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												Phylogenetic Relationships and Historical Biogeography of Tribes and Genera in the Subfamily Nymphalinae (Lepidoptera: Nymphalidae)
Blackwell Science, LtdOxford, UKBIJBiological Journal of the Linnean Society 0024-4066The Linnean Society of London, 2005? 2005 862 227251 Original Article PHYLOGENY OF NYMPHALINAE N. WAHLBERG ET AL Biological Journal of the Linnean Society, 2005, 86, 227–251. With 5 figures . Phylogenetic relationships and historical biogeography of tribes and genera in the subfamily Nymphalinae (Lepidoptera: Nymphalidae) NIKLAS WAHLBERG1*, ANDREW V. Z. BROWER2 and SÖREN NYLIN1 1Department of Zoology, Stockholm University, S-106 91 Stockholm, Sweden 2Department of Zoology, Oregon State University, Corvallis, Oregon 97331–2907, USA Received 10 January 2004; accepted for publication 12 November 2004 We infer for the first time the phylogenetic relationships of genera and tribes in the ecologically and evolutionarily well-studied subfamily Nymphalinae using DNA sequence data from three genes: 1450 bp of cytochrome oxidase subunit I (COI) (in the mitochondrial genome), 1077 bp of elongation factor 1-alpha (EF1-a) and 400–403 bp of wing- less (both in the nuclear genome). We explore the influence of each gene region on the support given to each node of the most parsimonious tree derived from a combined analysis of all three genes using Partitioned Bremer Support. We also explore the influence of assuming equal weights for all characters in the combined analysis by investigating the stability of clades to different transition/transversion weighting schemes. We find many strongly supported and stable clades in the Nymphalinae. We are also able to identify ‘rogue’ - 
												
												List of Animal Species with Ranks October 2017
Washington Natural Heritage Program List of Animal Species with Ranks October 2017 The following list of animals known from Washington is complete for resident and transient vertebrates and several groups of invertebrates, including odonates, branchipods, tiger beetles, butterflies, gastropods, freshwater bivalves and bumble bees. Some species from other groups are included, especially where there are conservation concerns. Among these are the Palouse giant earthworm, a few moths and some of our mayflies and grasshoppers. Currently 857 vertebrate and 1,100 invertebrate taxa are included. Conservation status, in the form of range-wide, national and state ranks are assigned to each taxon. Information on species range and distribution, number of individuals, population trends and threats is collected into a ranking form, analyzed, and used to assign ranks. Ranks are updated periodically, as new information is collected. We welcome new information for any species on our list. Common Name Scientific Name Class Global Rank State Rank State Status Federal Status Northwestern Salamander Ambystoma gracile Amphibia G5 S5 Long-toed Salamander Ambystoma macrodactylum Amphibia G5 S5 Tiger Salamander Ambystoma tigrinum Amphibia G5 S3 Ensatina Ensatina eschscholtzii Amphibia G5 S5 Dunn's Salamander Plethodon dunni Amphibia G4 S3 C Larch Mountain Salamander Plethodon larselli Amphibia G3 S3 S Van Dyke's Salamander Plethodon vandykei Amphibia G3 S3 C Western Red-backed Salamander Plethodon vehiculum Amphibia G5 S5 Rough-skinned Newt Taricha granulosa - 
												
												Dieteria [Machaeranthera] Cansecens
HOARY TANSYASTER Dieteria (Machaeranthera) canescens (Pursh) Nutt. Asteraceae – Aster family Corey L. Gucker & Nancy L. Shaw | 2018 ORGANIZATION NOMENCLATURE Dieteria canescens (Pursh) Nutt. until recently Names, subtaxa, chromosome number(s), hybridization. (2010) was known as Machaeranthera canescens (Pursh) A. Gray (ITIS 2017; USDA NRCS 2017). This species belongs to the Astereae tribe of the Asteraceae family (Morgan Range, habitat, plant associations, elevation, soils. 2006) and will hereafter be referred to by its common name, hoary tansyaster. NRCS Plant Code. MACA2 (USDA NRCS 2017). Life form, morphology, distinguishing characteristics, reproduction. Subtaxa. The Flora of North America (Morgan 2006) recognizes ten varieties of hoary tansyaster: Dieteria canescens var. Growth rate, successional status, disturbance ecology, importance to canescens, ambigua, aristata, glabra, incana, animals/people. leucanthemifolia, nebraskana, sessiliflora, shastensis, and ziegleri. Current or potential uses in restoration. Synonyms (Morgan 2006). Dieteria canescens: Machaeranthera canescens (Pursh) A. Gray; Aster canescens Seed sourcing, wildland seed collection, seed cleaning, storage, Pursh testing and marketing standards. D. c. var. ambigua: M. canescens (Pursh) A. Gray var. ambigua B.L. Turner D. c. var. aristatus: A. canescens Pursh var. Recommendations/guidelines for producing seed. aristatus Eastwood; M. canescens (Pursh) A. Gray var. aristata (Eastwood) B.L. Turner; M. rigida Greene D. c. var. canescens: M. divaricata (Nuttall) Recommendations/guidelines for producing planting stock. Greene; M. laetevirens Greene; M. latifolia A. Nelson; M. pulverulenta (Nuttall) Greene; M. viscosa (Nuttall) Greene D. c. var. glabra: M. canescens (Pursh) A. Gray Recommendations/guidelines, wildland restoration successes/ var. glabra A. Gray; A. canescens Pursh var. failures. viridis A. Gray; M. linearis Greene D. - 
												
