APPENDIX 3: List of Autapomorphic Characters for Taxa Involved in Phylogenetic Analysis. Millerettidae -No Known Autapomorphies
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JVP 26(3) September 2006—ABSTRACTS
Neoceti Symposium, Saturday 8:45 acid-prepared osteolepiforms Medoevia and Gogonasus has offered strong support for BODY SIZE AND CRYPTIC TROPHIC SEPARATION OF GENERALIZED Jarvik’s interpretation, but Eusthenopteron itself has not been reexamined in detail. PIERCE-FEEDING CETACEANS: THE ROLE OF FEEDING DIVERSITY DUR- Uncertainty has persisted about the relationship between the large endoskeletal “fenestra ING THE RISE OF THE NEOCETI endochoanalis” and the apparently much smaller choana, and about the occlusion of upper ADAM, Peter, Univ. of California, Los Angeles, Los Angeles, CA; JETT, Kristin, Univ. of and lower jaw fangs relative to the choana. California, Davis, Davis, CA; OLSON, Joshua, Univ. of California, Los Angeles, Los A CT scan investigation of a large skull of Eusthenopteron, carried out in collaboration Angeles, CA with University of Texas and Parc de Miguasha, offers an opportunity to image and digital- Marine mammals with homodont dentition and relatively little specialization of the feeding ly “dissect” a complete three-dimensional snout region. We find that a choana is indeed apparatus are often categorized as generalist eaters of squid and fish. However, analyses of present, somewhat narrower but otherwise similar to that described by Jarvik. It does not many modern ecosystems reveal the importance of body size in determining trophic parti- receive the anterior coronoid fang, which bites mesial to the edge of the dermopalatine and tioning and diversity among predators. We established relationships between body sizes of is received by a pit in that bone. The fenestra endochoanalis is partly floored by the vomer extant cetaceans and their prey in order to infer prey size and potential trophic separation of and the dermopalatine, restricting the choana to the lateral part of the fenestra. -
Sauropareion Anoplus, with a Discussion of Possible Life History
The postcranial skeleton of the Early Triassic parareptile Sauropareion anoplus, with a discussion of possible life history MARK J. MACDOUGALL, SEAN P. MODESTO, and JENNIFER BOTHA−BRINK MacDougall, M.J., Modesto, S.P., and Botha−Brink, J. 2013. The postcranial skeleton of the Early Triassic parareptile Sauropareion anoplus, with a discussion of possible life history. Acta Palaeontologica Polonica 58 (4): 737–749. The skeletal anatomy of the Early Triassic (Induan) procolophonid reptile Sauropareion anoplus is described on the basis of three partial skeletons from Vangfontein, Middelburg District, South Africa. Together these three specimens preserve the large majority of the pectoral and pelvic girdles, articulated forelimbs and hindlimbs, and all but the caudal portion of the vertebral column, elements hitherto undescribed. Our phylogenetic analysis of the Procolophonoidea is consonant with previous work, positing S. anoplus as the sister taxon to a clade composed of all other procolophonids exclusive of Coletta seca. Previous studies have suggested that procolophonids were burrowers, and this seems to have been the case for S. anoplus, based on comparisons with characteristic skeletal anatomy of living digging animals, such as the presence of a spade−shaped skull, robust phalanges, and large unguals. Key words: Parareptilia, Procolophonidae, phylogenetic analysis, burrowing, Induan, Triassic, South Africa. Mark J. MacDougall [[email protected]], Department of Biology, Cape Breton University, Sydney, Nova Scotia, B1P 6L2, Canada and Department of Biology, University of Toronto at Mississauga, 3359 Mississauga Road, Ontario, L5L 1C6, Canada; Sean P. Modesto [[email protected]], Department of Biology, Cape Breton University, Sydney, Nova Scotia, B1P 6L2, Canada; Jennifer Botha−Brink [[email protected]], Karoo Palaeontology, National Museum, P.O. -
