two sites. The observation solv:d to the satisfaction of most and are concisely reviewed by Holman (I 973) that sediment clast-size is greater at Norden by ~kinner , Skinner, and Gooris (1968). Bridge Quarry than at Egelhoff Quarry supports this sugges­ tion; however, this clast-size difference has not been demon­ ·\ttention of vertebrate paleontologists familiar with the strated quantitatively. Vak'ltine Formation has been focused upon the fossil quarries Withn the formation since Hibbard (1960) announced the The purpose of this paper is to present measured sections disc very of fossil vertebrates from the now well-known of the stratigraphy at the Egelhoff and Norden Bridge quarries Nor,' en Bridge Quarry (Fig. 1). In 1964 Morris Skinner dis­ and to illustrate sedimentological differences between them by cOve:ed another site, the Egelhoff Quarry, approximately means of description and a sieve analysis of sediments. Cross- 67 68 C. F. Wellstead strata and elongate bone orientations are evaluated as indica­ TABLE I. (Continued). tors of paleocurrent direction at the quarries.

Order Squamata TABLE I. Fossil vertebrate taxa identified from Norden Bridge Quarry (from Falk, Osborn, Pepped, and Voorhies, Family Xenosauridae 1980). Nordenosaurns magnus - large extinct lizard Family Iguanidae Sceloporns sp. Leiocephalus sp. - undescribed of extinct tropidurine Class Osteichthyes lizard Family Lepisosteidae Family Anguidae Lepisosteus sp. - garpike Gerrhonotus sp., cf G. mungerornm - extinct alligator lizard Family Amiidae Ophisaurns ventralis - eastern glass lizard Amia sp., cf A. calva - bowfin cf Peltosaurns - extinct lizard Family Ictaluridae Family Scincidae letalurns lambda - large extinct catfish Eumeces sp. - striped skink I. punctatus - channel catfish Family Amphisbaenidae Family Centrarchidae unidentified genus and species of worm lizard Lepomis sp., cf L. microlophus - sunfish Family Boidae

Class Amphibia Charina prebottae - extinct rubber boa Order Urodela Family Colubridae Family Cryptobranchidae cf Thamnophis - garter snake cf Neonatrix e/ollg{lta - extinct water snake Andrias matthewi ~- Matthew's giant salamander Paleoheterodon tiheni - ancestral hognose snake Family Ambystomatidae Nebraskensis skinneri - archaic colubrid snake Ambystoma minshalli - extinct mole salamander Salvadora paleolineata - extinct patch-nose snake Lampropeltis similis - ex tinct small king snake Order Anura Elaphe nebraskensis - extinct rat snake Family Pelobatidae Scaphiopus (Scaphiopus) wardornm - extinct spade foot toad Qass Mammalia S. (Spea) sp., cf S. bombifrons - plains spade foot Order Insectivora Family Bufonidae Family Erinaceidae Bufo sp., cf B. hibbardi - extinct toad B. valentinensis - extinct toad Parvericius montanus - small extinct hedgehog B. kuhrei - extinct toad Untermannerix copiosus - medium-sized extinct hedgehog Metechinus amplior - large extinct hedgehog Family Hylidae Family Plesiosoricidae Acris sp., cf A. crepitans - cricket frog Pseudacris sp., cf P. clarki - spotted chorus frog Plesiosorex sp., cf P. donroosai - large extinct insectivore P. nordensis - extinct chorus frog Family Soricidae Hyla sp., cf H. gratiosa - barking tree frog H. sp., cf H. squirella - tree frog Alluvisorex sp. - extinct shrew H. sp., cf H. versicolor - gray tree frog Family Talpidae Family Ranidae Mystipterns sp. - extinct shrew-mole Rana sp., nr. R. pipiens - leopard frog Domninoides valentinensis - extinct mole Qass Reptilia Scalopoides sp. - extinct mole Order Testudinata Order Lagomorpha Family Emydidae Family Leporidae Chrysemys sp., cf C. picta - painted turtle Hypolagus sp. - archaic rabbit Family Testudinidae Family Ochotonidae Geochelone onhopygia - giant land tortoise Hesperolagomys sp. - archaic pika G. nordensis - midget land tortoise Order Rodentia Family Trionychidae Family Aplodontidae Trionyx sp. - softshell turtle Allomys sp., cf A. stirtoni - extinct sewellel Sedimentology of Norden Bridge and Egelhoff quarries 69

TABLE I. (Continued). TABLE I. (Continued).

Family Family Equidae Mylagaulus sp. - horned Hypochippus sp. - browsing horse sp. Family Sciuridae Archaeohippus Merychippus sp. - primitive grazing horse Tamias sp. - extinct chipmunk Calippus sp. - small grazing horse Family Castoridae Protohippus perditus - tridactyl grazing horse Monosaulax sp. A - large primitive beaver "Hipparion" sp. - tridactyl grazing horse M. sp. B - small primitive beaver Order Artiodactyla Family Eomyidae Family Tayassuidae Paradjidaumo stirtoni - extinct archaic rodent cf. Prosthennops - extinct peccary Family Zapodidae Family Merycoidodontidae Megasminthus tiheni - large archaic jumping mouse Plesiosminthus sp. - small archaic jumping mouse cf. Ustatochoerus sp. - oreodont Family Heteromyidae Family Cervidae Perognathus sp., cf, P. furlongi - extinct pocket mouse Cranioceras (Procranioceras) sp., cf, C. (P.) skinneri - extinct P. trojectioansrum - small extinct pocket mouse three-horned "Diprionomys" sp., cf, P. agrarius - archaic pocket mouse Blastomeryx sp. - small sabertoothed deer D. sp. - large extinct pocket mouse Family Cupidinimus nebraskensis - small extinct pocket mouse Procamelus sp. - ancestral camel Family Cricetidae Protolabis sp. - small camel Copemys niobrarensis - extinct deer mouse Family Antilocapridae Tregomys sp. - extinct deer mouse Merycodus sp. - forked-horned prongbuck Family Geomyidae Ramoceros sp. - large-horned prongbuck undetermined genus and species of extinct pocket gopher undetermined genus and species of small prongbuck

