Studiesin the genusAcacia (Mimosaceae)-8 A revisionof the Uninerves-Triangulares,in part (the tetramerousspecies) By B. R. Maslin

Abstract A taxonon,ic revision of parl of Series Urr'fte/yeris presented. Includedare those species with 4-merousflowers and triangular-shapedphyllodes. The seventeenspecies comprising this group are endemicto south-westWestern . Eight new speciesand one new variety are descrlbe A. delphina sp. !9y., ,4./i/tolea sp. nov. (syn. A. decipiensaluct.), A. phaeocalyxsp. no\., A, phlebopetalasp. nov., A. phlebopetalavar. pube:cens var. nov., A. pycnocephala sp. nov., ,4. tobinae sp. noy., A. Mmitrullata sp. nov. and A. Lliginosasp. lroy. A new name, A, chtl'stcephala,is gilen to the taxon A. biflora var. airca E, Pritzel. Three names previously in common use have been relegatedto syn- onymy'.A. cuneataBenth.and A. decipiens(C. Koen.) R.Bt. are no.\'/A. tunca,a (Burm.f.) Hort. ex Hoffmannseggwhile,4. l'errlcosaW. V. Fitzg. is A. i crossam Hook. Keys io speciesand varietiesare provided,all taxa are illustratedand their distributionmapped.

Introduction The presentrevision is basedon Bentham's 1864and 1875treatments of "UnineNes-Triangulares" the (see below) but treats only those specieswith 4-merousflowers. Thus only Western Australian ta\a are considered. The speciesincluded by Bentham but excluded here are A. acqnthocladaF. Muell., A. bidentata Benth., l. dilatata Benth. a\d. A. gunnii Benth. (syn A. yomeri- .formis A. Cunn. ex Benth.). Three speciesand two varieties described since 1875 are included here, viz. A. inops Maidet et Blakely, A. mooreena W. Y. Fitzg., A. yernicosaW. Y. Fitzg. (: A. inuassata Hook.), l. bifiora var. aurea E. Pritzel (: A. chrysocephsldnom. et stat. nov.) and A. horridula rar. hastul- stoides E. Prttzel ( A. uliginosasp. no\.). ln addition. eighr new specie' and one new variety are described. The terminologyused to describeplant communitiesis that presentedby Aplin (1976). This is a slightly modified versionof the systemproposed by Specht(1970).

. . All illustrations appearing in this work have been drawn from pressed herbanum specrmens. Only a selectionof specimensis cited under each taxon. A list of the numbered specimensseen is given at the end of this paper. Some of the un- numbered specimensseen are cited in the text.