Ontogenetic Change in the Temporal Region of the Early Permian Parareptile Delorhynchus Cifellii and the Implications for Closure of the Temporal Fenestra in Amniotes
RESEARCH ARTICLE Ontogenetic Change in the Temporal Region of the Early Permian Parareptile Delorhynchus cifellii and the Implications for Closure of the Temporal Fenestra in Amniotes Yara Haridy*, Mark J. Macdougall, Diane Scott, Robert R. Reisz Department of Biology, University of Toronto Mississauga, Mississauga, Ontario, Canada * [email protected] a11111 Abstract A juvenile specimen of Delorhynchus cifellii, collected from the Early Permian fissure-fill deposits of Richards Spur, Oklahoma, permits the first detailed study of cranial ontogeny in this parareptile. The specimen, consisting of a partially articulated skull and mandible, exhib- OPEN ACCESS its several features that identify it as juvenile. The dermal tuberosities that ornament the dor- Citation: Haridy Y, Macdougall MJ, Scott D, Reisz sal side and lateral edges of the largest skull of D. cifellii specimens, are less prominent in RR (2016) Ontogenetic Change in the Temporal the intermediate sized holotype, and are absent in the new specimen. This indicates that the Region of the Early Permian Parareptile new specimen represents an earlier ontogenetic stage than all previously described mem- Delorhynchus cifellii and the Implications for bers of this species. In addition, the incomplete interdigitation of the sutures, most notably Closure of the Temporal Fenestra in Amniotes. PLoS ONE 11(12): e0166819. doi:10.1371/journal. along the fronto-nasal contact, plus the proportionally larger sizes of the orbit and temporal pone.0166819 fenestrae further support an early ontogenetic stage for this specimen. Comparisons Editor: Thierry Smith, Royal Belgian Institute of between this juvenile and previously described specimens reveal that the size and shape of Natural Sciences, BELGIUM the temporal fenestra in Delorhynchus appear to vary through ontogeny, due to changes in Received: July 18, 2016 the shape and size of the bordering cranial elements. -
Analysis of Millerettid Parareptile Relationships in the Light of New Material of BROOMIA PERPLEXA Watson, 1914, from the Permian of South Africa
Journal of Systematic Palaeontology 6 (4): 453–462 Issued 24 November 2008 doi:10.1017/S147720190800254X Printed in the United Kingdom C The Natural History Museum ! Analysis of millerettid parareptile relationships in the light of new material of BROOMIA PERPLEXA Watson, 1914, from the Permian of South Africa Juan Carlos Cisneros Departamento de Paleontologia† e Estratigrafia, Universidade Federal do Rio Grande do Sul, Porto Alegre, CP 15001, 91501-970, Brazil; and Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, Private Bag 3, WITS 2050, Johannesburg, South Africa Bruce S. Rubidge Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, Private Bag 3, WITS 2050, Johannesburg, South Africa Richard Mason Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, Private Bag 3, WITS 2050, Johannesburg, South Africa Charlton Dube Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, Private Bag 3, WITS 2050, Johannesburg, South Africa SYNOPSIS A second specimen of the poorly known millerettid Broomia perplexa is reported. It consists of a cranium and partially articulated postcranium. As indicated by its small size (ca.35% shorter skull than the holotype) and unfused pelvis and limbs, the new specimen is a sub-adult indi- vidual. Comparison with the holotype reveals only minor differences