Order Family Amphicyonidae cf, Pliocyon - large bear-dog TABLE II. Fossil vertebrate taxa identified from Egelhoff Family Canidae Quarry (from Falk, Osborn, Pepped, and Voorhies, 1980)_ Aelurodon sp. - large hyaenoid dog Tomarctus sp. - small hyaenoid dog Leptocyon vafer - fox-sized dog Cynarctus sp. - extinct raccoonlike dog aass Osteichthyes Family Mustelidae Family Ictaluridae Leptarctus primus - extinct mustelid leta/urus sp_ - catfish undetermined genus and species of small mustelid Family Centrarchidae Family Procyonidae undetermined genus and species of sunfish Bassariscus sp., cf, B. parvus - extinct miner's cat aass Amphibia Order Proboscidea Order Urodela Family Gomphotheriidae Family Ambystomatidae Gomphotherium sp., cf, G. productus - long-jawed mastodon Ambystoma minshalli - extinct mole salamander Order Anura Order Perissodactyla Family Pelobatidae Family Rhinocerotidae Scaphiopus sp_, cf, S. bombifrons - plains spadefoot toad Teleoceras sp. - short-limbed rhinoceros S. sp., cf, S. holbrooki - eastern spadefoot Aphelops sp. -long-limbed rhinoceros Family Bufonidae Family Tapiridae Hufo valentinensis - extinct toad Tapirus sp. - extinct tapir B. sp., cf, B. hibbardi - extinct toad 70 C. F. Wellstead

TABLE II. (Continued). TABLE II. (Continued).

Family Hylidae Family Heteromyidae Acris sp., cf A. crepitans - cricket frog Cupidinimus nebraskensis - small pocket mouse Pseudacris sp., cf P. clarki - spotted chorus frog Perognathoides sp_ - pocket mouse Hyla sp., cf H. cinerea - green frog Family Zapodidae H. sp., cf H. crucifer - spring peeper Megasminthus tiheni - archaic jumping mouse Family Ranidae Plesiosminthus sp. - small archaic deer mouse Rana sp. - frog Family Cricetidae

Class Reptilia Copemys sp_, cf C kel/ogae - archaic deer mouse Order Testudinata Order Carnivora Family Testudinidae Family Procyonidae Geochelone sp. - giant land tortoise Bussariscus sp. - extinct miner's cat Family Trionychidae Family Mustelidae Trionyx sp. - softshelled turtle unidentified weasellike carnivore Order Squamata Order Perissodactyla Family Anguidae Family Equidae Ophisaurus ventralis - eastern glass lizard unidentified genus and species of horse Ge"honotus sp., cf G. mungerorum - extinct alligator lizard Order Artiodactyla extinct lizard Peltosaurus minimus - Family BIastomerycidae Family Scincidae Blastomeryx sp_ - sabertoothed small deer Eumeces sp. - striped skink Family Antilocapridae Family Boidae Merycodus sp_ - forked-horned prongbuck Charina prebottae - ancestral rubber boa

Family Colubridae Neonatrix elongata - archaic water snake Paleoheterodon tiheni - ancestral hognose snake Nebraskophis skinneri - archaic colubrine snake Salvadora paleolineata - extinct patch-nosed snake Elaphe nebraskensis - extinct rat snake METHODSANDEQUWMENT Qass Mammalia Order Insectivora Stratigraphy Family Erinaceidae Parvericios montanus - small archaic hedgehog The stratigraphic sections were measured using the clino· Untermannerix copiosus - medium-sized archaic hedgehog meter of the Keuffel and Esser pocket transit as an eye level. Metechinus amplior - large archaic hedgehog Family TaIpidae Mechanical Analysis of Sediments Domninoides valentinensis - mole Order Lagomorpha Laboratory equipment used in processing sediment sam· Family Leporidae pIes and in execution of sieve analyses is as follows: Hypolagus sp. - archaic rabbit Family Ochotonidae 1. Model CL-280-A mechanical sediment separator, Soil Test, Incorporated cf Hesperolagomys - archaic pika Order Rodentia 2. Torbal PL·800 Balance, Torsion Balance Company Family Sciuridae Tamias sp_ - extinct chipmunk 3. Mettler H54 Balance Family Castoridae Monosaulax sp_ - primitive beaver 4. Ro·Tap no. 8017, W. S. Tyler Company Sedimentology of Norden Bridge and Egelhoff quarries 71

The sedimentary analysis procedure was adapted from versity of Michigan, Michigan State F \lk (1974). University (MSU), and Univer­ sity of Nebraska State Museum C ientation Data (UNSM). Keya Paha County, Ne­ braska. The strike and apparent dips of cross-strata foresets and tl ~ plunge and trend of elongate fossil bones were read using Bw 106 Norden Bridge Quarry of the tl ,~ clinometer and compass of the pocket transit. Strike and American Museum of Natural ar parent dips of cross-strata were converted to true strike and History, University of Michigan, dJJ following Billings (1972). Mean vectors and mean angular MSU, Notre Dame University, and d( Jiations were calculated following Till (1974). A hybrid UNSM. Brown County, Nebraska. F'est (Griffiths and Rosenfeld, 1953), modified by Potter and Statistics. M Graphic mean. Pettijohn (1977), was used to test the significance of orienta­ z tiln data. uG Graphic standard deviation. Inclusive graphic standard devia­ tion. Localities. Kp 101 Egelhoff Quarry of the American Museum of Natural History, Uni- Null hypothesis.

o 2 to I I N Norden Km. \ \ ....

" \ NEBRASKA \ , "\ \ \ 0100 \ \ Km. "\ \ , \ I I I I I A '0 Kp 101 o I I \ , \ \ B BROWN

T 33 N T32N

SCALE BASE: ,,- I NORDEN, NEBR.-S. OAK. 15' QUAD (1950) to Johnstown 10005'

FIGURE 1. Regional locator of Norden Bridge Quarry (Bw 106) and Egelhoff Quarry (Kp 101). 72 C. F. Wellstead

Alternative hypothesis. System

F-test statistic. Pierre Shale. The oldest formation exposed in the region t t-test statistic. is the Cretaceous Pierre Shale, seen at Meadville, some 18 km downstream (east) from Norden Bridge.