Historical perspective

Benthan (1841) described lhe Triangularcs as one of the eleven major subdivisions of his SeriesPiyl/odineae and incltded specieswith both uninerved and plurinerved phyllodes. . Meisner (1844 and 1848).Bentham (1855) and Mueller (1859) later describednew taxa which were referred to the Trians;lare;. In 1864 Bentham reconsidered his earlier classification of Acaciq. 1n this work he raised the SeriesPhyllodineae to the rank of Division and at the same time divided the species previously included in the Triangulares among the 266 L9Z 'DSoutS!lt't pLvtoJolln.!l!Luas 'DlDLldaJouzId k, V V'oppr,tJotl t, zr^ sercadspelalel ,{lssolcJnoJ ur:ncco sepol1,{qd elEllnrlrules (surSreu I?rxeq" pue Jerxep"eql qtoq uo polSu€eje ,{eqtl"ql ldocxe etu"s oqt er" sepolJr(rldel?ltnrl) ^luo urSrtur eql uo pe13ur 'sseq letxepe 'e]surrunre putt oqt e^oqe scu"lsrp elrlos t? olpplru aq1 ,4\oleqlsepEoJq eJBqcrq^\ sepollfqd o1 perlddr sr al"llnllruros ruJs] oql 'uorlrppe uI (tf6l) '1, l, proJp€l{ ur ue,tr8 srurel oql pasn e^eq I olqrssod ererlrl6 sedeqs eseql eqtJJSopol {3olouru:a1 psqsrTq"lseSursn ut osuE ueUo serllllcurp'IJloorru^sP 'pecnpold srql Jo esneJeg lrrleurru,{se ,{te^ eje q.tg,r elllos ol? sod€r{s 'seprs Jo 8ur{re,r ,{1epru $rql oqt Jo elnls^Jnc pue ql8ue1e^rt?ler eql ur secue -reJIp ol enp lnq JBIn8ueulsr odrqs crseqoqJ sdrlolSqnsol1'. .^arDf8uo!.tJ -sa,t.lautuneql epr^rpol pesnsJalJe.r"{c eqlJo euo sr eclrqsepo11,{q4 l€drJuud 'V ( 1 arn8tg ur peleJlsnJlrer? ,{\oleq ue,rr8 surrel eq1 Jo IuBt\) :sapo llqJ 'lue8und ssol.lo orolu eq u?J sos"qluetsrs;ed eql aDulqorv pue Dlolddoqalqd tr uI lue8und ere ,{3111e.rerl,$ sua8ta,ttp y uetJo pu? xllwoaoqd y roj ldecxe sesEqpeue{crrll pu€ eBurul"l slrou€os e,\€rl selndllseseqJ seqcue.rq3uno,( aql uo sepoll,{qdtsoLu ss"q oql lp lueseJdsr 'ou!4dpp Jo selndrlsgo -rrrd z ,, solurleulosp\rB Da.totjq J roj tdecxg :safd!$ t 'sepo11,4qdsnorqrlS e,r.uq,{11ensn sarceds eqt 'D.to4[!q Jo :lsereqJ Jrlcco op slenpr^rpursno;q213 auos V ]oJ leurJou oJu sopoll,{qd,tureq q8noqllv sepoll,{qd,{rr"q ^lluelsrsuocqtr,{\ uox"t .(luo orll st su saqnd Je^ DpD(lo(plL!(l oDo)l/ fJrcr] ol snoJq€I8 uo.rJ ,t.r"A Jo freq s^B,ulereqlle e.re"xel reqto orll ur soqJuslq eqJ- xilDoaDqd ur punoJ osle 's€qcuerq f fll€tuJoLr€r? ,{eql q8noqllu snorqelS ,{lluolsrsuoJs1z\ Da,lottll V {lug (688 d '9161 urlse1,1),{lsnor^erd pessnrsrp oeaq s?q ereq pesn ,{3o1o -ulurJot runlueunpur eqJ "x"t et"rlueJelllp ol ,{lln3sseccns,fte,r pe,{olduraeq uEcSuue^or Jo ed,{1eq1 rou srr?qJo ecuesqEJo oJuasordeq} roqlreN sepol 1,(qd snorqelS pup soqJu"rq ,{req e,teq sercedseql Jo lsolN :unltuawnpLq Z 'tq8req ur ru g 7 Surqcre;'dno.r8 cql sreqLueu lssll"l oql ete Dw)untt pLtEoanlltl Dt)D)/ uotle]r?a^ Jo V 'sdou|v pelurcoss"eql ur polSu?luo,rol3 serurteuos Jo esorJtuuqt tsnqol erou Sureq q8noqqe 'qcrq,tr soqJuttq :lJalo qll,r sqnJrls uE Dsout8tln y pu? 'DlolnJso\ 'sercads Dloqd oLtJId V z porll? ee:qg uor1e1a8o,rpall?rcosse eql uI se^losureq)el?uelue puu re^o olqluBIJSqrrq,\\ serlJuElq luroJrlg '{Ee,r seq rdrr1.l o\cocy suoqdacxo euros olu eroql'Ja^oA\oH lq8roq ur tu seurleuos lo ru 'lcere 'pr8u 'llBtus I 9.0 €.0 sqnrqsltruue:ed ere ser.edseql Jo lsolN :tlqDH I {3o1oqd.ro111

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a bm

A. truncata- type A. bif lora - type g

Figure l. Principrl phyllode types of the Uninerves-Triangulares. A A. hoffi.lula-type (phyllode semitrultrte). B-A. billorc-type(phyllode obliquely obdeltate). C-A. tntncata- type (phyllode obtriangular). abm-abaxial margin; adm-adaxial distal margin; apm- adaxialproximal margin; g gland; m apicalmucro; p pulvinus;pn principalnerve; sn secondarynerve.

The phyllodes have a principal nerve that extends from the pulvinus (which is normally rather obscure) to the normally pungent apical mucro. Often a less obvious secondary nerve extends towards the gland from near the base of the pdncipal nerve. A gland occurs at the apex of an angle on the adaxial margin of the phyllode. That portion of the adaxial margin between the gland and the pulvinus is termed the adaxial proximal margin while the portion between tho gland and the apical mucro is termed the adaxial distal margin. The abaxial margin is the side below the principal nerve between the pulvinus and the mucro.

5. : charactem are very useful in distinguishing species within the Uninert,es-Triqngulares. The most impoftant character is the number of flowers per head, but other useful characters include peduncle length, bud shape and petal nervature. The majority of species have simple (i.e. not compound) inflorescences which are normally solitary at the nodes along the upper branches. Only the two closely related taxa, A. littorea and A. truncata, have racemose in- florescences. Here the raceme axis is very reduced. 692 'sdnoJ8 lnoJ ur peur€luocen safip&uo!,tJ-sauauunaW Sursr-rdLuors:rc:ds ua:tu:nrs aLlI .a ernBrJur (;lerrieuu_Lei8erppjruiserii eJEsdrLlsuorlpliJ JgrJedsJ:lul ,{qsarpnts 3ur;paor petrurl 'qd Jles,{Lu pueleutnD Aq-sJlpnls -uJllod ru_o4pJ^rJep ecuepl/\e Surlroddns qlr,r\ ,{SoJorldJo[u e^rlereoujoJuo AJI?dlJuudpeqsrtq"lse eJ? sorlruuE sorcods .dnoJ8cttrlouoxel IBJnlEu" 1ou flurcgec or" eJeqpeugep se sioln8uutj-saatauun e\II saraF8uBlrl-saraulun oql ulqlpi sdlqsuouBlau

