NOPES Newsletter 3 20Small
Total Page:16
File Type:pdf, Size:1020Kb
Load more
Recommended publications
-
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE
Guide to the Flora of the Carolinas, Virginia, and Georgia, Working Draft of 17 March 2004 -- LILIACEAE LILIACEAE de Jussieu 1789 (Lily Family) (also see AGAVACEAE, ALLIACEAE, ALSTROEMERIACEAE, AMARYLLIDACEAE, ASPARAGACEAE, COLCHICACEAE, HEMEROCALLIDACEAE, HOSTACEAE, HYACINTHACEAE, HYPOXIDACEAE, MELANTHIACEAE, NARTHECIACEAE, RUSCACEAE, SMILACACEAE, THEMIDACEAE, TOFIELDIACEAE) As here interpreted narrowly, the Liliaceae constitutes about 11 genera and 550 species, of the Northern Hemisphere. There has been much recent investigation and re-interpretation of evidence regarding the upper-level taxonomy of the Liliales, with strong suggestions that the broad Liliaceae recognized by Cronquist (1981) is artificial and polyphyletic. Cronquist (1993) himself concurs, at least to a degree: "we still await a comprehensive reorganization of the lilies into several families more comparable to other recognized families of angiosperms." Dahlgren & Clifford (1982) and Dahlgren, Clifford, & Yeo (1985) synthesized an early phase in the modern revolution of monocot taxonomy. Since then, additional research, especially molecular (Duvall et al. 1993, Chase et al. 1993, Bogler & Simpson 1995, and many others), has strongly validated the general lines (and many details) of Dahlgren's arrangement. The most recent synthesis (Kubitzki 1998a) is followed as the basis for familial and generic taxonomy of the lilies and their relatives (see summary below). References: Angiosperm Phylogeny Group (1998, 2003); Tamura in Kubitzki (1998a). Our “liliaceous” genera (members of orders placed in the Lilianae) are therefore divided as shown below, largely following Kubitzki (1998a) and some more recent molecular analyses. ALISMATALES TOFIELDIACEAE: Pleea, Tofieldia. LILIALES ALSTROEMERIACEAE: Alstroemeria COLCHICACEAE: Colchicum, Uvularia. LILIACEAE: Clintonia, Erythronium, Lilium, Medeola, Prosartes, Streptopus, Tricyrtis, Tulipa. MELANTHIACEAE: Amianthium, Anticlea, Chamaelirium, Helonias, Melanthium, Schoenocaulon, Stenanthium, Veratrum, Toxicoscordion, Trillium, Xerophyllum, Zigadenus. -
The Vascular Plants of Massachusetts
The Vascular Plants of Massachusetts: The Vascular Plants of Massachusetts: A County Checklist • First Revision Melissa Dow Cullina, Bryan Connolly, Bruce Sorrie and Paul Somers Somers Bruce Sorrie and Paul Connolly, Bryan Cullina, Melissa Dow Revision • First A County Checklist Plants of Massachusetts: Vascular The A County Checklist First Revision Melissa Dow Cullina, Bryan Connolly, Bruce Sorrie and Paul Somers Massachusetts Natural Heritage & Endangered Species Program Massachusetts Division of Fisheries and Wildlife Natural Heritage & Endangered Species Program The Natural Heritage & Endangered Species Program (NHESP), part of the Massachusetts Division of Fisheries and Wildlife, is one of the programs forming the Natural Heritage network. NHESP is responsible for the conservation and protection of hundreds of species that are not hunted, fished, trapped, or commercially harvested in the state. The Program's highest priority is protecting the 176 species of vertebrate and invertebrate animals and 259 species of native plants that are officially listed as Endangered, Threatened or of Special Concern in Massachusetts. Endangered species conservation in Massachusetts depends on you! A major source of funding for the protection of rare and endangered species comes from voluntary donations on state income tax forms. Contributions go to the Natural Heritage & Endangered Species Fund, which provides a portion of the operating budget for the Natural Heritage & Endangered Species Program. NHESP protects rare species through biological inventory, -
Phylogenetic Placement of the Enigmatic Orchid Genera Thaia and Tangtsinia: Evidence from Molecular and Morphological Characters