that are ascribed to ontogenetic or individual variation. Accordingly we consider the new specimen to represent the second record of Broomia,discovered90yearsafterthedescriptionoftheholotype.Apreliminaryphylogenetic analysis of millerettid inter-relationships identifies Broomia as a millerettid related to the genus Millerosaurus.TheenigmaticparareptileEunotosaurus africanus is nested within Millerettidae as the sister taxon of Milleretta rubidgei.WhereasmillerettidsarewellknownfromthelatestPermian Dicynodon Assemblage Zone of the Beaufort Group of South Africa, Broomia and Eunotosaurus are found in the Middle Permian Tapinocephalus Assemblage Zone. -
Reptile Family Tree
Reptile Family Tree - Peters 2015 Distribution of Scales, Scutes, Hair and Feathers Fish scales 100 Ichthyostega Eldeceeon 1990.7.1 Pederpes 91 Eldeceeon holotype Gephyrostegus watsoni Eryops 67 Solenodonsaurus 87 Proterogyrinus 85 100 Chroniosaurus Eoherpeton 94 72 Chroniosaurus PIN3585/124 98 Seymouria Chroniosuchus Kotlassia 58 94 Westlothiana Casineria Utegenia 84 Brouffia 95 78 Amphibamus 71 93 77 Coelostegus Cacops Paleothyris Adelospondylus 91 78 82 99 Hylonomus 100 Brachydectes Protorothyris MCZ1532 Eocaecilia 95 91 Protorothyris CM 8617 77 95 Doleserpeton 98 Gerobatrachus Protorothyris MCZ 2149 Rana 86 52 Microbrachis 92 Elliotsmithia Pantylus 93 Apsisaurus 83 92 Anthracodromeus 84 85 Aerosaurus 95 85 Utaherpeton 82 Varanodon 95 Tuditanus 91 98 61 90 Eoserpeton Varanops Diplocaulus Varanosaurus FMNH PR 1760 88 100 Sauropleura Varanosaurus BSPHM 1901 XV20 78 Ptyonius 98 89 Archaeothyris Scincosaurus 77 84 Ophiacodon 95 Micraroter 79 98 Batropetes Rhynchonkos Cutleria 59 Nikkasaurus 95 54 Biarmosuchus Silvanerpeton 72 Titanophoneus Gephyrostegeus bohemicus 96 Procynosuchus 68 100 Megazostrodon Mammal 88 Homo sapiens 100 66 Stenocybus hair 91 94 IVPP V18117 69 Galechirus 69 97 62 Suminia Niaftasuchus 65 Microurania 98 Urumqia 91 Bruktererpeton 65 IVPP V 18120 85 Venjukovia 98 100 Thuringothyris MNG 7729 Thuringothyris MNG 10183 100 Eodicynodon Dicynodon 91 Cephalerpeton 54 Reiszorhinus Haptodus 62 Concordia KUVP 8702a 95 59 Ianthasaurus 87 87 Concordia KUVP 96/95 85 Edaphosaurus Romeria primus 87 Glaucosaurus Romeria texana Secodontosaurus -
A Reassessment of the Taxonomic Position of Mesosaurs, and a Surprising Phylogeny of Early Amniotes
GENERAL COMMENTARY published: 03 December 2018 doi: 10.3389/feart.2018.00220 Response: Commentary: A Reassessment of the Taxonomic Position of Mesosaurs, and a Surprising Phylogeny of Early Amniotes Michel Laurin 1* and Graciela Piñeiro 2 1 CR2P (UMR 7207), CNRS/MNHN Sorbonne Université, “Centre de Recherches sur la Paléobiodiversité et les Paléoenvironnements”, Muséum National d’Histoire Naturelle, Paris, France, 2 Departamento de Paleontología, Facultad de Ciencias, Montevideo, Uruguay Keywords: Mesosauridae, Parareptilia, Synapsida, Sauropsida, Amniota, Paleozoic, temporal fenestration A Commentary on Commentary: A Reassessment of the Taxonomic Position of Mesosaurs, and a Surprising Phylogeny of Early Amniotes by MacDougall, M. J., Modesto, S. P., Brocklehurst, N., Verrière, A., Reisz, R. R., and Fröbisch, J. (2018). Front. Earth Sci. 6:99. doi: 10.3389/feart.2018.00099 INTRODUCTION Edited by: Corwin Sullivan, University of Alberta, Canada Mesosaurs, known from the Early Permian of southern Africa, Brazil, and Uruguay, are the oldest known amniotes with a primarily, though probably not strictly, aquatic lifestyle (Nuñez Demarco Reviewed by: et al., 2018). Despite having attracted the attention of several prominent scientists, such as Wegener Tiago Simoes, University of Alberta, Canada (1966), who used them to support his theory of continental drift, and the great anatomist and paleontologist von Huene (1941), who first suggested the presence of a lower temporal fenestra *Correspondence: Michel Laurin in Mesosaurus, several controversies still surround mesosaurs. One concerns the presence of the [email protected] lower temporal fenestra in mesosaurs, which we accept (Piñeiro et al., 2012a; Laurin and Piñeiro, 2017, p. 4), contrary to Modesto (1999, 2006) and MacDougall et al. -