												MOTHS and BUTTERFLIES LEPIDOPTERA DISTRIBUTION DATA SOURCES (LEPIDOPTERA) * Detailed Distributional Information Has Been J.D
MOTHS AND BUTTERFLIES LEPIDOPTERA DISTRIBUTION DATA SOURCES (LEPIDOPTERA) * Detailed distributional information has been J.D. Lafontaine published for only a few groups of Lepidoptera in western Biological Resources Program, Agriculture and Agri-food Canada. Scott (1986) gives good distribution maps for Canada butterflies in North America but these are generalized shade Central Experimental Farm Ottawa, Ontario K1A 0C6 maps that give no detail within the Montane Cordillera Ecozone. A series of memoirs on the Inchworms (family and Geometridae) of Canada by McGuffin (1967, 1972, 1977, 1981, 1987) and Bolte (1990) cover about 3/4 of the Canadian J.T. Troubridge fauna and include dot maps for most species. A long term project on the “Forest Lepidoptera of Canada” resulted in a Pacific Agri-Food Research Centre (Agassiz) four volume series on Lepidoptera that feed on trees in Agriculture and Agri-Food Canada Canada and these also give dot maps for most species Box 1000, Agassiz, B.C. V0M 1A0 (McGugan, 1958; Prentice, 1962, 1963, 1965). Dot maps for three groups of Cutworm Moths (Family Noctuidae): the subfamily Plusiinae (Lafontaine and Poole, 1991), the subfamilies Cuculliinae and Psaphidinae (Poole, 1995), and ABSTRACT the tribe Noctuini (subfamily Noctuinae) (Lafontaine, 1998) have also been published. Most fascicles in The Moths of The Montane Cordillera Ecozone of British Columbia America North of Mexico series (e.g. Ferguson, 1971-72, and southwestern Alberta supports a diverse fauna with over 1978; Franclemont, 1973; Hodges, 1971, 1986; Lafontaine, 2,000 species of butterflies and moths (Order Lepidoptera) 1987; Munroe, 1972-74, 1976; Neunzig, 1986, 1990, 1997) recorded to date. - 
												
												CA Checklist of Butterflies of Tulare County
Checklist of Buerflies of Tulare County hp://www.natureali.org/Tularebuerflychecklist.htm Tulare County Buerfly Checklist Compiled by Ken Davenport & designed by Alison Sheehey Swallowtails (Family Papilionidae) Parnassians (Subfamily Parnassiinae) A series of simple checklists Clodius Parnassian Parnassius clodius for use in the field Sierra Nevada Parnassian Parnassius behrii Kern Amphibian Checklist Kern Bird Checklist Swallowtails (Subfamily Papilioninae) Kern Butterfly Checklist Pipevine Swallowtail Battus philenor Tulare Butterfly Checklist Black Swallowtail Papilio polyxenes Kern Dragonfly Checklist Checklist of Exotic Animals Anise Swallowtail Papilio zelicaon (incl. nitra) introduced to Kern County Indra Swallowtail Papilio indra Kern Fish Checklist Giant Swallowtail Papilio cresphontes Kern Mammal Checklist Kern Reptile Checklist Western Tiger Swallowtail Papilio rutulus Checklist of Sensitive Species Two-tailed Swallowtail Papilio multicaudata found in Kern County Pale Swallowtail Papilio eurymedon Whites and Sulphurs (Family Pieridae) Wildflowers Whites (Subfamily Pierinae) Hodgepodge of Insect Pine White Neophasia menapia Photos Nature Ali Wild Wanderings Becker's White Pontia beckerii Spring White Pontia sisymbrii Checkered White Pontia protodice Western White Pontia occidentalis The Butterfly Digest by Cabbage White Pieris rapae Bruce Webb - A digest of butterfly discussion around Large Marble Euchloe ausonides the nation. Frontispiece: 1 of 6 12/26/10 9:26 PM Checklist of Buerflies of Tulare County hp://www.natureali.org/Tularebuerflychecklist.htm - 
												