STRATIGRAPHY Tertiary System

Measured stratigraphic sections at Egelhoff and Norden Chadron Formation. A stratigraphic unit which Skinner Bridge quarries are presented in Figure 2 along with one of the (Skinner and Hibbard, 1972) referred to the Chadron Forma­ most complete sections of local Tertiary stratigraphy avail­ tion is exposed north of the Niobrara River in roadcuts along able for reference and comparison. Geographic positions of Nebraska Highway 183 approximately 25 km east of the study these measured sections are presented in Figure 1. area. Skinner described the lithology of the formation at this exposure as a yellowish, buff clay with zones of brown iron Brief descriptions of the local stratigraphy are offered oxide stain and bearing clear quartz and grains. Although the below to review the relevant units. Reference is made to more unit is not exposed in the Norden Bridge vicinity, a similar comprehensive works of the regional stratigraphy. unit was encountered between the Pierre Shale and Rosebud

FEEl MEASURED SECTIONS Qd (GEOGRAPHIC REF.: FIG 1)

T,~·y.,-'_y_ ~-r~...,;...: __Yo _. _'Y_. 7'-':--T-'f 2500

':-Y'-:-'-T :-:-:"T_:~""_'" ASH HOLLOW FM. y·_,y..:... -,.,.:.... "'1_ 1'~ CAP ROCK MBR. LEGEND ~~=~=~'~. f-:::-:::-:::j CLAY

T 1"'- ~"--''7"-,_, -c::::;:::,.....; <::::::::::> ------"F-~,,-=~or-;;~!~~~~7~P~~_;ii-I'- ~ PALEOSOL T- 7 _.,... -:l""-,""",:- b-= -=-=j SILT 2400

CJ SAND 1= =ILEDGEY VALENTINE FM. ______CROOKSTON BRIDGE MBRr----c-_-_~_-_7":-"- ~GRAVEL g ~ I~ 1.°""' } ~,~,~, EGELHOFF SECTION r------, 230d 2300 ro:::o-ol COBBL ES & ~BOULDERS ~ TUBULAR NORDEN BRIDGE Q-=---iQ--O~ SECTION ;-::::- :::~---L)-- I---<-~ l CLAY - -"- STRINGER

.•.. ..• _:.··.. '.•. -.·.-.·.••_.•. ·.O .. "..'.- .... = ...... •.: ...-... _...... : ..•.- ..... _...._.••.... 0.•. •..•.•... •.· •....•.·•·. 2249 =:::= ~ ROOT ::;:.'} CONTACTS ~.6._.-·.·.~ ~ ~~ ~:::=~- ~ CASTS ~ ~-=~=-?~ r;:;;;::t:~;2::t;:~} Kp 101 ·'.. ".<;>.':;.·;:'·d ... ,··· x Bw 106 2200 Co __ 2~ :::-_~~~~~~~_-::: 1_ -- -I INfERRED ROSEBUD FM. <::::> I SAND r---l COVERED 1 .c:Ji7 LENSES L---.J SECTION ~----~ 2~;: IL-----' '------~ 2117' 2100 FIGURE 2. Regional stratigraphy. Refer to Figure 1 for location of particular section. Sedimentology of Norden Bridge and Egelhoff quarries 73

I ormation in drill holes near the Norden damsite (Anony­ cross-bedded or horizontally laminated with thick and thin I ,ous, 1977). Fragmentary mammalian remains recovered beds following one another in rapid succession. The propor­ f om the highway 183 outcrop are insufficient to corroborate tions of sand, silt, and clay vary as well. Cross-bedding, lami­ t le accepted Chadronian provincial age assigned to the forma­ nation, and textural variability indicate rapidly fluctuating t In. With respect to the European time scale, the formation flow regimes during deposition of the member. At its type i~ considered to be Oligocene (Tedford, et al., 1981). section in northern Cherry County, Nebraska, it is from 46 to 54 m (150 to 175ft) thick (Skinner, Skinner, and Gooris, Rosebud Formation. Disconformably overlying the 1968). It is believed that the Egelhoff and Norden Bridge (iladron Formation is a pinkish, gray or tan, fme sandy or quarries occur in the Crookston Bridge Member, a subdivision c lyey siltstone, which forms high bluffs along stream courses. of the Valentine Formation. However, Holman (1976) re­ Tlis report follows Skinner, Skinner, and Gooris (1968) and ported that M. F. Skinner is considering a revision of the V. ebb (1969) in considering the unit to be a southerly exten­ Valentine stratigraphy. si III of the Rosebud Formation of Gidley (1904). The form­ a ion has nearly continuous exposures along the Niobrara The thickness of Valentine Formation depicted in section Ever between Valentine, Nebraska, and the Old Bruce Mill A (Fig. 2) is considered to represent only the Crookston (:

;Ie elevation of 691 m (2,249 ft). Above this point the section A fourth episode of sedimentation is represented by a covered by vegetation. bed of rounded, well-indurated, clayey, intraformational cobbles and boulders (maximum diameter 25 cm) as well as At no time was the entire quarry wall exposed for photog­ the disarticulated remains of large ungulates and probosci­ I_lphy, as it was impossible to keep pace with the slumping of deans (Fig. 4B and C; Table III). This bed varies in thickness 1 \e quarry face. Figure 3 is a composite diagram of the quarry from approximately 10 to 50 cm. Upper and lower contacts j .Ice compiled from individual exposures of portions of the of the bed are gradational for a distance of a centimeter or cuarry. two.

Sedimentary units recording five distinct episodes of The highest bed exposed in the quarry is a bed of uncon­ c3position were exposed at the quarry in 1976. The lowest solidated, cross-bedded, gray sands approximately 3.0 m thick. I-lit exposed was a cross-bedded, coarse gray sand with crystal­ The quarry sediments are covered by 25 to 50 cm of sandy soil. LIe gravel and gravel-sized clay balls (Fig. 4A). The coarsest of t,is gravel was retained on a ~ -3.5 sieve. Some 50 cm of this Egelhoff Quarry. The Egelhoff Quarry is located on the \:lit were excavated, but its base was not found as slumping east side of the Norden-Johnstown Road across from the 0" the unconsolidated sediment frustrated further digging. 2,188 ft bench mark at the Egellioff Ranch driveway (SWI4 SE'l4 NE'l4 SW'l4 Sec. 29, T. 33 N., R. 23 W., Keya Paha Coun­ The sand and gravel layer has sharp contact with an over- ty, Nebraska). 1:. ing white-weathering layer of silt, indurated by interstitial C:ly (Fig. 4A). The silty layer is approximately 15 cm thick The stratigraphy is not well exposed at the quarry (Fig. a d is blocky in appearance. 2). Therefore, the position of the quarry relative to the upper and lower contacts of the Valentine Formation, and the Overlying and in sharp contact with the white siltstone Norden Bridge Quarry as well, are speculative, particularly is a bed of unconsolidated, medium- to fine-grained, cross­ considering the relief present upon the Valentine-Rosebud b,dded, gray sand, some 35 to 40 cm thick (Fig. 4B). formational contact.

