Wetland Fields in the Maya Lowlands

Total Page:16

File Type:pdf, Size:1020Kb

Load more

Wetland Fields in the Maya Lowlands:
Archaeobotanical Evidence from Birds of Paradise, Belize

A thesis submitted to the
Graduate School of the University of Cincinnati in partial fulfillment of the requirements for the degree of
MASTER OF ARTS in the Department of Anthropology of the College of Arts and Sciences
2019 by
Martha M. Wendel
B.A., University of Cincinnati, 2015
Committee: Susan E. Allen, Ph.D., Chair
Sarah E. Jackson, Ph.D. David L. Lentz, Ph.D.
Abstract
Discoveries of rectangular canal patterns in the margins of wetlands in the ancient Maya lowlands of Guatemala, Belize, and Mexico shed light on a previously unknown agricultural practice: raised wetland fields. One example of wetland fields is found at the site Birds of Paradise (BOP) in the Rio Bravo region of northwestern Belize. For my research project, I have analyzed macrobotanical remains from BOP to: 1) identify the plants that were growing in the canals, 2) identify the plants that were growing in the features identified as raised fields, 3) assess their ecological preferences, 4) assess changing frequencies of different types of plants over time, and 5) provide some insight on how the canals and fields were used. Because this is the first time any systematic macrobotanical analysis has been done at BOP, it makes an important contribution to understanding how the Maya were interacting with their landscape with the use of these features. Additionally, innovative recovery methods such as use of a sonicator and sorting through all geological sieve fractions allowed for more robust quantitative analysis of data. These methods have wide application for use at other sites conducting archaeobotanical research.
Acknowledgments
There are so many people whom I have to thank for loving me and encouraging me during this incredibly stressful time and have given me so much support to continue in the field of archaeology. First, I would like to thank Dr. Susan Allen, who has given me so many opportunities to explore the field of archaeobotany and whose interest and enthusiasm for what she does continue to inspire me every day. I would be lost without your guidance!

Second, I would like to thank Dr. David Lentz who has taught me a lot about Maya paleoethnobotany and provided me with many opportunities for this research project. Third, I would like to thank Dr. Timothy Beach and Dr. Sheryl Luzzadder-Beach for providing me funding and for providing the opportunity to conduct research at Birds of Paradise. I would also like to thank Dr. Sarah Jackson for being on my committee and for teaching most of what I know about the ancient Maya. Thanks to Dr. Alan Sullivan for giving me great advice about life and archaeology and for providing me with the opportunity to explore the Southwest.

Additionally, I would like to thank Venicia Slotten for all of her help with collecting flotation samples, assistance with identification, and just listening to me when I needed to vent. Your friendship means so much to me. Thanks to Mariana Vazquez for help with identification of the plants collected from the survey. I would also like to thank the graduate students within my cohort at the University of Cincinnati, especially Emily Phillips and Ashley-Devon Williamston. Thanks to my parents for their support and for trying to understand what I do. Lastly, I would like to thank my partner, Rob Stambaugh, whose love and support gave me the strength I needed to complete one of the largest challenges in my life. Thank you for reading over chapters, listening to my ideas, and supporting my decisions.

i
Table of Contents
Acknowledgments ...........................................................................................................................i

Table of Contents ...........................................................................................................................ii List of Figures ................................................................................................................................iii List of Tables .................................................................................................................................vi Chapter 1: Introduction ..................................................................................................................1 Chapter 2: Methodology ...............................................................................................................14 Chapter 3: Results and Discussion of Collected Taxa ..................................................................25 Chapter 4: Interpretations and Conclusions ..................................................................................61 References Cited ...........................................................................................................................70 Appendix 1....................................................................................................................................78

ii
List of Figures
Figure 1.1 Map of BOP Wetlands in relation to other Maya sites. Photo from Beach et al. 2009..................................................................................................................................................2

Figure 1.2 SubOp A, Lots 1, 2, and 3 in progress.........................................................................10 Figure 1.3 Northwest baulk of SubOp B Lot 1, Closing...............................................................11 Figure 1.4 Southeast baulk of SubOp B Lot 3, Closing................................................................12 Figure 1.5 SubOp A profile sketch showing stratigraphic layers and location of seeds from AMS dating. Image provided by Samantha Krause................................................................................13

Figure 2.1 View of south baulk of SubOp A canal after column sampling..................................15

Figure 2.2 BOP Field Season 2016 Images: a) Flote-Tech Model A flotation machine at Programme for Belize, b) agitating the heavy fraction soil during the flotation process, c) heavy and light fractions drying after flotation, d) sorting the heavy fraction at Maya Research Program..........................................................................................................................................17

Figure 2.3 VWR Ultrasonic Bath Model 75T ..............................................................................20 Figure 2.4 BOP 10025 sieve fractions after ultrasonic bath.........................................................20 Figure 2.5 Archaeobotanical remains recovered from sorting after ultrasonic bath.....................21