Developmental Genetic Bases Behind the Independent Origin of the Tympanic Membrane in Mammals and Diapsids
ARTICLE Received 8 Nov 2014 | Accepted 5 Mar 2015 | Published 22 Apr 2015 DOI: 10.1038/ncomms7853 OPEN Developmental genetic bases behind the independent origin of the tympanic membrane in mammals and diapsids Taro Kitazawa1,2,*, Masaki Takechi3,4,*,w, Tatsuya Hirasawa3, Noritaka Adachi3, Nicolas Narboux-Neˆme5, Hideaki Kume1, Kazuhiro Maeda6, Tamami Hirai3, Sachiko Miyagawa-Tomita6, Yukiko Kurihara1,2, Jiro Hitomi4, Giovanni Levi5, Shigeru Kuratani3 & Hiroki Kurihara1,2,7 The amniote middle ear is a classical example of the evolutionary novelty. Although paleontological evidence supports the view that mammals and diapsids (modern reptiles and birds) independently acquired the middle ear after divergence from their common ancestor, the developmental bases of these transformations remain unknown. Here we show that lower-to-upper jaw transformation induced by inactivation of the Endothelin1-Dlx5/6 cascade involving Goosecoid results in loss of the tympanic membrane in mouse, but causes duplication of the tympanic membrane in chicken. Detailed anatomical analysis indicates that the relative positions of the primary jaw joint and first pharyngeal pouch led to the coupling of tympanic membrane formation with the lower jaw in mammals, but with the upper jaw in diapsids. We propose that differences in connection and release by various pharyngeal skeletal elements resulted in structural diversity, leading to the acquisition of the tympanic membrane in two distinct manners during amniote evolution. 1 Department of Physiological Chemistry and Metabolism, Graduate School of Medicine, The University of Tokyo, 7-3-1, Hongo, Bunkyo-ku, Tokyo 113-0033, Japan. 2 Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency (JST), Chiyoda-ku, Tokyo 102-0076, Japan. -
Physical and Environmental Drivers of Paleozoic Tetrapod Dispersal Across Pangaea
ARTICLE https://doi.org/10.1038/s41467-018-07623-x OPEN Physical and environmental drivers of Paleozoic tetrapod dispersal across Pangaea Neil Brocklehurst1,2, Emma M. Dunne3, Daniel D. Cashmore3 &Jӧrg Frӧbisch2,4 The Carboniferous and Permian were crucial intervals in the establishment of terrestrial ecosystems, which occurred alongside substantial environmental and climate changes throughout the globe, as well as the final assembly of the supercontinent of Pangaea. The fl 1234567890():,; in uence of these changes on tetrapod biogeography is highly contentious, with some authors suggesting a cosmopolitan fauna resulting from a lack of barriers, and some iden- tifying provincialism. Here we carry out a detailed historical biogeographic analysis of late Paleozoic tetrapods to study the patterns of dispersal and vicariance. A likelihood-based approach to infer ancestral areas is combined with stochastic mapping to assess rates of vicariance and dispersal. Both the late Carboniferous and the end-Guadalupian are char- acterised by a decrease in dispersal and a vicariance peak in amniotes and amphibians. The first of these shifts is attributed to orogenic activity, the second to increasing climate heterogeneity. 1 Department of Earth Sciences, University of Oxford, South Parks Road, Oxford OX1 3AN, UK. 2 Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstraße 43, 10115 Berlin, Germany. 3 School of Geography, Earth and Environmental Sciences, University of Birmingham, Birmingham B15 2TT, UK. 4 Institut -