SOIL 1M

5

SLUMP

\;'i;,i5~_-_;_btJff;frfl_c3.-o~_;oa:~_-.-)-_~ao_·P_,~~~_.o_80··~_;~O_·:_U_'i9,_·_.~_oO£l?O_-_-~~_b_·_-·-' ~ Go ,~.,',,' ,',CO , ,,' '.. - -' .. ' -.... -,0, Q' . 0 o~ -b~:·~ (j6,-: ' .•: ~,. g, ~ -, '~-_ --o,:~~' '0-' .• " _ -' : - "".: CJ . ~.-~~, _<_g_(j_O_O-_-o.c.o.O'~.O.-'6.'.~.:~,:' o-o~-~O:-- ~ >... _>. ... '. -.?_ ~ ~rY~~}5;·~L;i;:;:.:2~;ij;§·;:~;\·;·;:.:~\';Sf~";::"~:::';'J~ .., ..:~i·}~;;fif'

FIGURE 3. Diagram of Norden Bridge Quarry. 1. Sand and gravel bed. 2. Silty bed. 3. Lower cross-bedded sand. 4. Cobble anu boulder bed. 5. Upper cross-bedded sand. 76 C. F. Wellstead

A

c

FIGURE 4. A and B. Norden Bridge Quarry. A. Sand and gravel (1) and silty bed (2) of Figure 3. Scale: ice pick, 10 cm long. B. Beds 3, 4, and 5 of Figure 3. Scale: shovel handle, 75 cm. C. Skull of Protohippus perditus, UNSM 56063. Scale: ruler in cm. D. Egelhoff Quarry with sediment sampling grid for lower (1) and upper (2) silty beds and Egelhoff Quarry sands (3). Scale: grid unit is 30 cm.

TABLE III. Examples of large vertebrate remains from The sediments of the quarry comprise a single bed of un­ Norden Bridge Quarry boulder bed, summer 1976. consolidated, cross-bedded, gray sands approximately 1.6 m thick, bounded above and below by unfossiliferous beds of Maximum fine-grained sand and silt indurated by interstitial clay (Fig. Identification Length (cm) Museum Number 4D).

The lower fine-grained layer is olive drab in color when Protohippus skull 27 UNSM 56053 fresh, but dries to an off-white shade. Only the top of the bed Camelid scapula 43 UNSM 56055 was exposed in 1976. The bed bears a sharp erosional contact Proboscidean tusk fragment 44 UNSM 56056 with the quarry sands at 675 m (2,195 ft). Proboscidean tusk fragment 33 UNSM 56057 Proboscidean dentary with 2 molars 40 MSU Specimen The quarry sands themselves are a group of interfering trough cross-strata, each relatively uniform in grain size. Sedimentology of Norden Bridge and Egelhoff quarries 77

)uccessive cross-strata become finer-grained upward, however. Laboratory Procedure \t their upper limit the quarry sands grade over a distance of i 0 to 15 cm into the overlying fme-grained layer, which is The goal of the analysis was to investigate a pOSSible, ;imilar to the fine-grained bed underlying the quarry sands, statistically significant, clast-size difference between two sand Jnd overhangs the quarry as a ledge. The quarry exposure is beds rather than a finer sedimentary environmental distinc­ lpproximately 9 m wide. tion. Toward this end, James B. Swineheart, Conservation and Survey Division of the University of Nebraska-Lincoln, Discussion suggested that the project could be justifiably expedited by working with one-half of the collected samples from each unit. From their descriptions it should be apparent that the The handling of the samples up to the actual mechanical analy­ '

Because in practice the random sample is difficult to ob­ 2. Graphic standard deviation, aG , which is very similar tain, a systematic, mechanical scheme was employed to collect to the statistical standard deviation and used here to an initial 25 (1 kg estimated) sediment samples (number of calculate the standard error. samples set arbitrarily). A square grid was erected against the face of the bed to be sampled. This grid was constructed from 3. Inclusive graphic standard deviation, aI' the best es­ thick string with horizontal and vertical axes at 30 cm inter­ timator of overall sorting. vals. Each intersection of axes was projected visually against the quarry face and the sample was taken at that point. Slump­ Formulae for these parameters are presented in Folk (1974). ing of the quarry face made the lower cross-bedded sand at Norden Bridge generally inaccessible to the sampling scheme. The graphic mean, Mz' and the graphic standard devia­ Therefore, samples were taken from the Egelhoff sands and tion, aG , of the coarse sample from each of the two beds only from the upper bed of cross-bedded sand at Norden were used to perform Student's t-test of the null hypothesis Bridge Quarry. that there is no significant grain size difference between the 78 C. F. Wellstead

99.99 9999

99.95 99.95 999 99.9

99.5 995

--- COARSE SPLIT 99 --- COARSE SPLIT 99 --- HOMOGENEOUS SPLIT 1 ------HOMOGENEOUS SPLIT 1 J 98 ------HOMOGENEOUS SPLIT 2 / 98 ------HOMOGENEOUS SPLIT 2 / HOMOGENEOUS SPLIT 3 / HOMOGENEOUS SPLIT 3 ,'// ., 95 / ---- FINE SPLIT '// ----- FINE SPLIT j / 95 /// g:>'./ // 90 . / 90 1/ ! / /1 / ;; 80 f so ... ) Ii II ~ / 70 ,1/ 70 iii I ~ 60 t 60 ... / I r il I Q 50 50 J i/ I ~ ,) 40 40 " I ~ ;1 I 30 if I 30 3 // I 20 ! 20 !": I u i I 10 l::,1 I 10 " I / / I ,i ,! / / :' / / ,/ / / " /