Figure 3.1 Absolute counts of Eleocharis sp., Cladium jamaicense, and Spilanthes cf. acmella

and all other identified non-wood seeds in SubOp A Canal and Field..........................................28

Figure 3.2 Absolute counts of Eleocharis sp., Cladium jamaicense, and Spilanthes cf. acmella

and all other identified non-wood seeds in SubOp B Canal and Field..........................................29 iii
Figure 3.3 Example of an E. prostrata achene found in only two flotation samples...........................................................................................................................................32

Figure 3.4 Example of a Spilanthes cf. acmella achene from BOP..............................................33 Figure 3.5 Example of a Celtis sp. fruit from BOP.......................................................................34 Figure 3.6 Example of a Chenopodium ambrosioides seed from the canal of BOP.....................35 Figure 3.7 Example of a Cayaponia sp. seed from SubOp B.......................................................36 Figure 3.8 Example of a Carex polystachya seed from BOP.......................................................37 Figure 3.9 Example of a Cladium jamaicense seed from BOP....................................................38 Figure 3.10 Example of a Cyperus sp. seed from BOP................................................................39 Figure 3.11 Example of an Eleocharis sp. seed from BOP..........................................................40 Figure 3.12 Example of a Fimbristylis cf. dichotoma achene from BOP Canal...........................41 Figure 3.13 Example of a Scirpus sp. seed from BOP..................................................................42 Figure 3.14 Example of a Physalis sp. seed found in only one sample (BOP 10004) from SubOp B.....................................................................................................................................................43

Figure 3.15 Example of a Solanum sp. seed recovered from BOP...............................................44 Figure 3.16 Example of a Najas guadalupensis seed recovered from BOP.................................45 Figure 3.17 Example of a Juncus sp. seed found in two samples (BOP 10026 and BOP 10027) from SubOp A Canal.....................................................................................................................46

iv
Figure 3.18 Example of an Andropogon sp. seed found in only one sample (BOP 10004) from SubOp B Canal..............................................................................................................................47

Figure 3.19 Example of a Panicum sp. seed found in two samples (BOP 10015 and BOP 10017) from SubOp A Field.......................................................................................................................48

Figure 3.20 Example of a Paspalum sp. seed found in only one sample (BOP 10004) of SubOp B ....................................................................................................................................................49

Figure 3.21 Example of a Polygonum sp. seed recovered from BOP...........................................50 Figure 3.22 Example of a Portulaca sp. seed recovered from BOP.............................................51 Figure 3.23 Wood density from SubOp A flotation sample depths..............................................53 Figure 3.24 Wood density from SubOp B flotation sample depths..............................................53 Figure 3.25 Transverse (100x) and tangential (200x) section of Bucida buceras........................55 Figure 3.26 Transverse (100x) and tangential (120x) sections of H. campechianum..................56 Figure 3.27 Transverse (50x) section of Mimosa sp.....................................................................58 Figure 3.28 Transverse (100x) and tangential (100x) sections of Licaria sp...............................59 Figure 3.29 Transverse section of Ficus sp. at 100x magnification.............................................60 Figure 4.1 SubOp A: Relative frequency of seeds associated with wet, wet/dry, and dry ecological conditions.....................................................................................................................62

Figure 4.2 SubOp B: Relative frequency of seeds associated with wet, wet/dry, and dry ecological conditions.....................................................................................................................63

v
List of Tables
Table 1.1 List of 40 out of 44 identified herbarium specimens collected in 2016 qualitative field survey...............................................................................................................................................7

Table 3.1 Radiocarbon dates from BOP SubOp A canal and field...............................................26 Table 3.2 Summary of identified plant remains recovered from flotation samples during the 2016 field season at BOP, organized alphabetically by scientific name. C/F denote the context of recovery as Canal (C) or Field (F).................................................................................................27

Table 3.3 Analyzed hand-collected wood specimens with species, depth, SubOp, and context............................................................................................................................................54

Table 4.1 SubOp A species classified by ecological preferences: wet, wet/dry, or dry...............61 Table 4.2 SubOp B species classified by ecological preferences: wet, wet/dry, or dry................63

vi

Chapter 1: Introduction

The ancient Maya civilization spanned most of Mexico, Guatemala, Honduras, Belize, and El Salvador from 1800 BCE – 1530 CE (Coe and Houston 2015). The complex political and cultural organization of the Maya relied on a strong agriculturally based society adapted to a wide range of ecological and topographical settings. While their management was by no means uniform, they were able to adapt agricultural methods based on soil type, size of the land, location, etc. In order to sustain their growing population in the Late Classic Period (250 – 800 CE), the Maya used a variety of agricultural methods such as terracing, milpas, slash and burn, and small household garden plots (Lentz 1991; Beach and Dunning 1995). However, these forms of cultivation could not be used in all of the diverse environmental settings occupied by the ancient Maya, especially the swampy lowlands. Discoveries of rectangular canal patterns in wetland margins in the Maya lowlands of Guatemala, Belize, and Mexico shed light on a previously unknown agricultural practice: raised wetland fields (Turner and Harrison 1983; Jacob 1995; Fedick 1996; Pohl et. al 1996; Beach et al. 2009). These narrow, rectangular plots, elevated above the low-lying, seasonally inundated land bordering rivers or bajos, enabled the Maya to grow crops in these otherwise uncultivable areas (Coe and Houston 2015:20). The use of these wetlands allowed for the Maya to be resilient during droughts and other hardships and allowed for independence of smaller communities (Beach et al. 2009).