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A. biflora; A. robinae;A mooreana; A-horridula; A. hastulata; A. chrysocephala;a. phlebopetala A. semitrullata;A. uliginosa; A. divergen s; A- incrassata A. pycnocephala; a. inops

A.truncata; A. littorea

ACACIATRUNCATA GROUP

Figure 2. Affinities t1t the lJninetves-Triargrrldfes.Species blocked are closelyrelated; a solid line indicatesprobable close relationship; a brokenline indicatesuncertain alinity \, group Included species:-A. hqstulatq, '4, horridula" A. inops, A. pycnocephala, A. ltemit tllcttq and A. uliginosa. (On account of its legume morphology, ,4. plrueocalyx (seegroup 4 below) has some alinities here, but this speciesdiffers markedly in its vegetativeand floral morphology.) The A. horridulagroup is a distinctiveone and its membersare not closely related to any of the other speciesconsidered in this levisior. The unifying charactersof the speciesin this group are mainly those of the phyllodesand legumes. The phyllodes are frequently semitrullateor sometimesvery nar- rowly triangular or asymmetricallytrullate, they are broadestbelow the middle and taper into sharp apices,they rare]y exceed l0 nm in length and their principal nervesare more or lesscentrally situated. The legumes(not known for A. inops)are very distinctivein being terete, tapered at both ends, red- brown jn colour and longitudinally striate. Seedlingstudies, althottgh of a prelinrinarynature, re\eal the followirrgtrends in the sequenceof develop- inent 1'eedlingsrol known lor 4. int'p: and 4. rrcno,(pltala): first leaI alwayssolitary and pinnatewith 2 pails ofpinnules; next I 3 (5) lcavesbipinnate with one pair of pinnae; subsequentleaves reduced to phyllodes. Po)len stndiesby Ph. Guinet (pers.conrm.) confitm the uniformity of the A. horridtrla group as definedhele (N.8. pollen of .4. inopswas not studied). Within the A. horridula group, the speciesL horridtrlq, A. pycnocephala, A, uliginosa are nrore closely related to one another than A. semitrullatq and 'fhis to A. hastulatq and A. inops. first subgroup forms an intefielated species- complex the members of which are rather similar vegetativelybut differ in floral morphologyand distlibutiorr. It could be arguedthat thesetaxa should be treated lrs subspeciesof the one variable species,A. horridulq. 2. Acqciq biflora group Included species: A. biflora, A. chrysocephala,A. dit'ergens,A. incrusstla, A. mooteqnq, A, phlebopetala and A. robinae. The membersof this group are united priocipally by their phyllode and legume morphology. Acacia ditergens and A. incrassata both possesssome characters which render them a little atypical for the A. bifloro group. How- ever, consideredon a broad basis,it seemsbest to include thesetwo species here rather thar with lbe A. lruncate group to which they independently show someamnitY. - (r'BrJ) sdoul'V 'Z (lJulsrprr^rd 'relnSuprrl lelei -rel l) epr^{u,Illl I-S.0'poxouep ,{llurututord f,|^\or]eu= sepolllqd .q (€ 8rd) €l€lnlsaq'V 'I .. uo 'luetEd'el€llnjl oquad l"eu ol fueqlv) opr^r ruur (S.t) t Z ,{lle.uleu!u,{se s6poll,{qd .et (-tt pexauap I 'atellnrtrllles > ) ,{LlqSrtsro lueled .{l^\orr€uol Jeln8u"ul ,{1,^\orrEusapolL,{qd :,(rreq saLrunp.d Surre \ol{ .q n Cgt <) pexogap xllueuruord dO ot"lln.It .{ll€Ju1 uru,{s€sepoll,{qd :snorqelS solcunpedSsire^\olJ .e€ - € p€eq rod 8l uEql sselsraAlog :e^oqD sD tou edEqsepoll,(qd :orinul e Iq peddoUe^olou puElC .q (61 3ll) uulqdpp'v 'rI (,{eBelrleErsl ot ro^ru dnurllpd:lseoc qtnos) pEeqrad Z€-gl srr^\ou:utBleurlEtxepe uo:nds ielniuei:l snon.Idsuoae ql!^\ podeqs-luorsari+ sopolllqd :olrnur luo8ind,{[]qBr1spire (tutu .eZ 8.0-9.0) lroqs e .{q peddotre^o(epoll^qd Jo ur8reurlprxDpD SuoLp prlenlrs) puelC " 8 snouE^ edEqsopoll,tqd :olpprrr rroql e^oq€ lsepeoJqsopol - , .q l.(qd ueqt (etet) lertuea T Jr UO ruluoaxe Xlsnor^qosopoll,{qd Jo a res lecb.uud Z elpprur(Je0u ro) ,'^oleqlsepeotq lue8undpuEaleur[rncesopoll,{qd:polen|s.{llErtu"]+sopoll,{qdJoo^reuledbuuj.el selcads o1 ,{ay