0.5 /.// 0.5 / / ,/ / 0.2 .oX / 0.2 0.1 0.1 /' / - 0.05 ,-;{ // 0.05 , f _.---_--,-__-,- __.---_--,- __-,- __.---_--,-_--L- 0.01 -r-----,---....,--,---,--..,----,---,---'-001 -3 -2 -1 o .1 +2 +3 -3 -2 -1 +1 +2 +3 +4

PHI SCALE PHI SCALE

FIGURE 5. Cumulative curves: upper cross-bedded sand FIGURE 6. Cumulative curves: Egelhoff Quarry. at Norden Bridge Quarry. cross-bedded sands of Egelhoff Quarry and the upper bed The probability of obtaining a t value less than 2.131 of cross-bedded sands at Norden Bridge Quarry. The level of merely by chance were Ho true is greater than 0.05. The null probability chosen for the test was taken at 0.05. N is the hypothesis cannot be rejected. No significant difference in number of 0 intervals. grain size between the two beds of cross-bedded sands could be demonstrated. Calculation of standard e"or: Average sorting, aI' for the sediment samples from S(f = aG Norden Bridge + aG Egelhoff = 0.029 (N = 16) Norden Bridge Quarry is 0.538; that for the samples from N Egelhoff Quarry is 0.666. Both averages are within the moder­ t-test: ately well sorted category of Folk (1974), but are significantly HO: MZNorden Bridge = MZEgelhoff different: H : MzN. B. 4oMzE. l Calculation of standard error: MZE - MZN BOIS t-calc: . S(f " = 0:209 = 0.718 S2 = si + s~ = 0.0002135 + 0.00346 - 0.000735 d N 5 t(0.05, 15) = 2.131. Sd = 0.02711

.....J Sedimentology of Norden Bridge and Egelhoff quarries 79

t-test: The results of this last test suggest that, while there is no significant difference in average grain size between the two H:u =a o I Norden Bridge I Egellioff beds of sand, there is a wider size range about the mean at Egelhoff Quarry, as is borne out by aI values for samples HI: u1 F~ from Egellioff Quarry in Table IV. N.B. E. t =0.666 - 0.538 =0.128 =4 722 Analysis of Variance. An analysis of variance of the sieve (calc) Sd" 0.02711· analysis can be performed, considering the sieve analysis as a block design in which the quarries are treated as blocks, the t(4, 0.05) = 2.776 samples as units within blocks, and f/J size as treatment. Be­ cause each sample was processed through the same stack of The probability of obtaining a t value greater than 2.776 sieves, the assignment of treatment to each unit was not ran­ ,nerely by chance if Ho were true is less than 0.05. The null dom, but was decided prior to the sieve analysis. !lypothesis can be rejected and it can be concluded that there '$ a significant difference in sorting in the two cross-bedded The analysis of variance (Table V) shows no Significant lands compared. variation in f/J-class sediment weights attributable to differ­ ences between the quarries or to differences between samples. TABLE IV. Sediment sample statistics. The significant variation in ~-class sediment weights is a direct result of the sieve analysis, as more clasts were retained on

Mz aG aJ some screens than on others. More difficult to explain is the Quarry Sample (0) (0) (0) variation indicating significant interactions between ~ size and samples (s) and ~ size and quarries (q). These interactions occurred during the sieving and reveal some differential effect Norden Bridge coarse 1.95 0.525 0.535 of the sieving on some samples, but not on others, which homogeneous 1 2.48 0.475 0.525 affected samples of one quarry differently from those of the homogeneous 2 2.48 0.475 0.533 other. homogeneous 3 2.48 0.475 0.533 frne 2.53 0.055 0.563 Since the samples were treated alike, it is difficult to imagine what caused these interactions. Perhaps clast shape Fgelhoff coarse 2.10 0.650 0.719 and roundness, the result of subtle intra- and inter-quarry homogeneous 1 2.25 0.550 0.616 differences in source area, and distance travelled by the homogeneous 2 2.17 0.650 0.719 clasts affected their negotiation of the sieve openings. Dif­ homogeneous 3 2.18 0.600 0.686 ferences in mineralogy, thus specific gravity, of the clasts frne 2.65 0.550 0.593 may have had some effect also. However, no appropriate mineralogical study of the sediments at the two quarries has

TABLEV. Analysis of variance for sediment samples.