Throughout the Maya Lowlands, wetland agricultural use shows intensification contemporary with population increase of the ancient Maya, especially during the Classic period (Beach et al. 2009). There are three types of wetland fields: raised, drained, and recessional (Beach et al. 2009:1715). At Birds of Paradise (BOP) (Figure 1.1), features interpreted as raised fields and canals have been detected (Beach et al. 2009). Macrobotanical evidence provides one
1avenue for testing the hypothesis that these features at BOP were used as raised agricultural fields. Whereas past research has focused on pollen and stable carbon isotope evidence to understand these features, this thesis aims to use macrobotanical analysis to address questions that have long been asked about BOP. Prior to this study, only a limited preliminary macrobotanical study had been undertaken at BOP (Goldstein 2007). Macrobotanical analysis, in contrast to pollen analysis, provides a more local index of vegetation changes which can help to support or refute the hypothesis that these features were managed fields.

Figure 1.1 Map of BOP Wetlands in relation to other Maya sites. Photo from Beach et al. 2009:1711.

The goals of this research are to: 1) identify the plants that were growing in the canals, 2) identify the plants that were growing in the features identified as raised fields, 3) assess their ecological preferences, 4) assess changing frequencies of different types of plants over time, and
2
5) provide some insight into how the canals and fields were used. The reported findings are based on 28 flotation samples and 14 hand-collected charcoal specimens. This research is significant because it marks the first time any standardized macrobotanical analysis has been done at BOP, and therefore makes an important contribution to understanding Maya interaction with the fields and canals and changes in land use over time. Moreover, the lab protocols adopted here provide a model for future studies where small seeds dominate the assemblage.

Thesis Organization

The first chapter will give an overview of the site Birds of Paradise, provide background information on the environmental settings at BOP and discuss past excavations conducted at BOP. Additionally, it also includes information on archaeobotanical analysis in Mesoamerica and discusses the BOP 2016 excavation. The following chapter, Chapter 2, outlines the research methodologies that were used in the recovery, identification, and quantification of archaeobotanical remains from BOP. Chapter 3 presents the macrobotanical results and findings. Chapter 4 discusses the implications of this research for BOP in particular and the outcomes of methodological practices implemented in this thesis and potential avenues for further research. Appendix 1 includes a complete inventory of the plant remains.

Birds of Paradise Overview

BOP is a wetland field site located between three to five km of the large Maya center at
Blue Creek and Gran Cacao in northwest Belize. It occupies at least 1km2 of the savanna floodplain and more in the surrounding forest (Beach et al. 2015b:1622). The features identified as Maya wetland fields at this site are rectilinear areas with one to two m wide canals running east to west and north to south at interval of approximately ten m (Beach et al. 2015b:1622).

3
Timothy Beach and colleagues have been excavating at BOP since 2005 trying to understand the significance of these anthropogenic fields. On the basis of radiocarbon dates from the BOP

wetland fields and canals, the site’s occupation is dated from the Classic (250 – 900 CE) to

Postclassic (900 -1200 CE) periods (Beach et al. 2009; Luzzadder-Beach et al. 2012). Dr. Beach and Dr. Sheryl Luzzadder-Beach propose that the use of raised fields was one way that people adapted to environmental change, specifically as a response to rising sea level and the attendant higher water table and lower water quality (Beach et al. 2009; Luzzadder-Beach et al. 2012; Beach et al. 2015; Coe and Houston 2015). Understanding the archaeology behind wetland fields will add key insights to interpreting how humans have had to adapt to changing environments throughout time (Beach et al. 2009).

Environmental Setting of BOP

Geography

BOP is situated in northwestern Belize in an active floodplain near the confluence of

three rivers (Rio Azul, Rio Bravo, and Booth’s River) within the Rio Bravo Conservation and

Management Area in Northwestern Belize (Beach et al. 2009). Near the borders with Mexico and Guatemala, all three of these countries include well-drained upland karst ridge, bajos, river valleys, and the coastal plain (Beach et al. 2013). Topography in the Rio Bravo includes level and gently rolling areas, hills, and escarpments with deep ravines (Brokaw and Mallory 1993).

The local bedrock of this region is mostly limestone marl and clay, the surface of which is deeply weathered into a calcareous material known locally as sascab (Darch 1981). The floodplain and depressions are filled with Quaternary calcareous clays, marls, and peats, the upper portions of which contain abundant secondary carbonate and especially gypsum (Pohl et

4al. 1990). In the canals and fields at BOP, the soil is composed of clay and moderate quantities of gypsum, carbonate, and organic matter (Beach et al. 2009). Gypsum in the soil is derived from a geological unit that the groundwater encountered in the Rio Bravo fault zone as it rose to this elevation (Luzzadder-Beach et al. 2012:3649).

