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From the Lower Cretaceous of South Korea and Its Paleobiogeographic Implication

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See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/259510971 A new ichthyodectiform (Pisces, Teleostei) from the Lower Cretaceous of South Korea and its paleobiogeographic implication Article in Cretaceous Research · January 2014 DOI: 10.1016/j.cretres.2013.11.007 CITATIONS READS 2 321 4 authors, including: Haangmook Kim Mee-mann Chang Pusan National University Chinese Academy of Sciences 1 PUBLICATION 4 CITATIONS 48 PUBLICATIONS 631 CITATIONS SEE PROFILE SEE PROFILE Feixiang Wu Chinese Academy of Sciences 10 PUBLICATIONS 146 CITATIONS SEE PROFILE Available from: Feixiang Wu Retrieved on: 22 July 2016 Cretaceous Research 47 (2014) 117e130 Contents lists available at ScienceDirect Cretaceous Research journal homepage: www.elsevier.com/locate/CretRes A new ichthyodectiform (Pisces, Teleostei) from the Lower Cretaceous of South Korea and its paleobiogeographic implication Haang-Mook Kim a, Mee-Mann Chang b,*, Feixiang Wu b, Yang-Hee Kim a a College of Sciences, Department of Geological Environments, Pusan National University, Busan 609-735, Republic of Korea b Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China article info abstract Article history: Here we report the first ichthyodectiform in the Jinju Biota from the Lower Cretaceous Jinju Formation, Received 5 August 2013 Shindong Group, at Hyojagyo, Jinju City, South Korea. It shows the following ichthyodectiform charac- Accepted in revised form 12 November 2013 ters: 1) jaw teeth in single series; 2) coracoid expanded ventrally; 3) dorsal fin situated posteriorly, with Available online its origin opposite to that of long anal fin. Besides, its caudal skeleton also exhibits certain features often seen in basal ichthyodectiforms and some other primitive teleosts. The fish is noticeably similar to the Keywords: species referred to Chuhsiungichthys from the upper Lower Cretaceous Dobaru and Kumagai formations Jinjuichthys of the Wakino Subgroup of Kanmon Group in Kyushu, Japan, and Chuhsiungichthys tsanglingensis from Ichthyodectiformes Late Early Cretaceous the Jiangdihe Formation of similar age from Chuxiong, Yunnan Province, China. Chuhsiungichthys was, in South Korea turn, comparable with Mesoclupea showchangensis from the upper Lower Cretaceous Shouchang For- Cosmopolitan fishes indicating West Pacific mation in Shouchang and Linhai, and Guantou Formation in Zhuji, Zhejiang Province, China. The latter rising two genera also show the ichthyodectiform characters mentioned above. Nevertheless, the fish from Korea, those referred to Chuhsiungichthys from Japan and China, and Mesoclupea can easily be distin- guished from other ichthyodectiforms in their higher body depth and shorter length; proportionally longer head; longer dorsal and anal fins; lower number of vertebrae, with abdominal less than caudal; vertebrae deeper than long. Among them the Jinju fish is distinct from the other two genera in even higher and shorter body, and presence of urodermal. In addition, the Jinju fish is different from Meso- clupea showchangensis in having a much longer dorsal fin and lower number of vertebrae, but different from Chuhsiungichthys tsanglingensis in having a much higher number of vertebrae and more ridges and grooves on the lateral face of vertebrae. Thus, the Jinju fish cannot be referred to either Chuhsiungichthys or Mesoclupea, and establishing a new genus and species for it is warranted. The Asian ichthyodectiforms, Mesoclupea, Chuhsiungichthys, and Jinjuichthys, may most probably form a monophyletic group e Chuhsiungichthyidae, known so far restricted to the upper Lower Cretaceous fresh and/or brackish waters of East Asia. The new ichthyodectiform materials from Korea are better preserved than those from Japan, thereby allowing a relatively complete description of the fish, providing more information for discussion of its phylogenetic position, enriching the diversity of the local fish fauna, and helping us better understand the paleobiogeographical distribution of the group and its geological background. Ó 2013 Elsevier Ltd. All rights reserved. 