												Papilio (New Series) # 25 2016 Issn 2372-9449
PAPILIO (NEW SERIES) # 25 2016 ISSN 2372-9449 ERNEST J. OSLAR, 1858-1944: HIS TRAVEL AND COLLECTION ITINERARY, AND HIS BUTTERFLIES by James A. Scott, Ph.D. in entomology University of California Berkeley, 1972 (e-mail: [email protected]) Abstract. Ernest John Oslar collected more than 50,000 butterflies and moths and other insects and sold them to many taxonomists and museums throughout the world. This paper attempts to determine his travels in America to collect those specimens, by using data from labeled specimens (most in his remaining collection but some from published papers) plus information from correspondence etc. and a few small field diaries preserved by his descendants. The butterfly specimens and their localities/dates in his collection in the C. P. Gillette Museum (Colorado State University, Fort Collins, Colorado) are detailed. This information will help determine the possible collection locations of Oslar specimens that lack accurate collection data. Many more biographical details of Oslar are revealed, and the 26 insects named for Oslar are detailed. Introduction The last collection of Ernest J. Oslar, ~2159 papered butterfly specimens and several moths, was found in the C. P. Gillette Museum, Colorado State University, Fort Collins, Colorado by Paul A. Opler, providing the opportunity to study his travels and collections. Scott & Fisher (2014) documented specimens sent by Ernest J. Oslar of about 100 Argynnis (Speyeria) nokomis nokomis Edwards labeled from the San Juan Mts. and Hall Valley of Colorado, which were collected by Wilmatte Cockerell at Beulah New Mexico, and documented Oslar’s specimens of Oeneis alberta oslari Skinner labeled from Deer Creek Canyon, [Jefferson County] Colorado, September 25, 1909, which were collected in South Park, Park Co. - 
												
												NORTH AMERICAN BUTTERFLY ASSOCIATION 4 Delaware Road, Morristown, NJ 07960
NORTH AMERICAN BUTTERFLY ASSOCIATION 4 Delaware Road, Morristown, NJ 07960 tel. 973-285-0907 fax 973-285-0936 web: www.naba.org 41st ANNUAL NABA BUTTERFLY COUNT - 2015 INSTRUCTIONS (CANADA) COUNTERS & COUNT CIRCLES Please report your 2015 count results directly to NABA through Unless your count is an existing 1st of July count started before our online count form at www.naba.org. The online form 2008, at least 4 adult observers must participate. One or more allows compilers to enter all data for their counts through the groups of counters always produce better results. It is usually Web and also allows the regional editors to review and edit the desirable to visit several habitats and areas within a count circle, reports more efficiently. Most importantly, Butterfly Count which may be done best by several parties. Since counts are information will be entered and stored in a database which in the open for public participation, we encourage you to publicize future will allow it to be available online to NABA members your count plans! and the public. All compilers are now requested to enter the data A count is held at one or more selected sites within a for their counts through the online count form. If this presents 15-mile diameter CIRCLE. Groups starting a new count MUST any difficulty, please contact NABA for assistance. designate this 15-mile diameter circle. No count circles may overlap—that is, count centers must be a minimum of 15 miles COUNT PROGRAM OVERVIEW apart. Groups repeating a count held a previous year MUST use In order to encourage increased participation in the NABA the same circle, and count the same sites and habitats as before Butterfly Count Program and to encourage even more so far as practical. - 
												