Climate

BOP has a tropical wet and dry climate with an average annual temperature is 26.4°C
(79.52°F) and receives about 1500 mm (60 in) of precipitation annually, mostly during the rainy season between June and December (Beach et al. 2009; Beach et al. 2015b:1613). The highest rainfall occurs in June and September (Beach et al. 2009). Rio Bravo lies between 17° and 18° N latitude, in the outer- or sub-tropics. There is minor seasonal variation in temperature and strong seasonality of precipitation, which puts Rio Bravo in the subtropical moist life zone of the Holdridge Life Zone System (Holdridge and Poveda 1975; Brokaw and Mallory 1993:5). Belize experiences a lot of humidity, due in part to the influence of the trade winds that collect moisture over the Caribbean Sea (Bridgewater 2012). The fluctuations in rain and temperature encourage the growth of vegetation in the general category of subtropical moist forests, with broadleaf, semi-deciduous forests (Brokaw and Mallory 1993). Because climate has not changed significantly for the past 3,000 years within Mesoamerica, modern conditions reflect a climate that is similar, with some fluctuations, to that during the occupation of the ancient Maya (Markgraf 1989).

Vegetation

During the 2016 field season, Lentz, Wendel, and Slotten conducted a qualitative field survey of several habitats surrounding the BOP excavations, both to better understand the local

5vegetation and to collect comparative materials for use in identifying materials recovered from flotation samples (Wendel et al. 2017). Two specimens of each plant were collected, with one given to the Belize herbarium in Belmopan and the other exported to the University of Cincinnati. Plants were collected from the following zones: savanna, scrub forest or floodplain forest, riparian forest, and the wetland vegetations adjacent to Cacao Creek. When a plant was in flower or had fruits it was collected, identified, dried, and then pressed for mounting on herbarium sheets. Although collection took place during the dry season, few plants were in flower at the time. During survey, we recorded the coordinates, frequency, and a list of features (habitat, flower color, bark texture, etc.) for each specimen collected. A total of 44 specimens were collected in the field and 40 were identified in the lab (Table 1.1). Five species (Bucida

buceras, Cyperus sp., Ficus sp., Panicum sp., Solanum sp.) collected during our field survey

were also found in the flotation samples at BOP.
6
Table 1.1 List of 40 out 44 identified herbarium specimens collected in 2016 qualitative field survey.

Collection number

Recommended publications
  • Native Habitat Restoration in Eastern Washington Wine Vineyards

    Native Habitat Restoration in Eastern Washington Wine Vineyards

    NATIVE HABITAT RESTORATION IN EASTERN WASHINGTON WINE VINEYARDS AS A PEST MANAGEMENT STRATEGY By KATHARINE DENISE BUCKLEY A dissertation submitted in partial fulfillment of the requirements for the degree of DOCTOR OF PHILOSOPHY WASHINGTON STATE UNIVERSITY Department of Entomology MAY 2019 © Copyright by KATHARINE DENISE BUCKLEY, 2019 All Rights Reserved © Copyright by KATHARINE DENISE BUCKLEY, 2019 All Rights Reserved To the Faculty of Washington State University: The members of the Committee appointed to examine the dissertation of KATHARINE DENISE BUCKLEY find it satisfactory and recommend that it be accepted. _______________________________ David James, Ph.D., Chair _______________________________ Elizabeth Beers, Ph.D. _______________________________ Joan Davenport, Ph.D. ii ACKNOWLEDGMENTS I thank Lorraine Seymour and Gerry Lauby for their expertise, their organizational skills, their excellence as sounding boards, and hopefully their ability to pass some of that on to me. I thank Cole Provence and my family who were always supportive. I thank all the people who helped me with my statistics, especially Bernardo Chaves. I thank the computer technician who saved my computer’s data and my life. I thank Michael Aquilino. He knows what he did. Finally, I’d like to thank everyone who served on my committee, as well as Laura Lavine, for their guidance along the way. iii NATIVE HABITAT RESTORATION IN EASTERN WASHINGTON WINE VINEYARDS AS A PEST MANAGEMENT STRATEGY Abstract by Katharine Denise Buckley, Ph.D. Washington State University May 2019 Chair: David James Perennial crop systems such as wine grapes have begun using cover crops and hedgerows to increase beneficial insects and promote sustainable vineyard management in areas such as New Zealand and California.
  • Chromosome Numbers in Compositae, XII: Heliantheae