Sc-urce df SS MS F calc. Results of F-test EMS rnal 159 15,855.716 QHarry 15.194 15.194 2.374 not significant at p 0.05 a 2 + 80 g2 = q Sample 4 48.592 12.148 not significant at p 0.05 a 2 + 32 g2 1.898 = s (jI·dass 15 13,245.502 883.034 137.991 significant at p < 0.05 a2+10~ q I' S 4 10.605 2.651 0.414 not significant at p = 0.05 a 2 +16g2 q x ~ 2 s>. 0 60 1,883.115 31.385 4.905 Significant at p < 0.05 a + 2 ~ x 0 2 q;d~ 15 268.758 17.917 2.800 significant at p < 0.05 a + 5~ x 0 q:xsxc,l 60 383.950 6.399 a 2 80 C. F. Wellstead been attempted. Because the explanations offered are only Lacking access to the trough axis, the strike and apparent guesses, the interactions in the analysis of variance remain a dip of the foresets of both limbs of a trough cross-strata set difficulty. may be combined to calculate the azimuth of the set. How­ ever, because the characteristic interfering pattern of trough cross-strata sets originates from a scour-and-fill style of deposi­ ANALYSIS OF PALEOCURRENT INDICATORS tion, both limbs of a trough set are rarely preserved at Norden AT NORDEN BRIDGE Bridge and Egelhoff quarries. As a result of this limitation, AND EGELHOFF QUARRIES the field data collected from each set of trough cross-strata were readings of strike and apparent dip of foresets revealed The results of a study of directional features at Norden in each of two vertical, planar surfaces cut into the preserved Bridge and Egelhoff quarries are presented in this section. trough set limb. These raw data were converted into a true Directional readings were taken from cross-strata sets (sensu dip and strike for each cross-strata set following the method McKee and Weir, 1953) at both quarries and from elongate described by Billings (1972). Azimuths were then calculated fossil bones at Norden Bridge Quarry. No suitably large bones for each dip and strike reading. In all, azimuths were calcu­ were encountered in the course of this portion of the study at lated for 47 sets of cross-strata at Norden Bridge Quarry and Egelhoff Quarry. for 58 sets at Egelhoff Quarry. The raw data, corrected data, and azimuth for each cross-strata set are recorded in work by Cross-strata as Paleocurrent Indicators Wellstead (1977). Circular histograms of the azimuths are presented in Figure 7 A and B. Potter and Pettijohn (1977) concluded that cross-bedding studies are good indicators of local flow direction. However, Fossil Bones as Paleocurrent Indicators limitations imposed upon field work by the outcrop must be acknowledged. For example, the criteria of Allen's (1963) In what Dodson (1980) recognized as the classic work in detailed classification of cross-strata types are of little practi­ vertebrate taphonomy, Voorhies (1969: 11) used fossil bone cal use if the cross-strata studied cannot be exposed in three orientations to estimate current direction. More recently dimensions. When it is necessary to work with trough cross­ similar estimates have been made in a study by Hunt (1978). strata [see Michelson and Dott (1973) and Pi and Nu types of Allen (1963)1 as at Norden Bridge and Egelhoff quarries During the collection of cross-strata orientation data for (discussed below), it is preferable to measure the axis of the the present study, orientations of any elongate bones (mini­ trough rather than the dip of the foresets (Dott, 1973; Potter mum 10 cm in greatest length) discovered were recorded. and Pettijohn, 1977). Unfortunately, the nature of many These orientation data consist of the plunge of the bone and outcrops renders the axis of the trough inaccessible. This is the trend of its horizontal projection. Eleven such bones were the case at Norden Bridge Quarry, where unpredictable slump­ encountered at Norden Bridge Quarry at this time as no large ing of the quarry face precluded excavation of the cross-strata scale fossil excavations were conducted simultaneously. No sets, and at Egelhoff Quarry where the ponderously overhang­ elongate bone fragment was encountered at Egelhoff Quarry ing, white siltstone layer made such excavation not only during this study. A circular histogram for the bone orienta­ impossible in the absence of heavy equipment, but also dan­ tion data is presented in Figure 7C. gerous. Analysis of Orientation Data Forced to evaluate the type of cross-strata at the two fossil quarries in two dimensions only, the sets are interpreted Statistical Tests. Several authors (e.g., Curray, 1956; to be the trough type upon the following criteria: Pincus, 1956; Pelletier, 1958; Potter and Pettijohn, 1977) discussed the uncertainty inherent in representing the pre­ 1. Upper and lower surfaces of the sets are convergent. ferred orientation of sedimentary structures and fossils by an arithmetic mean and standard deviation. Calculation of 2. Lower surfaces of the sets are curved, concave upward. the mean vector and the mean angular deviation for the body of data is one solution. Till's (1974) method was used for this 3. The sets are present as interfering groups. calculation and the resulting mean vectors and mean angular deviations for cross-strata azimuths and bone orientations These criteria are in general accordance with discussions of are presented in Figure 7. trough cross-strata presented by McKee and Weir (1953), Allen (1963), Blatt, Middleton, and Murray (1972), Petti­ A hybrid F-test was used to test the null hypothesis john, Potter, and Siever (1972), and Potter and Pettijohn that the distribution of a set of directional features, such as (1977, Fig. 4.2). cross-strata azimuths from either quarry or the fossil bone Sedimentology of Norden Bridge and Egelhoff quarries 81 A N B N

'"... ("") 6 7 c N

FIGURE 7. Rose diagrams of directional features at Norden Bridge and Egelhoff quarries. A. Cross-strata azimuths at Norden Bridge Quarry. Mean vector, 305 ± 75, mean angular deviation, 75.489°. B. Cross-strata azimuths at Egelhoff Quarry. Mean vector 202 ± 56, mean angular deviation, 56.047°. C. Bone trends at Norden Bridge Quarry. Mean vector 222 ± 77, mean angular devia­ tion, 76.933°. orientations, does not differ significantly from a uniform dis­ In this test, degrees of freedom for both denominator and tribution. If the set of directional features does differ signifi­ numerator are n-1. The probability level, chosen arbitrarily, c;,ntly from the uniform distribution, the test assumes it to be is 0.05. unimodal. The test does not discriminate bimodal or poly­ modal distributions. This potential limitation is not serious 1. Norden Bridge Quarry, cross-strata data. if the Valentine Formation sediments are fluvial, as generally agreed, and thereby, unimodal. N=47

The test term (error term) in the F-test is (180°)2 (= the (180°)2 = 1.895 sc;uare of the maximum by which an azimuth may vary from (± 75.489°) tre true stream flow direction). The mean angular deviation (I ig. 7) is used as the denominator in the F-test. The tests F(O .05, 46, 46) = 1.67. ard their results follow: The probability of obtaining an F-value greater than 1.67 For each F-test: merely by chance if Ho were true is less than 0.05. Grounds exist to reject Ho and to conclude that the data differ signifi­ HO: The body of data does not differ significantly cantly from a uniform distribution. from a uniform distribution. 2. Norden Bridge Quarry, fossil bone orientations. HI: The body of data differs significantly from a uni­ form distribution. N=l1 82 C. F. Wellstead

SUMMARY

The measured section presented in this report confirms that Egelhoff Quarry is topographically higher than Norden F(0.05, 10, 10) = 2.98 Bridge Quarry, but that a demonstration of their relative stratigraphic positions is impossible as the relationship of Egel­ The probability of obtaining an F-value less than 2.98 merely hoff Quarry to the Valentine-Rosebud formational contact by chance if Ho were true is greater than 0.05. No grounds is not exposed and because the Egelhoff Quarry horizon can­ exist to reject Ho or to conclude that the data differ signifi­ not be traced toward Norden Bridge due to vegetation cover cantly from a uniform distribution. and erosion of the section.

3. Egelhoff Quarry, cross-strata data. Description of the sediments at the two quarries supports Holman's (1973) contention that coarser sediments are found N= 58 at Norden Bridge. However, it also reveals that these coarse sediments and fossil remains of large vertebrates are restricted to two beds and do not characterize the entire quarry. Beds of sediments finer than those at Egelhoff Quarry and at least one bed which shows no significant difference in mean grain size from Egelhoff Quarry sediments, exist as well at Norden F (0.05,57, 57) = 1.96 Bridge Quarry.