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Flowers consistenlly4 per head ...... 6 b. Flowersmore than4 per head ...... 7 6a. Flower heads golden yellow and + sessile(peduncles l 5-3 nln long); seeds minutely roughened;legumes to 40 (45) mnr long; phyllodes (l .5) 2,4 mm wide. (Narrogin-Kojonup to Lake King-Ravensthorpe) .... 4. A. pycnocephala(Fig. 6) b. Flower heads very pale yellow and distinctly pedunculate(pedrrncles 5-10 mm long); se€dssmooth; legumesto 75 mm long; phyllodes1.5 2.5 mm wide. (Maddington to Serpentine) .... 3. A. horridula(Fig. 5) 7a. Stipulesrather spreading;petals hairy (sometinressparsely) and 1-nerved;flowering peduncles 3-8 mrn long; flower heads globular with 5-8 flowers. (Busselton- Donnybrook to Harvey) .... 5. A. semitrullata(Fig. 7) b. Stipules distinctly ascending;petals glabrous and nerveless;flowering peduncles 1 3 mm long; flower headsgloblla. to slightly obloid with 8 15 ffowers; Iegumes glabrousor glabrcscent. (Nea( Albany to near Augusta) .... 6. A. uliginosa(Fig. 8) 8a. Flowerc2 per head b. Flowers 4 or more per head .... I I 9a. Petals3 7-nerved*; peduncles(2) 3 8 mm long; flower headswhite (normally drying orange-brown);phyllodes hairy or sometimesglabrous; aril not shiny. (Albany to near Esperance) ...... 7. A. biflora (Fig.9) b. Petals l-nerved; peduncles0.5-2 (4) mm long; phyllodesnormally glabrous .. 10 10a. Flower headsgolden yellow;mature phyllodesfrequently barely pungent,distinctly ascendingAND/OR with a discerniblesecondary Detve in addition to principal nerve; legumeshairy or sometimesglabrous; afil slightly shiny but not waxy looking. (Nanogin-Mount Barker to near Esperaflce) ...... 12. A. chrysocephala(Fig. 14) b. Flower heads + white (oormally drying orange-brown); nature phyllodes very pungent,patent, secondarynerve absent; legumes glabrous; aril obviouslyshiny and waxy looking. (Near Albany to Mount Manypeaks) 8. A. robinaa(Fjg. 10) lla. Flowers 4-6 per head .... 12 b. Flowers more than 6 per head ...... 16 l2a. Petals 3 ?-nerved*; flower heads creamy white; peduncles7 12 mm long; upper branchesmoderately to densely(often minutely) h;iry and rather obscurelyirervi:d; phyllodes5-10 mm longand 5-ll ll8) mm !'vide. (Southcoast: patlinutRiver to Munglinup) 9. A. phtebopetala(Fig. lt) b. Petals1-nerved (nerve sometimes obscure); othea characters not conbined as above 13 13a. Flowers4 per head;peduncles I 3 mm long ...... 14 b. Flowersmore than 4 per head;peduncles 3.5-10 mm long ...... 15 14a. Calyx.lobes_a_nd bracteoles prominently acunrinate; flower headspale yellow; branchletsglabrous to sparsely(rarely densely)hairy. (Near Bunbury ro near Augusta) .... 11, A. mooreana(Fig. 13) b. Calyx lobes and bracteolesnot acuminate; flower heads bright Bolden 'yellow. Theseare 4-floweredvariants of a specieswith normally 2 floweis pe; head. (Nar- rogin-Cranbrook to near Ravensthorpe) ...... 12. A. chrysocephali(Fig. 14) l5a. Phyllodesbroadest below the middle and taperedinto acuminate,straighr, pungent apices:flower headscream ro pale yello$: shrub lo 2 j m Lall. rWaJdUin!to Augusta and Stirliog Range) 10. A. divergens(Fig. 12) b. Phyllodesbroadest above the middle, + obliquely truncate at apex: flower heads goldenyellowl duarfshrub 20 J0 cm tall. tNew Norcia(o Serpentine) 13. A. incrassafa(Fig. 15) 16a. Flower headsobloid with 18 32 flowersj gland (on angle along adaxial nargin of phyllode) overtopped by a short. slightly pungent nuiro. (South coast: Mount ^/ra\weltro $raelle $ay) 17. A, d€lphina(Fig. 19) b. Flower headsglobular with 5-16 flowers; gland not overtoppedby a mucro .... l7 17a. Phyllodesbroadesl below (or near) middle and acuminarewirh st.aight- and very pungentapices; stipules frequently pungent; brancbtets finely ribbed ...... 18 b. Phyllod€sbroadest above the middle and with + obliquely iruncate apices;apical mucro.leispungent than abo\el sripules{$hen presenilnor tor scarceiy)' pungenr; branchletsnormally coarsely ribbed ...... lq + Observeapex of petal when dry at not less than x 10 magnification;midrib normally morepromtnent than flanktngner!e5. 272 tLz 'prosdrtle. 'ounSol llq8rls ot prolqo uI pulpntl3uol spaas peuefclrltrtrou 21la& : oleu$,(1purpn1r8uo1'u,r,Lolq-pel'lu$selqEl8'snoec?I.tocfpq311s o1 lDalg,tow 'pa^lnc '3PI^\ '8uol snoeJ? 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Z '8uol ta ol : f.tBlrlos poJq &f)unPad :snoJq3lS tuu.l 7 salcunpatt /rsrg 'epou tr :sepollrqd Surpeecx3^pqSJls to uBqt.relJoqs ;ed I'eldw$ s aaJSaJoLluI 'lueu 'opoll.(qdJoelSu? uo pel"nlrs luord I ou puo19 3uo1wul2.0 I.0 l?Ixef"'.tlst,./'ollei '(,{.rp 'pol?nlls 'pacnper ,(ro^ snu^lnd uorJ,,!\)pesrBJ + f,IleJJueca'uau 'lq8rells'8uol 'luo8ond lotlpuu,[:ot o]g 1Ll3rL ruut I €J (e,treultdrcuud 3ut1e -ut||rat) or)nu prlrlz :101e;npun,(11qBrts soqol) peqol ,{ll?nbeun es€q '(,{rp ueq,{.) ueer8 lsepeoJq1" ePt/\{ IrBp'sno:qe13,{11uru.rou'polseSuoc'pt8u'lual"d'lulod'IrlroJlproo ulur (9.e) €-Z'8uol rutu 9-q.t 1sout1e,{llzuorsttcco:oe1e11nr1 f11u -cr.rleruur,{sesapo tLld u^\oJq 'elelllc ro snorqtlS 'pe,unc ,{11q3t1s'SurpueJst 'lue8und 'es"q 'luelslsled'3uo1 1ou ]€ peue{clql,{l}q8rls }nq snoltecs ruur g-9. 1 'snoec?les uauo 'r"ln8uBul ,{l,to.r:eu ,{:e,L ,laPdrls so,l:eu uee,tleq ueeJ8 ^lI"rurou '(snoJn.reqnd,(1asro-r1ue fJIeuorsBcJo) osolld- oqs ,(lel"JepouI ot '(pesrer ',{\olle,{ 'eleJel Xles:eds lou se,treu) pe,rreu,{lelncsqo sidH)uolq :u,r'orq -eauero oJ LlstppeJsaLpuDtq :spJ?,{\u,^\opSulucl€ uelJo olrnb pu? papl^IP ,{Ies;edsuelgo seqcuBlq pettrcoss? 3uoru" pelSu"lue 'll€1 l€uIurtol'gon"la3o,\ 'SuIpEeJds se[rr]euos ru z 01 qn ls eleJrJlur'{"4,\\ l3q1€J o1 lcelE 'pnu '(9r8r) 'q.,"1 'uun*..s rllou 6r:9n81u,ry'A seD-nequorrec il'4fi::l !t;it":;;"::l 'DtDln$Dq qns u,{sord '.. €t€proc (098t) d ^o{esrlv V'(0t81) rz'pouoH '-pnalsV,. 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BR.Anstw1177.