    Chromosome Numbers in Compositae, XII: Heliantheae

    SMITHSONIAN CONTRIBUTIONS TO BOTANY 0 NCTMBER 52 Chromosome Numbers in Compositae, XII: Heliantheae Harold Robinson, A. Michael Powell, Robert M. King, andJames F. Weedin SMITHSONIAN INSTITUTION PRESS City of Washington 1981 ABSTRACT Robinson, Harold, A. Michael Powell, Robert M. King, and James F. Weedin. Chromosome Numbers in Compositae, XII: Heliantheae. Smithsonian Contri- butions to Botany, number 52, 28 pages, 3 tables, 1981.-Chromosome reports are provided for 145 populations, including first reports for 33 species and three genera, Garcilassa, Riencourtia, and Helianthopsis. Chromosome numbers are arranged according to Robinson’s recently broadened concept of the Heliantheae, with citations for 212 of the ca. 265 genera and 32 of the 35 subtribes. Diverse elements, including the Ambrosieae, typical Heliantheae, most Helenieae, the Tegeteae, and genera such as Arnica from the Senecioneae, are seen to share a specialized cytological history involving polyploid ancestry. The authors disagree with one another regarding the point at which such polyploidy occurred and on whether subtribes lacking higher numbers, such as the Galinsoginae, share the polyploid ancestry. Numerous examples of aneuploid decrease, secondary polyploidy, and some secondary aneuploid decreases are cited. The Marshalliinae are considered remote from other subtribes and close to the Inuleae. Evidence from related tribes favors an ultimate base of X = 10 for the Heliantheae and at least the subfamily As teroideae. OFFICIALPUBLICATION DATE is handstamped in a limited number of initial copies and is recorded in the Institution’s annual report, Smithsonian Year. SERIESCOVER DESIGN: Leaf clearing from the katsura tree Cercidiphyllumjaponicum Siebold and Zuccarini. Library of Congress Cataloging in Publication Data Main entry under title: Chromosome numbers in Compositae, XII.
  • Literature Cited

    Literature Cited

    Literature Cited Robert W. Kiger, Editor This is a consolidated list of all works cited in volumes 19, 20, and 21, whether as selected references, in text, or in nomenclatural contexts. In citations of articles, both here and in the taxonomic treatments, and also in nomenclatural citations, the titles of serials are rendered in the forms recommended in G. D. R. Bridson and E. R. Smith (1991). When those forms are abbre- viated, as most are, cross references to the corresponding full serial titles are interpolated here alphabetically by abbreviated form. In nomenclatural citations (only), book titles are rendered in the abbreviated forms recommended in F. A. Stafleu and R. S. Cowan (1976–1988) and F. A. Stafleu and E. A. Mennega (1992+). Here, those abbreviated forms are indicated parenthetically following the full citations of the corresponding works, and cross references to the full citations are interpolated in the list alphabetically by abbreviated form. Two or more works published in the same year by the same author or group of coauthors will be distinguished uniquely and consistently throughout all volumes of Flora of North America by lower-case letters (b, c, d, ...) suffixed to the date for the second and subsequent works in the set. The suffixes are assigned in order of editorial encounter and do not reflect chronological sequence of publication. The first work by any particular author or group from any given year carries the implicit date suffix “a”; thus, the sequence of explicit suffixes begins with “b”. Works missing from any suffixed sequence here are ones cited elsewhere in the Flora that are not pertinent in these volumes.
  • El Género Sclerocarpus (Asteraceae, Heliantheae) En México

    El Género Sclerocarpus (Asteraceae, Heliantheae) En México

    Revista Mexicana de Biodiversidad 82: 51-61, 2011 El género Sclerocarpus (Asteraceae, Heliantheae) en México The genus Sclerocarpus (Asteraceae, Heliantheae) in Mexico José Luis Villaseñor* y Óscar Hinojosa-Espinosa Departamento de Botánica, Instituto de Biología, Universidad Nacional Autónoma de México., Apartado postal 70-367, 04510 México, D. F., México. *Correspondencia: [email protected] Resumen. El género Sclerocarpus (Asteraceae, Heliantheae) está constituido por 8 especies, 7 de ellas presentes en el territorio mexicano. Este taxón se caracteriza por sus páleas, que al madurar se tornan gruesas, duras y encierran por completo a las cipselas formando estructuras llamadas esclerocarpos. Se presenta una sinopsis del género para México, una clave para la identificación de las especies y mapas de distribución de cada una de ellas. Palabras clave: Asteraceae, esclerocarpo, Heliantheae, México, Sclerocarpus. Abstract.The genus Sclerocarpus (Asteraceae, Heliantheae) comprises 8 pecies, 7 of them recorded in Mexico. It is characterized by its paleae, that become thick and hard when mature, and completely enclosing the cypselae, forming structures called sclerocarps. A synopsis of the genus in the country is provided, including distribution maps of the species and a key to their identification. Key words: Asteraceae, Heliantheae, Mexico, sclerocarp, Sclerocarpus. Introducción y Rhysolepis S.F. Blake (actualmente incluido en Viguiera Kunth). En estos taxones, las cipselas maduras también se encuentran envueltas por completo por sus páleas; sin El género Sclerocarpus Jacq. es un miembro de Helian- embargo, la envoltura paleácea es relativamente delgada, theae, la tribu más grande y diversa de la familia Asteraceae papirácea y de superficie rugosa o corrugada. En los 3 (Stuessy, 1977).
  • New Smut Fungi (Ustilaginomycetes) from Mexico, and the Genus Lundquistia