TAXON 61 (1) • February 2012: 45–54 Xiang & al. • Phylogenetic placement of Thaia and Tangtsinia Phylogenetic placement of the enigmatic orchid genera Thaia and Tangtsinia: Evidence from molecular and morphological characters Xiao-Guo Xiang,1 De-Zhu Li,2 Wei-Tao Jin,1 Hai-Lang Zhou,1 Jian-Wu Li3 & Xiao-Hua Jin1 1 Herbarium & State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, P.R. China 2 Key Laboratory of Biodiversity and Biogeography, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650204, P.R. China 3 Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Menglun Township, Mengla County, Yunnan province 666303, P.R. China Author for correspondence: Xiao-Hua Jin, [email protected] Abstract The phylogenetic position of two enigmatic Asian orchid genera, Thaia and Tangtsinia, were inferred from molecular data and morphological evidence. An analysis of combined plastid data (rbcL + matK + psaB) using Bayesian and parsimony methods revealed that Thaia is a sister group to the higher epidendroids, and tribe Neottieae is polyphyletic unless Thaia is removed. Morphological evidence, such as plicate leaves and corms, the structure of the gynostemium and the micromorphol- ogy of pollinia, also indicates that Thaia should be excluded from Neottieae. Thaieae, a new tribe, is therefore tentatively established. Using Bayesian and parsimony methods, analyses of combined plastid and nuclear datasets (rbcL, matK, psaB, trnL-F, ITS, Xdh) confirmed that the monotypic genus Tangtsinia was nested within and is synonymous with the genus Cepha- lanthera, in which an apical stigma has evolved independently at least twice. -
Phylogeny, Character Evolution and the Systematics of Psilochilus (Triphoreae)
THE PRIMITIVE EPIDENDROIDEAE (ORCHIDACEAE): PHYLOGENY, CHARACTER EVOLUTION AND THE SYSTEMATICS OF PSILOCHILUS (TRIPHOREAE) A Dissertation Presented in Partial Fulfillment of the Requirements for The Degree Doctor of Philosophy in the Graduate School of the Ohio State University By Erik Paul Rothacker, M.Sc. ***** The Ohio State University 2007 Doctoral Dissertation Committee: Approved by Dr. John V. Freudenstein, Adviser Dr. John Wenzel ________________________________ Dr. Andrea Wolfe Adviser Evolution, Ecology and Organismal Biology Graduate Program COPYRIGHT ERIK PAUL ROTHACKER 2007 ABSTRACT Considering the significance of the basal Epidendroideae in understanding patterns of morphological evolution within the subfamily, it is surprising that no fully resolved hypothesis of historical relationships has been presented for these orchids. This is the first study to improve both taxon and character sampling. The phylogenetic study of the basal Epidendroideae consisted of two components, molecular and morphological. A molecular phylogeny using three loci representing each of the plant genomes including gap characters is presented for the basal Epidendroideae. Here we find Neottieae sister to Palmorchis at the base of the Epidendroideae, followed by Triphoreae. Tropidieae and Sobralieae form a clade, however the relationship between these, Nervilieae and the advanced Epidendroids has not been resolved. A morphological matrix of 40 taxa and 30 characters was constructed and a phylogenetic analysis was performed. The results support many of the traditional views of tribal composition, but do not fully resolve relationships among many of the tribes. A robust hypothesis of relationships is presented based on the results of a total evidence analysis using three molecular loci, gap characters and morphology. Palmorchis is placed at the base of the tree, sister to Neottieae, followed successively by Triphoreae sister to Epipogium, then Sobralieae. -
Vegetative Anatomy of Calypsoeae (Orchidaceae) William Louis Stern Florida International University
Eastern Illinois University The Keep Faculty Research & Creative Activity Biological Sciences January 2008 Vegetative anatomy of Calypsoeae (Orchidaceae) William Louis Stern Florida International University Barbara S. Carlsward Eastern Illinois University, [email protected] Follow this and additional works at: http://thekeep.eiu.edu/bio_fac Part of the Biology Commons Recommended Citation Stern, William Louis and Carlsward, Barbara S., "Vegetative anatomy of Calypsoeae (Orchidaceae)" (2008). Faculty Research & Creative Activity. 265. http://thekeep.eiu.edu/bio_fac/265 This Article is brought to you for free and open access by the Biological Sciences at The Keep. It has been accepted for inclusion in Faculty Research & Creative Activity by an authorized administrator of The Keep. For more information, please contact [email protected]. LANKESTERIANA 8(1): 105-112. 