Ftgurc 3. . A Portion of branch. B Flower. C Bracteole. D Inflorescenceshowing pedunclesubtended by a solitary basal bract (b). E Node showing \tipu.lesG) and patenl..aslmmerrica_llytrullare phllloda (gland posi(jon arrowed). f Seed: u. Legume{terele anO tongtludtnatty stflale), A, E from C. A. cardner 3323;B Df.omA. S. Ceorge2639;F-Gfrom A. M. Ashby4265.

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'\t0oa! v ldecxe dnorS eql Jo slequlotuloqlo IIE ul snolnraqnd)solcunped snolqEIS sll st ololnwoq y SittstuSocoruI InJesnJelJeJerlo leqlouY sepo11,{qde1t11nr1 slr ,{q dnor8 srql sJeqllleluloqlo eql uo{ peqslnbullslp ,(ll?f,uteullufse Jo 'dnor8 xjipuer sr onfJsDq onDrY Dqplltoq y setcods ql!.lr .{ll€lcedse _eql '3^Ilcullsrp ojiputotl p'5qt o1 dlqsuoqtler e s1se8?ns,{3o1oqd:o'u s}r ^re^ -s! itthwsoit ue1lodeqt alrq,lr l"ql pouoder oq,,lr( ruuoc sred) lourng rld fq v 1o 'V payoddns sr Llorsnl)uoJslt{I (0/Z d) dnot8 opp!t.toq eql uttllll olDlntrsDLl ;y 'erun33l :re;d s.rol.rcreqropoll{L1d JEleueBpu" ocuecse.IoUur ogJ (dls ',\N "131,'\l'-I '0t8t 't 'D{H'I3OD'D) 696 oN rtard qreH tdes IZ p (lsur8ellLPld)..erreJs.Suruqs '-l3l.^{\ pldgPlo punosenrorD nul) :lHIuSdl pe oe.eJrnl osouoreolos ul :(HIuaI *eI ,7 11 ') 'I^g) 6t9a 'S / 'ellltcqtroNlo S ltu 9 lnoqv :(HItSd tztt lauryDt t' J ra^Id )retuuoc :(..plorr\lfsesorurl ,,rtueu sur s.u\\org sreeq looqs sltll 80tt roqlunuuollnqrl]srp lleuueg 'l,,lgl 'lll ISrorD sr8!u Lunrrodedord ^\olC edeJ rnoq&H sso.ulra n i r"r.,g U 'ear!. '.{eqrofle le,{ou .rijur sr,rlrsuI :(HIU!la) S9& Iq\sV t! V .{ueqlv rEeN :(HIIad) 0t9I tqtfiy:ri'yjrcdtteiutqlytolsensepuneJV:v!'lvxrsnYN{rrsel6:statuDadsPaPalaS ',{r"nu3f pu? lequleJo(I -prcu ur pelcelloc ueeq el"q spees eJnl€ur q1t,tr serun8el :pottad 8w1tn.tg 'fuBnrqel pu€ requsJoc uee,^d.l3qpalcalloJ,^^eJ E ele e.I3rJllnq loqlue,\oN 01_,{ln{ 3l"l iuor; pe.reqltS eJe^\ pu?q lE suorurJedsSuue^{og lso1N :po ad 8ultalrolf 'Dlotlt,!o upuDlJ puu suolntusol oayotcy 'DllolJoanl '3y 'sdaJl,t.todstuoBv 'snsot.tD)ssndtotoldal qll^\ pelslcoss" qnJcs {crqt uf uauo seJeld fdlu€,lrs ur Pue sleeJc .reau u"ol to puES :lPllqDH '(uolroqued Jo tse^\ tu{ 0t) re,ttX ,{1leuroq eql ol lso^\rJuou-lso,^\,{uEqlV tuo.U suorEe.ruJ3qlnoS :"IIetlsnV ulelse/d lse,{\-qlnoS(lZ eJnBIJ) :uottnE.tlst(I '8uol ru(u I Bc ol qslllq^\ lo lu?eJc 'poploJ-€Juo,{qseg e olut,{lldn:qe ssel //r, lrreJnlsuerl+ 'tuJoJrlg pepuedxe 'sarnletll pees s" Suuol{ll.'!