A New Permian Temnospondyl with Russian Affinities from South America, the New Family Konzhukoviidae, and the Phylogenetic Status of Archegosauroidea
Journal of Systematic Palaeontology ISSN: 1477-2019 (Print) 1478-0941 (Online) Journal homepage: http://www.tandfonline.com/loi/tjsp20 A new Permian temnospondyl with Russian affinities from South America, the new family Konzhukoviidae, and the phylogenetic status of Archegosauroidea Cristian Pereira Pacheco, Estevan Eltink, Rodrigo Temp Müller & Sérgio Dias- da-Silva To cite this article: Cristian Pereira Pacheco, Estevan Eltink, Rodrigo Temp Müller & Sérgio Dias-da-Silva (2016): A new Permian temnospondyl with Russian affinities from South America, the new family Konzhukoviidae, and the phylogenetic status of Archegosauroidea, Journal of Systematic Palaeontology To link to this article: http://dx.doi.org/10.1080/14772019.2016.1164763 View supplementary material Published online: 11 Apr 2016. Submit your article to this journal View related articles View Crossmark data Full Terms & Conditions of access and use can be found at http://www.tandfonline.com/action/journalInformation?journalCode=tjsp20 Download by: [Library Services City University London] Date: 11 April 2016, At: 06:28 Journal of Systematic Palaeontology, 2016 http://dx.doi.org/10.1080/14772019.2016.1164763 A new Permian temnospondyl with Russian affinities from South America, the new family Konzhukoviidae, and the phylogenetic status of Archegosauroidea Cristian Pereira Pachecoa,c*, Estevan Eltinkb, Rodrigo Temp Muller€ c and Sergio Dias-da-Silvad aPrograma de Pos-Gradua c¸ ao~ em Ciencias^ Biologicas da Universidade Federal do Pampa, Sao~ Gabriel, CEP 93.700-000, RS, Brazil; bLaboratorio de Paleontologia de Ribeirao~ Preto, FFCLRP, Universidade de Sao~ Paulo, Av. Bandeirantes 3900, 14040-901, Ribeirao~ Preto, Sao~ Paulo, Brazil; cPrograma de Pos Graduac¸ ao~ em Biodiversidade Animal, Universidade Federal de Santa Maria, Av. -
Chinese Pareiasaurs
Benton, M. J. (2016). The Chinese pareiasaurs. Zoological Journal of the Linnean Society, 177(4), 813-853. https://doi.org/10.1111/zoj.12389 Peer reviewed version License (if available): Unspecified Link to published version (if available): 10.1111/zoj.12389 Link to publication record in Explore Bristol Research PDF-document This is the author accepted manuscript (AAM). The final published version (version of record) is available online via Wiley at http://onlinelibrary.wiley.com/doi/10.1111/zoj.12389/abstract. Please refer to any applicable terms of use of the publisher. University of Bristol - Explore Bristol Research General rights This document is made available in accordance with publisher policies. Please cite only the published version using the reference above. Full terms of use are available: http://www.bristol.ac.uk/red/research-policy/pure/user-guides/ebr-terms/ Page 1 of 85 Zoological Journal of the Linnean Society 1 1 2 3 1 [Abstract] 4 5 2 6 3 Pareiasaurs were important medium- to large-sized herbivores in the Middle and Late 7 8 4 Permian, some 268-252 million years (Myr) ago. They are best known from abundant remains 9 10 5 of several taxa each in South Africa and Russia, with isolated finds from other parts of the 11 6 world. Six genera and species of pareiasaurs have been described from China, and yet they 12 13 7 have not been reviewed. Of these six, Tsiyuania may be a synonym of Honania, but this taxon 14 15 8 is not further considered here. The other four, which were named for separate finds from the 16 9 Sunjiagou Formation (Changhsingian, 254-252 Myr) show considerable similarities. -