The probability of obtaining an F-value greater than 1.96 While no test of the hydraulic equivalency of fossil bones merely by chance if Ho were true is less than 0.05. Grounds from the Norden Bridge and Egelhoff quarries has been con­ exist to reject Ho and to conclude that the data differ signifi­ ducted, the isolation of remains of large vertebrates in the beds cantly from a uniform distribution. of coarser sediments at Norden Bridge and their general exclusion from the finer sediments of both quarries suggests Discussion of Test Results. The statistical tests indicate that the remains are hydraulically sorted. This probability that the cross-strata have a preferred orientation at both supports the recent work of Korth (1979) and also Holman's Norden Bridge and Egelhoff quarries. In both instances the (1976) suggestion that the faunal differences between the two implied current direction (N55W at Norden Bridge Quarry quarries are the result of depositional factors rather than and S33W at Egelhoff Quarry) is anomalous relative to the being due to actual differences in the local faunas. easterly regional dip of Tertiary strata in the area (Bentall, et aZ., 1971). However, as recognized by Potter and Pettijohn Mean vectors derived from cross-strata sets indicate that (1977), Steinmetz (1975), and Pettijohn, Potter, and Siever paleocurrents flowed northwesterly at Norden Bridge Quarry (1972), the directional features at any point along a stream are and southwesterly at Egelhoff Quarry. These results are sur­ not likely to correspond exactly with the overall direction of prising considering the easterly regional dip of local Tertiary streamflow in the basin. strata, but emphasize the desirability of a regional approach to paleocurrent analysis. To obtain an adequate indication of paleocurrents within the Valentine Formation, data must be collected and evaluated on a regional basis. The practical difficulties encountered in ACKNOWLEDGMENTS this brief study of paleocurrents stem from the lack of three­ dimensional access to the cross-strata and indicate that prior The patient guidance of Michael R. Voorhies throughout to beginning a regional study, a preliminary assessment of the completion of the master's thesis from which this report relative proportions of cross-strata type within the formation was extracted is gratefully acknowledged. The Conserva­ and the amount of excavation necessary to expose them prop­ tion and Survey Division of the University of Nebraska is erly should be completed. Steinmetz (1975) suggested a pre­ thanked for allowing access to its laboratory facilities. James liminary, regional sampling of orientation data to assess B. Swineheart, Research Geologist, provided advice and variability at each outcrop and to assist in estimating the instruction pertinent to the sedimentological portion of this number of orientation readings necessary for each site. report.

Absence of statistical significance in the sample of bone Robert L. Evander, now a graduate student at Columbia orientations may be due to actual lack of anyone preferred University, suggested the sediment sampling scheme used in orientation, or to small sample size. the field and lent critical support to the project. Sedimentology of Norden Bridge and Egelhoff quarries 83

Research was supported by Schramm and Shell scholar­ Local Fauna in north-central Nebraska. Contributions ;;hips from the University of Nebraska-Lincoln Department of from the Museum of Paleontology, University of Michi­ Geology, a research assistantship from U.S. Bureau of Reclam­ gan, 23(15):239-246. .ltion Contract No. 14-06-700-837 awarded to Carl R. Falk, \nthropology Department, University of Nebraska-lincoln, Curray, J. R. 1956. The analysis of two dimensional orienta­ ,md a Grant-in-Aid of Research from Sigma Xi. tion data.Journal of Geology, 64(2):117-131.

Morris F. Skinner, Michael R. Voorhies, and Brent B. Dodson, P. 1971. The significance of small bones in paleo­ ~~ickol reviewed the manuscript and offered many suggestions ecological interpretation. University of Wyoming, Con­ (or its improvement. tributions to Geology, 12:15-19.

__. 1980. Vertebrate burials. Paleobiology, 6(1):6-8. REFERENCES Dott, R. H., Jr. 1973. Paleocurrent analysis of trough cross Allen, J. R. L. 1963. The classification of cross stratified units stratification. Journal of Sedimentary Petrology, 43(3): with notes on their origin. Sedimentology, 2(2):93-114. 779-783.

Anonymous. 1977. Final environmental statement, O'Neill Falk, C. R., A. J. Osborn, R. E. Pepped, and M. R. Voorhies. Unit, Nebraska, Appendix A, geologic data. Washington, 1980. Cultural and paleontological investigations within United States Department of the Interior, Bureau of the proposed Norden Reservoir area, Nebraska: an in­ Reclamation. terim report. Division of Archeological Research, Depart­ ment of Anthropology, University of Nebraska, Lincoln, Hentall, R., and others. 1971. Water supplies and the land. Technical Report No. 80-05: 1-35 and appendix. Elkhorn River Basin of Nebraska. Resource Atlas 1. lincoln, Nebraska, Conservation and Survey Division: Folk, R. L. 1974. Petrology of sedimentary rocks. Austin, 1-50. Texas, Hemphill Publishing Company: 182p.

Berggren, W. A. 1972. A Cenozoic time scale. Some implica­ Gidley, J. W. 1904. New or little known mammals from the tions for regional geology and paleobiogeography. Lethaia, Miocene of South Dakota. Bulletin of the American 5:192-215. Museum ofNatural History, 22:241-268.

, and J. A. Van Couvering. 1974. The Late Neogene. Griffiths, J. C., and M. A. Rosenfeld. 1953. A further test of Biostratigraphy, geochronology, and paleontology of the dimensional orientation of quartz grains in Bradford sand. last 15 million years in marine and continental sequences. American Journal of Science, 152:192-214. Palaeogeography, Palaeoclimatology, Palaeoecology, 16 (1/2):1-216. Hibbard, C. W. 1960. An interpretation of Pliocene and Pleistocene climates in North America. Annual Report of BUlings, M. P. 1972. Structural geology, 3rd ed. Englewood the Michigan Academy of Sciences, Arts and Letters, Cliffs, New Jersey, Prentice Hall, Incorporated: 573p. 61 :5-30.