\'3uo1 (sse1lo) ultu .ro eroru E.0 'eug aprun! ',aprN rutu g .0 pu" 3uo1 ruut 'BJ apafi :unIIIq eql spre,"ro1uado 'PIStnl7 'apl\\ '3uol ruotSotnaltl: ^uII{s Jeqt€r'u,{\oJq ruul 9.t-l tuul S.t 9.2 peduncles.3.5 5 mrn long. .rery slender, glabrous; basal peduncularbract sollrary,-crrrorate;y'ov,er heqds cream to white, globular, with 5_g flowers,ca. just.prior i prn.9i1r. to anthesis. Brecteoiei slssile,'0.5 rrm long (equal rn rengrIr ro catyx)J ovate to obtolg, acuminate, clliolate. Flowers 4_ merous; bud\ attenuared; calyx )-1. Jengrhof corolla.dirided for .l .j its lengthinro coarsefycltlotate triangularlo6es: pctal\ 1.2_1.5 mm '/s .1. long. connatefor Tnerrlengrn. gtabrous or someltme\rery sparselystrigose. very obscurely l-nerved: olary ses\ile.slahrou\ lsparsely!apillose in riry your1glegumes)". Mature legunis and seedi not seen,

, 5O mm

BR llnsuu 1177. -Lpp:r Figute 4. Acaciainopt. A parr of planl showingweak, filiform branches. B- r\r and cteuexed. narrowtylriangutar ph) ode bearinga basal granc-oeafli,.9:::i-olliCng spur:!igy]es (arrowed). All from R. D. Royce 3913. 276 -1-qolaa ,(lueuruold !! 7 :snolqel?_oqnt ,{Fulncrlred1ou se,lrou) pe,rreu-g re;n8ueu; :ur .eiloLor 1o.19l9tnt9t1" ^lpporqolur qrBu:1irr'{. .,o1peprrrp jo qldueTi r.//aJ:pe)L'nuellE ({l)q8rl\ .ro) tou 5pnq:snolaLu-tr s.r,',ao7jr .sno.rqe;B 'elelorlrc'e^€Juoc ,(x,{lBc :ltfr"qto'elt\\e. u?qt re oqs) Buol ,uu, 1.g g.g,e1i^o szpat.)o.tB.:lJ^\ou qlr,r .q\Jqtue .ueip"Lutu .", -relnqof t or ioud t"nf g.1 n "rollJA eled spDdrl.tJtvol{.. lre]|o. tJDtq .lDrJunpad psDq : rn.u u r (lqereprsuoc .o.ro.rrue I uJq,^^ Suruolrrqr or ruiied s.rreqt.nolniaqndfljsujp br r1:ru)a -pout '8uol 'aidwrs .sndr,tlnd trtru 0l 9 rapunpad lepou -rod 1 satiatsiogiuJ' Luut,I Z 9.l epo[,{qd urS.rELu uoa13ue lq8rlie uo pe}en}rs l !9u .8uo[ Jo ;erxepe,lcur]srp toLrpuDtC LuuI S. 0-€. 0 snwtlnd i(itp uaqn) pesrd.r 'epollAqd.lillly"jd .peienlrd Jo rspq .rEeuurBrELu lerxpq€ spr?,rol po^rnc,{llq8rp file.rlriec a-ttay '8uol 'lua8uird tq8l€rt: utu-r 1 e5 (e,treu ierircur.rdSurteiirruiet) o.t)nwryd1.tu1ttI.: 'tq8rcrls 'pelsoSuoc'pr8ri ,lue1ridiluiod lDrtdD :snorqqa lsepzorq 1r Jpj.^\ '8uot urur g-9 ,el€llnJl[uas ,uwoJq 'snorqzl8_LutuS.a-9.1 '(qJuerq Xl,rorreu sapo ltld lq;ll qlr^\ .97 rc go ol8ue uri Aurpeecxe1'ou)..aurpee:Oi ,lue8und ,luelsrsred l"rl,{\etllosol burpuocse'esEq1? peue)crqt,{pq8rls lou ,Buo1 Lutu t 9.(':noJJElJ( ol Jeln8uptJt,{;nolleu(-r.:r s4r?d//S .u{olq qstppJJ '1:s-rorrue ol u$orq tq8rl (pleLu.rou ot ruored ".rruqt.irolni:qnd fla.irep bi xg:re -Japour'p.)\Jeu .eEJet ,{lJJnJLqo satpuDq:(pJlJoq$ ,{luouruo:pue pJpr\rp AlJcrn0s'burpuJJse s1r_.rrrep) | .rJddn ur peqJueJqqJnul rnq rueld 1o I tomijl ur porlcu"rq .{lesreds .pau[ue1s-e i11it ,{11erurou l8urs ru 11.19.gb1 gnr,15