1. Introduction numerous fossils, including vertebrates, invertebrates, and plants, were collected from the Jinju Formation (see Fig. 2B for the A large-scale paleontological excavation was carried out by geological section of this part of the formation), and the name “Jinju Haang-Mook Kim and his team from the Pusan University in the Biota” was first used for these fossils. The Jinju Biota was then period of 2005e2007, in the area close to Jinju city, during the correlated with the Early Cretaceous Jehol Biota from northwestern construction work for broadening the highway between Jibhyeon China, particularly with the fossils from the Yixian Formation. In and Saengbiryang (Kim, 2009; Figs. 1 and 2A). In the excavation, this paper we describe two teleost specimens collected from the Jinju Formation, Shindong Group, Gyeongsang Supergroup at Hyojagyo, Micheon-myeon, Jinju City, South Gyeongsang Province, * Corresponding author. South Korea. They show a close resemblance to the specimens E-mail addresses: [email protected] (H.-M. Kim), [email protected] described by Yabumoto (1994) from the Lower Cretaceous Dobaru (M.-M. Chang), [email protected] (F. Wu), [email protected] (Y.-H. Kim). 0195-6671/$ e see front matter Ó 2013 Elsevier Ltd. All rights reserved. http://dx.doi.org/10.1016/j.cretres.2013.11.007 118 H.-M. Kim et al. / Cretaceous Research 47 (2014) 117e130 survey of the Ichthyodectiformes, and considered the order as a basal teleostean group, sister to Tharsis dubius plus all extant tele- osts. A recent work by Cavin et al. (2012) has shed much new light on this group. Although Cavin et al. (2012) excluded Ascalabo- thrissops (Arratia, 2000) and Pachythrissops (Woodward, 1919) from the Ichthyodectiformes in their comprehensive phylogenetic anal- ysis we still cite some characters of these two taxa for the purpose of comparison, wherever information is available. Fish belonging to the Chuhsiungichthyidae (incorrectly spelt as “chuingichthiid”) (Order Ichthyodectiformes) was mentioned by Lee et al. (2001,p. 363) from “the Jinju Formation exposed in Donggogri (Habin- myeon, Dalseong-gun, North Gyeongsang Province)” among other fossil fishes, and listed in Lee et al. (2001), table 1 (p. 358). The age of the Shindong Group, the Jinju Formation being its youngest unit, has been designated as the HauterivianeBarremian by palynolo- gists and ostracod workers, and as Aptian to Albian by mollusc workers (Yang, 1982; Yi et al., 1994; and Choi, 1985 in Lee et al., 2001; Sha et al., 2012). Recently a UePb dating based on detrital zircons of the group was given by Lee et al. (2010) as ranging from the Aptian to Albian. In this paper, we describe the two well- preserved fossil fish specimens from Hyojagyo, Micheon-myeon and compare them with other known ichthyodectiforms, espe- cially Chuhsiungichthys and Mesoclupea. For comparison, a spec- imen of well-preserved impression of Mesoclupea showchangensis without head region (GMC V 1007-1) is also described in details. We conclude that the specimens from the Jinju Formation described here cannot be assigned to any of the known ichthyo- dectiforms. Therefore, we must treat them as a new genus and species of the order. However, we feel that a meaningful phyloge- netic analysis has to wait for sufficient morphological information of the three Asian forms to emerge from future discoveries. The Jinju fish fauna (Ichthyodectiformes, Osteoglossiformes etc.) is Fig. 1. The geological map of Gyeongsang Basin, the black dot indicates the fish lo- similar in composition to the fish fauna from Kyushu, Japan and cality, the circles indicate cities. southeast China, and forms one and the same ichthyofauna with those from the latter two areas. This ichthyofauna has a slightly younger age and different geological background than the fish fauna from the Jehol Biota, and contained several cosmopolitan fish and Kumagai formations of the Wakino Subgroup of Kanmon forms. Group in Kyushu, Japan (Yabumoto et al., 2006). Yabumoto (1994) established two new species, namely, Chuhsiungichthys yanagidai (from Dobaru Formation) and C. japonicus (from Kumagai Forma- 2. Materials and methods tion), referred to the genus Chuhsiungichthys described by Lew (1974) from the Jiangdihe Formation of similar age from Chux- The material described in this paper consists of two specimens. iong (the Pinyin spelling adopted after 1979 in mainland China that One is a fish skeleton without the caudal portion (PSU V 1011), and has replaced the old spelling of Chuhsiung in Wade-Giles romani- the other is a posterior part of a fish with well-preserved caudal zation), Yunnan Province, China. Yabumoto (1994) was the first to skeleton, caudal fin and relatively large parts of dorsal and anal refer the genus to the early teleost order Ichthyodectiformes fins (PSU V 1012). The specimens are preserved in laminated black (bulldog fish), and he established a new family Chuhsiungichthyi- siltstone, and both the matrix and skeletons are of similar black dae to include Chuhsiungichthys and another genus Mesoclupea, color, which caused great difficulty in preparation. Nevertheless, described by Ping and Yen (1933) from the Lower Cretaceous our preparator, using just thin needles, managed to do a beautiful Shouchang Formation, now designated as the basal part of Aptian job