Blatt, H., G. Middleton, and R. Murray. 1972. Origin of sedi­ Holman, J. A. 1973. Reptiles of the Egelhoff Local Fauna mentary rocks. Englewood Cliffs, New Jersey, Prentice (Upper Miocene) of Nebraska. Contributions from the Hall, Incorporated: 634p. Museum of Paleontology, University of Michigan, 24: 125-134. B,)ellstorff, J. 1978. Chronology of some Late Cenozoic de­ posits from the central United States and the Ice Ages. __. 1976. The herpetofauna of the Lower Valentine Forma­ Transactions of the Nebraska Academy of Sciences, tion, north-central Nebraska. Herpetologica, 32:262-268. 6:35-49. Hunt, R. M., Jr. 1978. Depositional setting of a Miocene , and M. F. Skinner. 1977. A fission track date from mammal assemblage, Sioux County, Nebraska (U.S.A.). post-Rosebud, Early Valentine rocks. Proceedings of the Palaeogeography, Palaeoclimatology, Palaeoecology," 24: Nebraska Academy ofSciences, 87:39-40. 1-52.

2hantell, C. J. 1971. Fossil amphibians from the Egelhoff Klingener, D. 1968. of the Mio-Pliocene Norden 84 C. F. Wellstead

Bridge Local Fauna, Nebraska. American Midland Natural­ Nebraska. Bulletin of the American Museum of Natural ist, 80:65-71. History, 148:1-148.

Korth, W. W. 1979. Taphonomy of microvertebrate fossil __, S. M. Skinner, and R. J. Gooris. 1968. Cenozoic rocks assemblages. Annals of the Carnegie Museum, 48:235- and faunas of Turtle Butte, south-central South Dakota. 285. Bulletin of the American Museum of Natural History, 138:379-436. Leidy, J. 1858. Notice of remains of extinct Vertebrata from the valley of the Niobrara River. Proceedings of the Aca­ Smith, C. L. 1962. Some Pliocene fishes from Kansas, Okla­ demy ofNatural Sciences, Philadelphia, 10:20-29. homa and Nebraska. Copeia, 1962:505-520. lindsay, E. H. 1972. Small mammal fossils from the Barstow Stanley, K. 0., and W. J. Wayne. 1972. Epeirogenic and cli­ Formation, California. University of California Publica­ matic controls of Early Pleistocene fluvial sediment dis­ tions in Geological Sciences, 93:1-104. persal in Nebraska. Bulletin of the Geological Society of America, 83 :3675-3690. Macdonald, J. R, and J. C. Harksen. 1968. Rosebud Forma­ tion in South Dakota. South Dakota Geological Survey Steel, R. G. D., and J. H. Torrie. 1960. Principles and pro­ Report ofInvestigations, 57. cedures of statistics. New York, McGraw-Hill Book Com­ pany: 481p. MacGinitie, H. D. 1962. The Kilgore Flora, a Late Miocene flora from northern Nebraska. University of California Steinmetz, R. 1975. Cross-bed variability in a single sand Publications in Geological Sciences, 35: 67-158. body. Memoirs of the Geological Society of America, 142:89-102. McKee, E. D., and G. W. Weir. 1953. Terminology for strati­ fication and cross-stratification in sedimentary rocks. Storer, J. E. 1973. The entoptychine geomyid Lignimus Bulletin of the Geological Society of America, 64:381- (Mammalia, Rodentia) from Kansas and Nebraska. Cana­ 390. dian Journal of Earth Sciences, 10(1):72-83.

Michelson, P. C., and R H. Dott, Jr. 1973. Orientation analy­ Tedford, R. H., T. Galusha, M. F. Skinner, B. E. Taylor, sis of trough cross-stratification in Upper R. Fields, J. R. Macdonald, T. H. Patton, J. M. Rens­ sandstones of western Wisconsin. Journal of Sedimentary berger, and D. P. Whistler. 1981. Faunal succession and Petrology, 43:784-794. biochronology of the Arikareean through Hemphillian interval (Late Oligocene through Late Miocene Epochs) Pelletier, B. R 1958. Pocono paleocurrents in Pennsylvania North America. University of California Publications in and Maryland. Bulletin of the Geological Society of Geological Sciences, In press. America, 69: 1033-1064. Tihen, J. A., and C. J. Chantell. 1963. Urodele remains from Pettijohn, F. J., P. E. Potter, and R Siever. 1972. Sand and the Valentine Formation of Nebraska. Copeia, 1963: sandstone. New York, Springer-Verlag: 618p. 505-510.

Pincus, H. J. 1956. Some vector and arithmetic operations on Till, R. 1974. Statistical methods for the earth scientist. New two dimensional orientation variation with application to York, John Wiley and Sons: 154p. geological data. Journal of Geology, 64(6):533-557. Voorhies, M. R. 1969. Taphonomy and population dynamics Potter, P. E., and F. J. Pettijohn. 1977. Paleocurrents and of an Early Pliocene vertebrate fauna, Knox County, basin analysis, 2nd. ed. Berlin, Springer-Verlag: 425p. Nebraska. University of Wyoming Contributions to Geology, Special Paper No.1: 1-69. Rich, T. H. V., and D. L. Rasmussen. 1973. New North Amer­ ican erinaceine hedgehogs (Mammalia: Insectivora). __. 1973. Early Miocene mammals from northeast Nebras­ Occasional Papers, Museum of Natural History, Univer­ ka. University of Wyoming Contributions to Geology, sity of Kansas, 21: 1-54. 12:1-10.

Skinner, M. F., and C. W. Hibbard. 1972. Early Pleistocene Webb, S. D. 1969. The Burge and Minnechaduza Claren­ pre-glacial and glacial rocks and faunas of north-central donian mammalian faunas of north-central Nebraska. Sedimentology of Norden Bridge and Egelhoff quarries 85

University of California Publications in Geological Sci­ Wood, H. C., R. W. Chaney, J. Clark, E. H. Colbert, G. L. ences, 78:1-91. Jepsen, J. B. Reeside, Jr., and C. Stock. 1941. Nomen­ clature and correlation of the North American continental Wellstead, C. F. 1977. Fossil lizards of the Valentine Forma­ Tertiary. Bulletin of the Geological Society of America, tion. Master of Science Thesis, University of Nebraska­ 52:1-48. lincoln: 159p.