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Selected specimens: WrsrERN AUSTRALTA: Narrogin, Aug. 1925. C. A. Gsfttner s.n. (PERTH)j South S^ttlingsRd,36 mi(5J s_knrJE ofBorden rd, ,1 S. C..,Je b253 (PERTH); l1-Ti.(15_lgrllS of Kojonuptowards Rocky cull), B. R. Mastin617 |CANB, pEirTH) and 3992(PERTH); ;l mi (0.5km) N of Highbur](berseen Wasin and Narroein).8. R. ,4i.as1rir 658(NSw; PERTH); 6 km N of Mounr Mcddenrowards Lake Kina. E. R. Ma"rtin4063 (P[RTH):l0km NN\ ofMounlLroper, /(. Vc!h,l,4lco{PhR tHJ. Meisner (1844)based the name A. horridulqon two collectionsviz. preiss 965 and Preissll5l. The former collectionrepresents A. pt,cnocenhalawbile the latleris A. ho-ridula\en5u lecrol)pico (.ee p. 2Zgt.

-. . Aca.ci,qpy,cno.cephala is placed in the A. horriLlulq group (p. 210) and is distinguishedby its golden yellow flower headslcrearn or whjte in tire other species)and its minutely rugose to r,erruculoseseeds {snlooth in the other species). Other charactersuseful itr recognisingA. pyinocephalainclude its short,densely puberulous peduncJes. its consistently4-frowered flowcr heads,its smooth, glabrous and nervelesspetals and its bl.ancheswhich are deniely white-puberulous towards their apices. Its general phyllode and flowei morphology render ,4. pycnocephali closer to 7. horriJiti than to the other specieswithin the A. horritlula group.

^ According to_Ken Newbey (pers. comn.) A. pycnocephalaregenerares after fire from suckers; seedlings are rarely seen. The specific ephithet refers to the inllorescences which are rLnrcrous and congestedtowards the extremitiesof the br.alchlets.