2008. VEGETATIVE ANATOMY OF CALYPSOEAE (ORCHIDACEAE) WILLIAM LOUIS STERN1 & BARBARA S. CARLSWARD2,3 1Department of Biological Sciences, Biscayne Bay Campus, MSB 357, Florida International University, North Miami, Florida 33181, USA 2Department of Biological Sciences, Eastern Illinois University, Charleston, Illinois 61920-3099, USA 3Corresponding author: [email protected] ABSTRACT. Calypsoeae represent a small tribe of anatomically little-known orchids with a wide distribution in the Western Hemisphere. Leaves are present in all genera, except Corallorhiza and Wullschlaegelia both of which are subterranean taxa. Stomata are abaxial (ad- and abaxial in Aplectrum) and tetracytic (anomocytic in Calypso). Fiber bundles are absent in leaves of all taxa examined except Govenia tingens. Stegmata are present in leaves of only Cremastra and Govenia. Roots are velamentous, except in filiform roots of Wullschlaegelia. Vegetative anatomy supports a relationship between Wullschlaegelia and Corallorhiza but does not support the grouping of winter-leaved Aplectrum and Tipularia nor proposed groupings of genera based on pollinarium features. -
2011 November Guess That Plant: Tipularia Discolor (Pursh) Nutt
2011 November Guess that Plant: Tipularia discolor (Pursh) Nutt. – Crane fly orchis November’s Guess that Plant was the cranefly orchis, know in other parts of the Southeast as the crippled cranefly. This unusual orchid is a denizen of the hardwood forests. The elegant feature of this species is it is easiest discovered during the dormant season, late October through early May. The plant’s reproductive strategy is interesting, providing photosynthetic tissue (leaves) during the dormant season for deciduous canopy trees. As canopy trees sleep the winter away, their open branches allow sufficient light to reach the forest floor. The thick and well protected leaves exist within a near-earth micro-climate that allows food production even on the coldest of days, as long as there is no snow cover. The temperature at ground level can reach 60o F by solar radiation, which is more than warm enough to allow the photosynthesis. As the season warms in late April and early May, our deciduous canopy trees return the dense leaf cover, signaling the cranefly to shed its basal leaves. From this point into summer, plant is subterranean. This habit changes in late July and early August when it pushes a 12-18” flower stalk from the buried buds. The plant is now ready to flower. The copper to flesh toned flower raceme is not the striking display visible on some of our other more flashy species. This neutral color makes observation by human eyes difficult. Once fertilized it becomes a multi-ribbed capsule that droops on the stem, distributing seeds until the capsule is either emptied or buried by the next winter snow event. -
A Two Year Population Ecology Study of Puttyroot Orchid (Aplectrum Hyemale (Muhl
2014. Proceedings of the Indiana Academy of Science 123(2):131–137 A TWO YEAR POPULATION ECOLOGY STUDY OF PUTTYROOT ORCHID (APLECTRUM HYEMALE (MUHL. EX WILLD.) TORR.) IN CENTRAL INDIANA Megan E. Smith1 and Alice L. Heikens2: Department of Biology, Franklin College, 101 Branigin Boulevard, Franklin, IN 46131 USA ABSTRACT. Aplectrum hyemale, puttyroot orchid, is a terrestrial, winter perennial found in rich, mesic forests throughout the Midwest as well as in Hougham Woods Biological Field Station (HWBFS) near Franklin in Johnson County, Indiana. This orchid overwinters as a single basal leaf that may produce a flowering shoot in spring. The size of the Aplectrum population at HWBFS remained relatively stable with 305 to 311 individuals during this 2012-2013 study. Only 2.9% of the plants flowered and only one plant produced fruits in 2012. No flowering or fruiting occurred in 2013. Plants that produced flowering shoots had basal leaves that were larger than the vegetative plants. However, basal leaf size was not correlated with the number of flowers per plant. It appears that weather, including the unusually warm spring and summer drought in 2012 and the dry spring in 2013, affected phenology and reproduction of Aplectrum. Keywords: Aplectrum hyemale, puttyroot, Adam-and-Eve orchid, mesic woods, phenology INTRODUCTION used as a paste to mend broken pottery (Correll Puttyroot orchid, Aplectrum hyemale,is 1950; Whiting & Catling 1986). It is also known a winter perennial that is found throughout as the Adam-and-Eve orchid because of the the eastern deciduous forest in mesic woods in paired corms (Porcher & Rayner 2001). -
The Vascular Flora of the Red Hills Forever Wild Tract, Monroe County, Alabama