    New Smut Fungi (Ustilaginomycetes) from Mexico, and the Genus Lundquistia

    Fungal Diversity New smut fungi (Ustilaginomycetes) from Mexico, and the genus Lundquistia Kálmán Vánky* Herbarium Ustilaginales Vánky (HUV), Gabriel-Biel-Str. 5, D-72076 Tübingen, Germany Vánky, K. (2004). New smut fungi (Ustilaginomycetes) from Mexico, and the genus Lundquistia. Fungal Diversity 17: 159-190. The genus Lundquistia is emended and widened. Twelve new species of smut fungi are described from Mexico: Lundquistia mexicana on Andropogon gerardii and Schizachyrium mexicanum, Entyloma aldamae on Aldama dentata, E. siegesbeckiae on Siegesbeckia orientalis, Jamesdicksonia festucae on Festuca tolucensis, Macalpinomyces tuberculatus on Bouteloua curtipendula, Sporisorium dacryoideum on Aristida adscensionis, S. ustilaginiforme on Muhlenbergia pulcherrima, Tilletia brefeldii on Muhlenbergia filiculmis, T. gigacellularis on Bouteloua filiformis, T. microtuberculata on Muhlenbergia pulcherrima, Ustilago circumdata on Muhlenbergia montana, and U. panici-virgati on Panicum virgatum. New combinations proposed: Lundquistia dieteliana, L. duranii and L. panici-leucophaei, with its three new synonyms, Ustilago bonariensis, Sorosporium lindmanii and L. fascicularis. Key words: Lundquistia, new combinations, new species, synonyms, taxonomy. Introduction The smut fungi of Mexico are relatively well-known, especially due to over 30 years of investigation by Prof. Ruben Durán (Washington State University, Pullman, USA). Numerous papers have been published by Durán, alone or in collaboration, in which many new species, a new genus and also the nuclear behaviour of the basidia and basidiospores of many North American smut fungi have been described (Durán and Fischer, 1961; Durán and Safeeulla, 1965, 1968; Durán, 1968, 1969, 1970, 1971, 1972, 1979, 1980, 1982, 1983; Durán and Cromarty, 1974, 1977; Cordas and Durán, 1976[1977]). Durán's work culminated in the publication of the profusely illustrated book Ustilaginales of Mexico (1987), containing 128 taxa of which 14 were new species.
  • Washington Flora Checklist a Checklist of the Vascular Plants of Washington State Hosted by the University of Washington Herbarium

    Washington Flora Checklist a Checklist of the Vascular Plants of Washington State Hosted by the University of Washington Herbarium

    Washington Flora Checklist A checklist of the Vascular Plants of Washington State Hosted by the University of Washington Herbarium The Washington Flora Checklist aims to be a complete list of the native and naturalized vascular plants of Washington State, with current classifications, nomenclature and synonymy. The checklist currently contains 3,929 terminal taxa (species, subspecies, and varieties). Taxa included in the checklist: * Native taxa whether extant, extirpated, or extinct. * Exotic taxa that are naturalized, escaped from cultivation, or persisting wild. * Waifs (e.g., ballast plants, escaped crop plants) and other scarcely collected exotics. * Interspecific hybrids that are frequent or self-maintaining. * Some unnamed taxa in the process of being described. Family classifications follow APG IV for angiosperms, PPG I (J. Syst. Evol. 54:563?603. 2016.) for pteridophytes, and Christenhusz et al. (Phytotaxa 19:55?70. 2011.) for gymnosperms, with a few exceptions. Nomenclature and synonymy at the rank of genus and below follows the 2nd Edition of the Flora of the Pacific Northwest except where superceded by new information. Accepted names are indicated with blue font; synonyms with black font. Native species and infraspecies are marked with boldface font. Please note: This is a working checklist, continuously updated. Use it at your discretion. Created from the Washington Flora Checklist Database on September 17th, 2018 at 9:47pm PST. Available online at http://biology.burke.washington.edu/waflora/checklist.php Comments and questions should be addressed to the checklist administrators: David Giblin ([email protected]) Peter Zika ([email protected]) Suggested citation: Weinmann, F., P.F. Zika, D.E. Giblin, B.
  • Eastern Washington Plant List

