The Ecology and Evolution of Melitaeine Butterflies Niklas Wahlberg Metapopulation Research Group Department of Ecology and Systematics Division of Population Biology University of Helsinki Finland Academic dissertation To be presented, with permission of the Faculty of Science of the University of Helsinki, for public criticism in the lecture room of the Department of Ecology and Systematics, P. Rautatiekatu 13, on October 27, 2000, at 12 o’clock noon. Helsinki 2000 © Niklas Wahlberg, pp. 7–26 Technical editing by Johan Ulfvens Author’s address: Metapopulation Research Group Department of Ecology and Systematics Division of Population Biology P.O. Box 17 (Arkadiankatu 7) 00014 University of Helsinki Finland e-mail:
[email protected] ISBN 952-91-2615-8 (nid) ISBN 952-91-2688-3 (pdf) Oy Edita Ab Helsinki 2000 Helsinki 2000 The Ecology and Evolution of Melitaeine Butterflies Niklas Wahlberg Metapopulation Research Group Department of Ecology and Systematics Division of Population Biology P.O. Box 17 (Arkadiankatu 7) 00014 University of Helsinki Finland The thesis is based on the following articles: I Wahlberg, N. & Zimmermann, M. 2000. Pattern of phylogenetic relationships among members of the tribe Melitaeini (Lepidoptera: Nymphalidae) inferred from mtDNA sequences. – Cladistics 16, in press. II Wahlberg, N. 2000. The phylogenetics and biochemistry of host plant specialization in melitaeine butterflies (Lepidoptera: Nymphalidae). – Submitted manuscript. III Wahlberg, N., Klemetti, T., Selonen, V. & Hanski, I. 2000. Metapopulation structure and movements in five species of checkerspot butterflies. – Manuscript. IV Wahlberg, N., Moilanen, A. & Hanski, I. 1996. Predicting the occurrence of endangered species in fragmented landscapes. – Science 273: 1536-1538. V Wahlberg, N., Klemetti, T.