Studies in the Genus Acacia (Mimosaccae),1
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Studiesin the genusAcacia (Mimosaceae)-8 A revisionof the Uninerves-Triangulares,in part (the tetramerousspecies) By B. R. Maslin Abstract A taxonon,ic revision of parl of Acacia Series Urr'fte/yeris presented. Includedare those species with 4-merousflowers and triangular-shapedphyllodes. The seventeenspecies comprising this group are endemicto south-westWestern Australia. Eight new speciesand one new variety are descrlbe A. delphina sp. !9y., ,4./i/tolea sp. nov. (syn. A. decipiensaluct.), A. phaeocalyxsp. no\., A, phlebopetalasp. nov., A. phlebopetalavar. pube:cens var. nov., A. pycnocephala sp. nov., ,4. tobinae sp. noy., A. Mmitrullata sp. nov. and A. Lliginosasp. lroy. A new name, A, chtl'stcephala,is gilen to the taxon A. biflora var. airca E, Pritzel. Three names previously in common use have been relegatedto syn- onymy'.A. cuneataBenth.and A. decipiens(C. Koen.) R.Bt. are no.\'/A. tunca,a (Burm.f.) Hort. ex Hoffmannseggwhile,4. l'errlcosaW. V. Fitzg. is A. i crossam Hook. Keys io speciesand varietiesare provided,all taxa are illustratedand their distributionmapped. Introduction The presentrevision is basedon Bentham's 1864and 1875treatments of "UnineNes-Triangulares" the (see below) but treats only those specieswith 4-merousflowers. Thus only Western Australian ta\a are considered. The speciesincluded by Bentham but excluded here are A. acqnthocladaF. Muell., A. bidentata Benth., l. dilatata Benth. a\d. A. gunnii Benth. (syn A. yomeri- .formis A. Cunn. ex Benth.). Three speciesand two varieties described since 1875 are included here, viz. A. inops Maidet et Blakely, A. mooreena W. Y. Fitzg., A. yernicosaW. Y. Fitzg. (: A. inuassata Hook.), l. bifiora var. aurea E. Pritzel (: A. chrysocephsldnom. et stat. nov.) and A. horridula rar. hastul- stoides E. Prttzel ( A. uliginosasp. no\.). ln addition. eighr new specie' and one new variety are described. The terminologyused to describeplant communitiesis that presentedby Aplin (1976). This is a slightly modified versionof the systemproposed by Specht(1970). All illustrations appearing in this work have been drawn from pressed herbanum specrmens. Only a selectionof specimensis cited under each taxon. A list of the numbered specimensseen is given at the end of this paper. Some of the un- numbered specimensseen are cited in the text. Historical perspective Benthan (1841) described lhe Triangularcs as one of the eleven major subdivisions of his SeriesPiyl/odineae and incltded specieswith both uninerved and plurinerved phyllodes. Meisner (1844 and 1848).Bentham (1855) and Mueller (1859) later describednew taxa which were referred to the Trians;lare;. 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Principrl phyllode types of the Uninerves-Triangulares. A A. hoffi.lula-type (phyllode semitrultrte). B-A. billorc-type(phyllode obliquely obdeltate). C-A. tntncata- type (phyllode obtriangular). abm-abaxial margin; adm-adaxial distal margin; apm- adaxialproximal margin; g gland; m apicalmucro; p pulvinus;pn principalnerve; sn secondarynerve. The phyllodes have a principal nerve that extends from the pulvinus (which is normally rather obscure) to the normally pungent apical mucro. Often a less obvious secondary nerve extends towards the gland from near the base of the pdncipal nerve. A gland occurs at the apex of an angle on the adaxial margin of the phyllode. That portion of the adaxial margin between the gland and the pulvinus is termed the adaxial proximal margin while the portion between tho gland and the apical mucro is termed the adaxial distal margin. The abaxial margin is the side below the principal nerve between the pulvinus and the mucro. 5. Inflorescences: Inflorescence charactem are very useful in distinguishing species within the Uninert,es-Triqngulares. The most impoftant character is the number of flowers per head, but other useful characters include peduncle length, bud shape and petal nervature. The majority of species have simple (i.e. not compound) inflorescences which are normally solitary at the nodes along the upper branches. Only the two closely related taxa, A. littorea and A. truncata, have racemose in- florescences. 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