    Eastern Washington Plant List

    The NatureMapping Program Revised: 9/15/2011 Eastern Washington Plant List - Scientific Name 1- Non- native, 2- ID Scientific Name Common Name Plant Family Invasive √ 1141 Abies amabilis Pacific silver fir Pinaceae 1 Abies grandis Grand fir Pinaceae 1142 Abies lasiocarpa Sub-alpine fir Pinaceae 762 Abronia mellifera White sand verbena Nyctaginaceae 1143 Abronia umbellata Pink sandverbena Nyctaginaceae 763 Acer glabrum Douglas maple Aceraceae 3 Acer macrophyllum Big-leaf maple Aceraceae 470 Acer platinoides* Norway maple Aceraceae 1 5 Achillea millifolium Yarrow Asteraceae 1144 Aconitum columbianum Monkshood Ranunculaceae 8 Actaea rubra Baneberry Ranunculaceae 9 Adenocaulon bicolor Pathfinder Asteraceae 10 Adiantum pedatum Maidenhair fern Polypodiaceae 764 Agastache urticifolia Nettle-leaf horse-mint Lamiaceae 1145 Agoseris aurantiaca Orange agoseris Asteraceae 1146 Agoseris elata Tall agoseris Asteraceae 705 Agoseris glauca Mountain agoseris Asteraceae 608 Agoseris grandiflora Large-flowered agoseris Asteraceae 716 Agoseris heterophylla Annual agoseris Asteraceae 11 Agropyron caninum Bearded wheatgrass Poaceae 560 Agropyron cristatum* Crested wheatgrass Poaceae 1 1147 Agropyron dasytachyum Thickspike wheatgrass Poaceae 739 Agropyron intermedium* Intermediate ryegrass Poaceae 1 12 Agropyron repens* Quack grass Poaceae 1 744 Agropyron smithii Bluestem Poaceae 523 Agropyron spicatum Blue-bunch wheatgrass Poaceae 687 Agropyron trachycaulum Slender wheatgrass Poaceae 13 Agrostis alba* Red top Poaceae 1 799 Agrostis exarata* Spike bentgrass
  • Appendix 6.8-A

    Appendix 6.8-A

    Appendix 6.8-A Terrestrial Wildlife and Vegetation Baseline Report AJAX PROJECT Environmental Assessment Certificate Application / Environmental Impact Statement for a Comprehensive Study Ajax Mine Terrestrial Wildlife and Vegetation Baseline Report Prepared for KGHM Ajax Mining Inc. Prepared by This image cannot currently be displayed. Keystone Wildlife Research Ltd. #112, 9547 152 St. Surrey, BC V3R 5Y5 July 2015 Ajax Mine Terrestrial Wildlife and Vegetation Baseline Keystone Wildlife Research Ltd. DISCLAIMER This report was prepared exclusively for KGHM Ajax Mining Inc. by Keystone Wildlife Research Ltd. The quality of information, conclusions and estimates contained herein is consistent with the level of effort expended and is based on: i) information available at the time of preparation; ii) data collected by Keystone Wildlife Research Ltd. and/or supplied by outside sources; and iii) the assumptions, conditions and qualifications set forth in this report. This report is intended for use by KGHM Ajax Mining Inc. only, subject to the terms and conditions of its contract with Keystone Wildlife Research Ltd. Any other use, or reliance on this report by any third party, is at that party’s sole risk. 2 Ajax Mine Terrestrial Wildlife and Vegetation Baseline Keystone Wildlife Research Ltd. EXECUTIVE SUMMARY Baseline wildlife and habitat surveys were initiated in 2007 to support a future impact assessment for the redevelopment of two existing, but currently inactive, open pit mines southwest of Kamloops. Detailed Project plans were not available at that time, so the general areas of activity were buffered to define a study area. The two general Project areas at the time were New Afton, an open pit just south of Highway 1, and Ajax, east of Jacko Lake.
  • ICBEMP Analysis of Vascular Plants

    ICBEMP Analysis of Vascular Plants

    APPENDIX 1 Range Maps for Species of Concern APPENDIX 2 List of Species Conservation Reports APPENDIX 3 Rare Species Habitat Group Analysis APPENDIX 4 Rare Plant Communities APPENDIX 5 Plants of Cultural Importance APPENDIX 6 Research, Development, and Applications Database APPENDIX 7 Checklist of the Vascular Flora of the Interior Columbia River Basin 122 APPENDIX 1 Range Maps for Species of Conservation Concern These range maps were compiled from data from State Heritage Programs in Oregon, Washington, Idaho, Montana, Wyoming, Utah, and Nevada. This information represents what was known at the end of the 1994 field season. These maps may not represent the most recent information on distribution and range for these taxa but it does illustrate geographic distribution across the assessment area. For many of these species, this is the first time information has been compiled on this scale. For the continued viability of many of these taxa, it is imperative that we begin to manage for them across their range and across administrative boundaries. Of the 173 taxa analyzed, there are maps for 153 taxa. For those taxa that were not tracked by heritage programs, we were not able to generate range maps. (Antmnnrin aromatica) ( ,a-’(,. .e-~pi~] i----j \ T--- d-,/‘-- L-J?.,: . ey SAP?E%. %!?:,KnC,$ESS -,,-a-c--- --y-- I -&zII~ County Boundaries w1. ~~~~ State Boundaries <ii&-----\ \m;qw,er Columbia River Basin .---__ ,$ 4 i- +--pa ‘,,, ;[- ;-J-k, Assessment Area 1 /./ .*#a , --% C-p ,, , Suecies Locations ‘V 7 ‘\ I, !. / :L __---_- r--j -.---.- Columbia River Basin s-5: ts I, ,e: I’ 7 j ;\ ‘-3 “.
  • Flora and Vegetation of the Huascarán National Park, Ancash, Peru: With