Evidence of Advanced Sensory Perception at an Early Stage of Amniote Evolution Johannes Mu¨ Ller*, Linda A
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by PubMed Central Impedance-Matching Hearing in Paleozoic Reptiles: Evidence of Advanced Sensory Perception at an Early Stage of Amniote Evolution Johannes Mu¨ ller*, Linda A. Tsuji Humboldt-Universita¨t zu Berlin, Museum fu¨r Naturkunde, Berlin, Germany Background. Insights into the onset of evolutionary novelties are key to the understanding of amniote origins and diversification. The possession of an impedance-matching tympanic middle ear is characteristic of all terrestrial vertebrates with a sophisticated hearing sense and an adaptively important feature of many modern terrestrial vertebrates. Whereas tympanic ears seem to have evolved multiple times within tetrapods, especially among crown-group members such as frogs, mammals, squamates, turtles, crocodiles, and birds, the presence of true tympanic ears has never been recorded in a Paleozoic amniote, suggesting they evolved fairly recently in amniote history. Methodology/Principal Findings. In the present study, we performed a morphological examination and a phylogenetic analysis of poorly known parareptiles from the Middle Permian of the Mezen River Basin in Russia. We recovered a well-supported clade that is characterized by a unique cheek morphology indicative of a tympanum stretching across large parts of the temporal region to an extent not seen in other amniotes, fossil or extant, and a braincase specialized in showing modifications clearly related to an increase in auditory function, unlike the braincase of any other Paleozoic tetrapod. In addition, we estimated the ratio of the tympanum area relative to the stapedial footplate for the basalmost taxon of the clade, which, at 23:1, is in close correspondence to that of modern amniotes capable of efficient impedance-matching hearing. -
Cisneros2008.Pdf
Journal of Systematic Palaeontology 6 (3): 345–366 Issued 22 August 2008 doi:10.1017/S1477201907002350 Printed in the United Kingdom C The Natural History Museum ! Phylogenetic relationships of procolophonid parareptiles with remarks on their geological record Juan Carlos Cisneros∗ Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, South Africa SYNOPSIS The phylogenetic intrarelationships of procolophonid parareptiles are determined via a comprehensive cladistic analysis using a data matrix of 21 taxa and 58 characters. Most taxa are in- cluded for the first time in a phylogenetic analysis and 27 characters are novel. The relationships within the group are more firmly resolved than in previous analyses. Procolophoninae and Leptopleuron- inae, two of the three traditional subdivisions of the Procolophonidae, are valid monophyletic groups, but Spondylolestinae is polyphyletic. The Chinese genera Pentaedrusaurus and Neoprocolophon are the most primitive members of the Leptopleuroninae. A new group, Theledectinae, is erected. The latter clade consists of small procolophonids with a reduced marginal dentition and wide bulbous monocuspid teeth. Eumetabolodon from China and the former genus ‘Thelegnathus’ from South Africa are shown to be polyphyletic. The successful radiation of the Procolophonidae during the Triassic is likely to be related to the development of feeding adaptations that allowed exploration of various ecological niches, particularly the exploitation of high-fibre herbivory. The scarcity of Permian records of procolophonids is examined and the genus Spondylolestes from the Upper Permian of South Africa is considered to be a valid taxon with procolophonid affinities. Finally, a review of the records from the Middle and Upper Triassic reveals a procolophonid global hiatus of more than 15 Ma in Ladinian–Lower Carnian rocks.