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  • Teleostei: Ichthyodectiformes) from the Cenomanian of Morocco

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    Palaeontologia Electronica http://palaeo-electronica.org A NEW SPECIES OF CLADOCYCLUS (TELEOSTEI: ICHTHYODECTIFORMES) FROM THE CENOMANIAN OF MOROCCO Peter L. Forey and Lionel Cavin Peter L. Forey. Department of Palaeontology, The Natural History Museum, Cromwell Road, London SW7 5BD, England. [email protected] Lionel Cavin. Department of Geology and Palaeontology, Muséum d'Histoire Naturelle, CP 6434 1211, Genève 6, Switzerland. [email protected] ABSTRACT A species of ichthyodectiform fish, Cladocyclus pankowskii sp.nov. is described based on a single well-preserved braincase specimen from the Cenomanian Kem Kem beds of southern Morocco. It differs from the type species, C. gardneri, in the propor- tions of the skull roof. The presence of a species of Cladocyclus in Morocco adds yet another vicariant taxon shared between Brazil and Africa in the mid-Cretaceous, when Africa was separating from South America during the formation of the South Atlantic. Key Words: new species; anatomy; actinopterygian; palaeobiogeography INTRODUCTION more derived ichthyodectiforms (see below). Ich- thyodectiforms are an extinct group of about 30 The braincase described here belongs to an species of primitive teleosts ranging in time from ichthyodectiform teleost. It shows a well-developed the Bathonian to Maastrichtian. Most were marine ethmopalatine (a synapomorphy of all but the most fishes and many had structural adaptations sug- primitive of ichthyodectiforms), as well as the fol- gesting they were fast open water swimmers. lowing combination of features: median parietals Some must have been the top predators of their meeting in front of a large supraoccipital, large day. The phylogeny of ichthyodectiforms has been intercalar that forms a significant part of the hyo- addressed on several occasions (Bardack 1965; mandibular facet as well as the skull roof, a jugular Patterson and Rosen 1977; Maisey 1991; Stewart canal opening into the floor of a deep subtemporal 1999; Taverne and Chanet 2000).
  • Family-Group Names of Fossil Fishes

    Family-Group Names of Fossil Fishes

    European Journal of Taxonomy 466: 1–167 ISSN 2118-9773 https://doi.org/10.5852/ejt.2018.466 www.europeanjournaloftaxonomy.eu 2018 · Van der Laan R. This work is licensed under a Creative Commons Attribution 3.0 License. Monograph urn:lsid:zoobank.org:pub:1F74D019-D13C-426F-835A-24A9A1126C55 Family-group names of fossil fishes Richard VAN DER LAAN Grasmeent 80, 1357JJ Almere, The Netherlands. Email: [email protected] urn:lsid:zoobank.org:author:55EA63EE-63FD-49E6-A216-A6D2BEB91B82 Abstract. The family-group names of animals (superfamily, family, subfamily, supertribe, tribe and subtribe) are regulated by the International Code of Zoological Nomenclature. Particularly, the family names are very important, because they are among the most widely used of all technical animal names. A uniform name and spelling are essential for the location of information. To facilitate this, a list of family- group names for fossil fishes has been compiled. I use the concept ‘Fishes’ in the usual sense, i.e., starting with the Agnatha up to the †Osteolepidiformes. All the family-group names proposed for fossil fishes found to date are listed, together with their author(s) and year of publication. The main goal of the list is to contribute to the usage of the correct family-group names for fossil fishes with a uniform spelling and to list the author(s) and date of those names. No valid family-group name description could be located for the following family-group names currently in usage: †Brindabellaspidae, †Diabolepididae, †Dorsetichthyidae, †Erichalcidae, †Holodipteridae, †Kentuckiidae, †Lepidaspididae, †Loganelliidae and †Pituriaspididae. Keywords. Nomenclature, ICZN, Vertebrata, Agnatha, Gnathostomata.