												North American Butterfly Association
NORTH AMERICAN BUTTERFLY ASSOCIATION 4 Delaware Road, Morristown, NJ 07960 tel. 973-285-0907 fax 973-285-0936 web: www.naba.org ANNUAL NABA BUTTERFLY COUNT - INSTRUCTIONS (USA) This printed count forms is for field use only. All counts must submit their results using the online data entry system. Please contact the NABA Count Program if you have questions or concerns. Please report your count results directly to NABA Timing/Requirements for United through our online count form at www.butterflycounts.org. Stated Counts The online form allows compilers to enter all data for their Count NABA 4th of July Butterfly Count: A minimum of four counts through the Web and also allows the regional editors adult observers AND 6 party-hours per count are Date of REQUIRED for all counts started after 2008; and, to review and edit the reports efficiently. Most importantly, June or except in extenuating circumstances, ALL counts should Butterfly Count information will be entered and stored in a July expend at least 6 party-hours of effort. database which in the future will allow it to be available Count online to NABA members and the public. If entering your Date other NABA Seasonal Butterfly Count: A minimum of four adult observers AND 6 party-hours per count is data through the online count form presents any difficulty, than June REQUIRED. please contact NABA for assistance. or July COUNT PROGRAM OVERVIEW DATE OF NEXT YEAR'S COUNT In order to encourage increased participation in the Please contact NABA (at address/phone above, or e-mail NABA Butterfly Count Program and to encourage even more to ) with the date of your next year's monitoring possibilities, the NABA Board of Directors [email protected] authorized the introduction of Seasonal Butterfly Counts in count and information on how to contact the compiler. - 
												
												Nevada Butterflies and Their Biology to Forward Such for Inclusion in the Larger Study
Journal of the Lepidopterists' Society 39(2). 1985. 95-118 NEV ADA BUTTERFLIES: PRELIMINARY CHECKLIST AND DISTRIBUTION GEORGE T. AUSTIN Nevada State Museum and Historical Society, 700 Twin Lakes Drive, Las Vegas, Nevada 89107 ABSTRACT. The distribution by county of the 189 species (over 300 taxa) of but terflies occurring in Nevada is presented along with a list of species incorrectly recorded for the state. There are still large areas which are poorly or not collected. Nevada continues as one of the remaining unknown areas in our knowledge of butterfly distribution in North America. Although a com prehensive work on the state's butterflies is in preparation, there is sufficient demand for a preliminary checklist to justify the following. It is hoped this will stimulate those who have any data on Nevada butterflies and their biology to forward such for inclusion in the larger study. Studies of Nevada butterflies are hampered by a paucity of resident collectors, a large number of mountain and valley systems and vast areas with little or no access. Non-resident collectors usually funnel into known and well worked areas, and, although their data are valu able, large areas of the state remain uncollected. Intensive collecting, with emphasis on poorly known areas, over the past seven years by Nevada State Museum personnel and associates has gone far to clarify butterfly distribution within the state. The gaps in knowledge are now more narrowly identifiable and will be filled during the next few sea sons. There is no all encompassing treatment of Nevada's butterfly fauna. The only state list is an informal recent checklist of species (Harjes, 1980). - 
												
												A SKELETON CHECKLIST of the BUTTERFLIES of the UNITED STATES and CANADA Preparatory to Publication of the Catalogue Jonathan P
A SKELETON CHECKLIST OF THE BUTTERFLIES OF THE UNITED STATES AND CANADA Preparatory to publication of the Catalogue © Jonathan P. Pelham August 2006 Superfamily HESPERIOIDEA Latreille, 1809 Family Hesperiidae Latreille, 1809 Subfamily Eudaminae Mabille, 1877 PHOCIDES Hübner, [1819] = Erycides Hübner, [1819] = Dysenius Scudder, 1872 *1. Phocides pigmalion (Cramer, 1779) = tenuistriga Mabille & Boullet, 1912 a. Phocides pigmalion okeechobee (Worthington, 1881) 2. Phocides belus (Godman and Salvin, 1890) *3. Phocides polybius (Fabricius, 1793) =‡palemon (Cramer, 1777) Homonym = cruentus Hübner, [1819] = palaemonides Röber, 1925 = ab. ‡"gunderi" R. C. Williams & Bell, 1931 a. Phocides polybius lilea (Reakirt, [1867]) = albicilla (Herrich-Schäffer, 1869) = socius (Butler & Druce, 1872) =‡cruentus (Scudder, 1872) Homonym = sanguinea (Scudder, 1872) = imbreus (Plötz, 1879) = spurius (Mabille, 1880) = decolor (Mabille, 1880) = albiciliata Röber, 1925 PROTEIDES Hübner, [1819] = Dicranaspis Mabille, [1879] 4. Proteides mercurius (Fabricius, 1787) a. Proteides mercurius mercurius (Fabricius, 1787) =‡idas (Cramer, 1779) Homonym b. Proteides mercurius sanantonio (Lucas, 1857) EPARGYREUS Hübner, [1819] = Eridamus Burmeister, 1875 5. Epargyreus zestos (Geyer, 1832) a. Epargyreus zestos zestos (Geyer, 1832) = oberon (Worthington, 1881) = arsaces Mabille, 1903 6. Epargyreus clarus (Cramer, 1775) a. Epargyreus clarus clarus (Cramer, 1775) =‡tityrus (Fabricius, 1775) Homonym = argentosus Hayward, 1933 = argenteola (Matsumura, 1940) = ab. ‡"obliteratus" - 
												