The Vascular Flora of the Red Hills Forever Wild Tract, Monroe County, Alabama T. Wayne Barger1* and Brian D. Holt1 1Alabama State Lands Division, Natural Heritage Section, Department of Conservation and Natural Resources, Montgomery, AL 36130 *Correspondence: wayne [email protected] Abstract provides public lands for recreational use along with con- servation of vital habitat. Since its inception, the Forever The Red Hills Forever Wild Tract (RHFWT) is a 1785 ha Wild Program, managed by the Alabama Department of property that was acquired in two purchases by the State of Conservation and Natural Resources (AL-DCNR), has pur- Alabama Forever Wild Program in February and Septem- chased approximately 97 500 ha (241 000 acres) of land for ber 2010. The RHFWT is characterized by undulating general recreation, nature preserves, additions to wildlife terrain with steep slopes, loblolly pine plantations, and management areas and state parks. For each Forever Wild mixed hardwood floodplain forests. The property lies tract purchased, a management plan providing guidelines 125 km southwest of Montgomery, AL and is managed by and recommendations for the tract must be in place within the Alabama Department of Conservation and Natural a year of acquisition. The 1785 ha (4412 acre) Red Hills Resources with an emphasis on recreational use and habi- Forever Wild Tract (RHFWT) was acquired in two sepa- tat management. An intensive floristic study of this area rate purchases in February and September 2010, in part was conducted from January 2011 through June 2015. A to provide protected habitat for the federally listed Red total of 533 taxa (527 species) from 323 genera and 120 Hills Salamander (Phaeognathus hubrichti Highton). -
Systematics, Phylogeography, Fungal Associations, and Photosynthesis
Systematics, Phylogeography, Fungal Associations, and Photosynthesis Gene Evolution in the Fully Mycoheterotrophic Corallorhiza striata Species Complex (Orchidaceae: Epidendroideae) Dissertation Presented in Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy in the Graduate School of the Ohio State University By Craig Francis Barrett, M. S. Evolution, Ecology, and Organismal Biology ***** The Ohio State University 2010 Dissertation Committee: Dr. John V. Freudenstein, Advisor Dr. John W. Wenzel Dr. Andrea D. Wolfe Copyright by Craig Francis Barrett 2010 ABSTRACT Corallorhiza is a genus of obligately mycoheterotrophic (fungus-eating) orchids that presents a unique opportunity to study phylogeography, taxonomy, fungal host specificity, and photosynthesis gene evolution. The photosysnthesis gene rbcL was sequenced for nearly all members of the genus Corallorhiza; evidence for pseudogene formation was found in both the C. striata and C. maculata complexes, suggesting multiple independent transitions to complete heterotrophy. Corallorhiza may serve as an exemplary system in which to study the plastid genomic consequences of full mycoheterotrophy due to relaxed selection on photosynthetic apparatus. Corallorhiza striata is a highly variable species complex distributed from Mexico to Canada. In an investigation of molecular and morphological variation, four plastid DNA clades were identified, displaying statistically significant differences in floral morphology. The biogeography of C. striata is more complex than previously hypothesized, with two main plastid lineages present in both Mexico and northern North America. These findings add to a growing body of phylogeographic data on organisms sharing this common distribution. To investigate fungal host specificity in the C. striata complex, I sequenced plastid DNA for orchids and nuclear DNA for fungi (n=107 individuals), and found that ii the four plastid clades associate with divergent sets of ectomycorrhizal fungi; all within a single, variable species, Tomentella fuscocinerea. -
Diversity and Evolution of Monocots