    Flora and Vegetation of the Huascarán National Park, Ancash, Peru: With

    Iowa State University Capstones, Theses and Retrospective Theses and Dissertations Dissertations 1988 Flora and vegetation of the Huascarán National Park, Ancash, Peru: with preliminary taxonomic studies for a manual of the flora David Nelson Smith Iowa State University Follow this and additional works at: https://lib.dr.iastate.edu/rtd Part of the Botany Commons Recommended Citation Smith, David Nelson, "Flora and vegetation of the Huascarán National Park, Ancash, Peru: with preliminary taxonomic studies for a manual of the flora " (1988). Retrospective Theses and Dissertations. 8891. https://lib.dr.iastate.edu/rtd/8891 This Dissertation is brought to you for free and open access by the Iowa State University Capstones, Theses and Dissertations at Iowa State University Digital Repository. It has been accepted for inclusion in Retrospective Theses and Dissertations by an authorized administrator of Iowa State University Digital Repository. For more information, please contact [email protected]. INFORMATION TO USERS The most advanced technology has been used to photo­ graph and reproduce this manuscript from the microfilm master. UMI films the text directly from the original or copy submitted. Thus, some thesis and dissertation copies are in typewriter face, while others may be from any type of computer printer. The quality of this reproduction is dependent upon the quality of the copy submitted. Broken or indistinct print, colored or poor quality illustrations and photographs, print bleedthrough, substandard margins, and improper alignment can adversely affect reproduction. In the unlikely event that the author did not send UMI a complete manuscript and there are missing pages, these will be noted. Also, if unauthorized copyright material had to be removed, a note will indicate the deletion.
  • Notes on Certain Type Specimens of American Asteraceae in European Herbaria

    Notes on Certain Type Specimens of American Asteraceae in European Herbaria

    NOTES ON CERTAIN TYPE SPECIMENS OF AMERICAN ASTERACEAE IN EUROPEAN HERBARIA By S. F. BLAKE In the course of the writer's work at several of the larger European herbaria in the summer of 1925, special effort was made to determine the identity of various American species of Asteraceae which were imperfectly described by early authors and have since been unrecog- nized or misinterpreted. In this paper four generic names (Parastre- phia, PhUactisj Anaitis, and Aschenbornia) and about 115 specific names which have been misunderstood or regarded as doubtful since the time of their original publication are first given their rightful status as valid names or synonyms. A few identifications here in- cluded have already been published.1 The genera are arranged in the systematic order of the accepted names, the species alphabetically under the genera. Special attention was given to the investigation of the Heliantheae, particularly in the Prodromus Herbarium of the DeCandolles, now forming a unit in the Delessert Herbarium, Geneva, and in the Schultz Bipontinus Herbarium, which includes Sprengel's types and forms a part of the Cosson Herbarium at the Museum d'Histoire Naturelle at Paris. Schultz's herbarium includes, besides his own types and those of Sprengel, a wealth of fragments of types and authentic specimens from other authors, and the Prodromus Herba- rium is rich in authentic specimens of earlier writers in addition to De Candolle's own types. A rather hurried examination was made of all the Asteraceae of the Humboldt and Bonpland Herbarium at Paris, Notes, photographs, and in many cases small fragments of types or significant specimens were obtained of hundreds of species of American Asteraceae, of which those discussed in this paper represent only the comparatively small part in which some change of interpretation has been found necessary.
  • University Micrcxilms International 300 N

    University Micrcxilms International 300 N

    INFORMATION TO USERS This reproduction was made from a copy of a document sent to us for microfilming. While the most advanced technology has been used to photograph and reproduce this document, the quality of the reproduction is heavily dependent upon the quality of the material submitted. The following explanation of techniques is provided to help clarify markings or notations which may appear on this reproduction. 1.The sign or “target” for pages apparently lacking from the document photographed is “Missing Page(s)”. If it was possible to obtain the missing page(s) or section, they are spliced into the film along with adjacent pages. This may have necessitated cutting through an image and duplicating adjacent pages to assure complete continuity. 2. When an image on the film is obliterated with a round black mark, it is an indication o f either blurred copy because o f movement during exposure, duplicate copy, or copyrighted materials that should not have been filmed. For blurred pages, a good image o f the page can be found in the adjacent frame. If copyrighted materials were deleted, a target note will appear listing the pages in the adjacent frame. 3. When a map, drawing or chart, etc., is part of the material being photographed, a definite method of “sectioning” the material has been followed. It is customary to begin filming at the upper left hand comer of a large sheet and to continue from left to right in equal sections with small overlaps. If necessary, sectioning is continued again—beginning below the first row and continuing on until complete.