												Variation in Wing Pattern and Palatability in a Female-Limited Polymorphic Mimicry System
UC Davis UC Davis Previously Published Works Title Variation in wing pattern and palatability in a female-limited polymorphic mimicry system. Permalink https://escholarship.org/uc/item/3xn706gr Journal Ecology and evolution, 4(23) ISSN 2045-7758 Authors Long, Elizabeth C Hahn, Thomas P Shapiro, Arthur M Publication Date 2014-12-01 DOI 10.1002/ece3.1308 Peer reviewed eScholarship.org Powered by the California Digital Library University of California Variation in wing pattern and palatability in a female- limited polymorphic mimicry system Elizabeth C. Long1,2,3, Thomas P. Hahn4 & Arthur M. Shapiro1,5 1Center for Population Biology, University of California Davis, Davis, California 2Entomology Section, Natural History Museum of Los Angeles County, Los Angeles, California 3UCLA La Kretz Center for California Conservation Science, Institute of the Environment and Sustainability, University of California, Los Angeles, California 4Department of Neurobiology, Physiology, and Behavior, University of California Davis, Davis, California 5Department of Evolution and Ecology, University of California Davis, Davis, California Keywords Abstract Batesian mimicry, Chlosyne hoffmanni, Chlosyne palla, Euphydryas chalcedona, Checkerspot butterflies in the genera Euphydryas and Chlosyne exhibit pheno- palatability. typic polymorphisms along a well-defined latitudinal and elevational gradient in California. The patterns of phenotypic variation in Euphydryas chalcedona, Correspondence Chlosyne palla, and Chlosyne hoffmanni suggest a mimetic relationship; in addi- Elizabeth Long, UCLA La Kretz Center for tion, the specific patterns of variation in C. palla suggest a female-limited poly- California Conservation Science, Entomology morphic mimicry system (FPM). However, the existence of polymorphic Section, Natural History Museum of Los Angeles County, 900 Exposition Boulevard, models runs counter to predictions of mimicry theory. - 
												
												Watching Western Colorado Butterflies
Watching Western Colorado Butterflies Dr. Warren H. “Herb” Wagner Why? • Enhance the outdoor experience • Intro to butterfly conservation • Fun Three Steps to Conservation Xerxes Society for Invertebrate Conservation 1) Identify the resources The butterflies Their larval host plants The adult food sources floral & other 2) Protect the resources The landscape & nectar, host plants 3) Enhance the resources Larval & adult host plants Gardens What are the threats? Invasive weeds Knapweeds, bindweed, cheatgrasses, spurges, … Large scale hot-burning fire More frequent, smaller fires not a threat People Serious collecting benefits long term conservation efforts A few rare species are threatened by over-collecting Habitat destruction is a real and serious threat to many butterfly populations Collecting Butterflies With proper labeling & care COLO Mesa Co • Indisputable historical records Black Ridge Rd • Essential for proper ID 6 Apr 1996 • Genetic records JA & RW Hammon • Fun! • Pretty Paonia Colo 7/23/1932 If you are going to use a field guide, this is the one Colorado Butterflies 289 Species West slope - > 200 species Gunnison County 159 species Mesa County 155 species • Skippers 36 species • Skippers 36 species • Swallowtails 6 species • Swallowtails 7 species • Lycaenids 40 species • Lycaenids 39 species • Pierids 20 species • Pierids 16 species • Metalmarks 1 species • Metalmarks 1 species • Nymphalids 56 species • Nymphalids 56 species Metamorphosis naturallycuriouswithmaryholland Lepidoptera = Scale wing The proboscis, clubbed antenna