Lilioids - petaloid monocots 4 main groups: Diversity and Evolution • Acorales - sister to all monocots • Alismatids of Monocots – inc. Aroids - jack in the pulpit ! • Lilioids (lilies, orchids, yams) – grade, non-monophyletic – petaloid . orchids and palms . ! • Commelinoids – Arecales – palms – Commelinales – spiderwort – Zingiberales –banana – Poales – pineapple – grasses & sedges Lilioids - petaloid monocots Asparagales: *Orchidaceae - orchids • finish the Asparagales by 1. Terrestrial/epiphytes: plants looking at the largest family - typically not aquatic the orchids 2. Geophytes: herbaceous above ground with below ground modified perennial stems: bulbs, corms, rhizomes, tubers 3. Tepals: showy perianth in 2 series of 3 each; usually all petaloid, or outer series not green and sepal-like & with no bracts 1 *Orchidaceae - orchids *Orchidaceae - orchids The family is diverse with about 880 genera and over 22,000 All orchids have a protocorm - a feature restricted to the species, mainly of the tropics family. Orchids are • structure formed after germination and before the mycotrophic (= fungi development of the seedling plant dependent) lilioids; • has no radicle but instead mycotrophic tissue some are obligate mycotrophs Cypripedium acaule Corallorhiza striata Stemless lady-slipper Striped coral root Dactylorhiza majalis protocorm *Orchidaceae - orchids *Orchidaceae - orchids Cosmopolitan, but the majority of species are found in the Survive in these epiphytic and other harsh environments via tropics and subtropics, ranging from sea -
Vegetation Community Monitoring at Congaree National Park: 2014 Data Summary
National Park Service U.S. Department of the Interior Natural Resource Stewardship and Science Vegetation Community Monitoring at Congaree National Park 2014 Data Summary Natural Resource Data Series NPS/SECN/NRDS—2016/1016 ON THIS PAGE Tiny, bright yellow blossoms of Hypoxis hirsuta grace the forest floor at Congaree National Park. Photograph courtesy of Sarah C. Heath, Southeast Coast Network. ON THE COVER Spiraling compound leaf of green dragon (Arisaema dracontium) at Congaree National Park. Photograph courtesy of Sarah C. Heath, Southeast Coast Network Vegetation Community Monitoring at Congaree National Park 2014 Data Summary Natural Resource Data Series NPS/SECN/NRDS—2016/1016 Sarah Corbett Heath1 and Michael W. Byrne2 1National Park Service Southeast Coast Inventory and Monitoring Network Cumberland Island National Seashore 101 Wheeler Street Saint Marys, GA 31558 2National Park Service Southeast Coast Inventory and Monitoring Network 135 Phoenix Drive Athens, GA 30605 May 2016 U.S. Department of the Interior National Park Service Natural Resource Stewardship and Science Fort Collins, Colorado The National Park Service, Natural Resource Stewardship and Science office in Fort Collins, Colorado, publishes a range of reports that address natural resource topics. These reports are of interest and applicability to a broad audience in the National Park Service and others in natural resource management, including scientists, conservation and environmental constituencies, and the public. The Natural Resource Data Series is intended for the timely release of basic data sets and data summaries. Care has been taken to assure accuracy of raw data values, but a thorough analysis and interpretation of the data has not been completed. -
Maryland Native Plant Society Plant Lists
Maryland Native Plant Society Plant Lists We offer these lists to individuals and groups to enhance the enjoyment and study of plants of different locations in Maryland and nearby states. Their accuracy has not been verified by the Maryland Native Plant Society. FLAG PONDS NATURE PARK, CALVERT COUNTY, MD This list records plants seen during five field visits to the Flag Ponds Nature Park in Lusby, Maryland in 2014 on March 14th, April 11th, May 9th, June 13th, and July 11th. Field trips were led by Karyn Molines. Plant lists by Karyn Molines. Nomenclature follows the USDA Plant Database at http://plants.usda.gov (February 2014). Synonyms are footnoted for some species. Acer rubrum Red maple Aceraceae Acer saccharinum Silver maple Aceraceae Achillea millefolium Yarrow Asteraceae Ailanthus altissima Ailanthus Simaroubaceae Allium sp. Field garlic Liliaceae Andropogon virginicus Broomsedge Poaceae Antennaria sp. Pussytoes Asteraceae Aplectrum hyemale Puttyroot Orchidaceae Aquilegia canadensis Wild columbine Ranunculaceae Aralia spinosa Devil's walking stick Araliaceae Arisaema triphyllum Jack-in the-pulpit Araceae Asimina triloba Pawpaw Annonaceae Asplenium platyneuron Ebony spleenwort Aspleniaceae Berberis thunbergii Japanese barberry Berberidaceae Betula nigra River birch Betulaceae Boehmeria cylindrica False nettle Urticaceae Botrychium sp. Grapefern Ophioglossaceae Botrychium virginianum Rattlesnake fern Ophioglossaceae Campsis radicans Trumpet creeper Bignoniaceae Cardamine concatenata1 Cut-leaved toothwort Brassicaceae Cardamine hirsuta