5. Acacia semitrullataMaslin sp. nov.-Figure 7. [Acaciahorrit]ulu auct. non Meisn.:Bentham,_Flora Austral. 2:350 (1g64) pro parre,as Io SwanRiver, Drummondand, Haryey F.i.ver,Oldfield.l . Ftut?x^+,debilis.0I I t1 5) n1altus; ramuli spa:sim ad denseantrorse puberuli ad slfrgosr.J//p///i./r per\t\tenles. parente\.PrJ //, d/ir plerunrqueangLrste Gmrtrullcta, rnterdu tflangul:lfra longd.lJ-2 n1n1lala. patenlia rel detlera, .5 l9'l'" pc Ilerr_rrnoue pungentra. ner\o pflncrpdtt ad centfunt por,1o. tttcuti J 8 mnr longi rtnodice, oen\e pubertrlt; capitula globulosa, 5 8 floribus. f/u,.J 4-ri,eri, callcis lobi triarrgulares; pel4to l 5lrln longa. modice rjparsimr pubefuld. t-nertia. Lepinina rereria. a--do0 rnnr , jtnala. 1-51 longa. 2-J mm lala. cur\a. Iladra. 5.rrrlr?din legLrmineIo|ltsiludinalia, obloiCeir aJ parum ellipcoider.4lltnt long:l, 2 mm lara; drl///,r conicua. ,,Shrub Type: BetweenHarvey and Myalup, Western Australia. to 0.3 m tall;,nornally gJowing erect anrong associatedvegetation; branches slender; ll:ads cr-eam..,Qark. grey lqrdf l_oapin semi-swamparca.,, 20 Aug. 1976, B. R. Mqslin 4193 (holo: PERTH; iso : CAN B, K, MEL). Rather weak, erccl shrub0.3-l (l ' 5) m tall, not scrambling,eirher single- stemmedor dividing at ground level in_toa number of slendei orange_brdwn branches; branchletsterele, finely nerved (nerves yellowish), sparsely t-odensely 282 €82 '(Hlddd) 'tcrrtsrp us rrry&/rl ) y'Z06I ,{rnqung: (tuetuBE4-HJUAd 'tlleqtuog^q uoas-'lft1{) us pt2{pto 're^ru ,{e^reH:(.]gr{) us ptog)W 'stw 'tL9l 're^ru poo.4^Ic"lg:(HJJEi) ullsDt1tr g rp"oUsoFC uo uollossng tu)] r]noqv 96It 'ladeJU. 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Dlnpt,tlol V egl Lr paaz(l st DtDlltl.lJ!LudsDFn)y 'Irr€J JgFedsluL'JrE,\\ ol rlSrrorro1.rurlsrp lr JeprsrioJI luosard eql JoJ'Dfpt,t,toLl / Jo rroxel cgrcedsr.r.;rirue se peleeJl )seq sr Dto n.tl -tLuas y2ql etEcrpur .{Eur ,{pn1s a:n1ng q8noqttv plegplg 're,r.r1 ,{e,trrg V 're^rx 'zt| prle puorurrrnJc uB,r DlDllut]atas / 0r? qcrq,{\ o^\l suollcolloa 'tqgt) S 'aldru"xeJo rnoJ pelrc (OSg d rLtgLlt\reg'Dlnp!.t.toqf'repun roC lsreltr Dfp!.troq V' qlrM pasnJuoJ ueeq uJfo serl nlolln.t tu)\' t, lsEd eql uI S,4rabeither scranbling (with long slender branchesto I m) or erect (30 60 cm tall) with normally 3-6 (sonetimesrnore) thin stemsarising from a subtertanean root stock; brqnchlets terete, finely nerved (nerves barely raised above branch surface,yellow), nornrally sparselyantrorsely puberulous with hairs restrictedto nelves (occasionallydensely puberulous with hairs not re- stricted to nerves),greer. Srirrlr,s very narowly triangular 1o sctaceous, 1 5-2 5 mtn long, persistert,scarious (but slightly thickened,especially near base),not pungent,distinctly ascendirlg, + straight,glabrous, brown. Phlllodes normally vely narrowly triangular(but baseunequal more roundedon adaxial margin than abaxial nargin), sometimesvery narrowly semitrullate,5 9 mm long, 1-l 5 mm wide at broadestpoint, + patent, rigid, frequently rather distant, straight to slightly curved, glabrous, medium green; apical mucro (termirating principal nerve) pungent, ca. 1 mm long, straight, light brown; principal nert,ecentrally situated and sJightlycurved towards abaxial margin nearphyllode base, sometimes raised (when dry);palunas very reduced. Gland not proninent, situatedon adaxial nrargin of phyllode (sometimeson a very slight angle)0.5 t mm abovethe pulvinus. Inf.orescencess\mple, I per node; pedunclesI 3 mm long (to 5 mm when in fruit), puberuJous;basal peduncular 6racl solitaryand cilidate1'llower heads pale yellow to crearn,globular to slighlly obloid, with 8 15 flowers,ca. 3 mm diam. just prior to anthesis. Brqcteolas more or lessseJsile, + I mm long (exceedingcalyx in length);laminae concave, acuminate,ciliolate. F/o\L,er-,r4-nrerous; buds attenuated;rallr + to almost ] length of corolla, divided for * its length into triangular ciliolatelobes, tube glabrous and nerveless;petals 11.5 mm long, glabrous, nerveless;r,rary denselytomentose. Legumesterete, to 70 mm long and 2 mm wide, curved, taperedat both ends,slightly contractedbetween seeds, firmly chartaceousto somewhat coriaceous, glabrous or glabrescant,red-brown, longitudinally sftrale: nlarginql nerle nol thickened. See.1slongitudinal in legume, obloid to slightlyellipsoid,3 4 mnr long, ca.2 mm wide, turgid, r'nediumbrown, slightly shiny; pleurograntopen towards the hilum, situated near periphery of seedon each face: areole2.5 3 x I 1.3 mm;. funicle minute (ca. 0 5 mm long), withering as seedmatures, expanded irto a once or twice-foldedconical slightly shiny pale yellow thickenedqril to 2 5 mrr long. Disttibutiotl: (Figure 20) South-westWesterr Australia: Near Witchclifle south to near Augusta then southeastthrough Northcliffe to Marbellup (about 15 km west of Albany). Habitqt: Seasonallyswampy corrditiorrsplevail ilr most places where this speciesgrows. Betwcerl WitchclilTeand Augusta it has been colleqtedfrorl shallow depressionsin open-forestdominated by Eucab)ptusmarginota and E. calophl,llo. The soil hereis sand,sandy loam or lateriticloam. To the south- east (betweeo Northcliffe and Marbellup) A. uliginosa occurs in loam in better defined,seasonally swampy areas. Here someofthe associatedspecies include Aclinodiunt cunninghantii, Agonis parriceps, ,lnqrthria scabru and Etandra ar$Iata. Flor)eri g pe od: August-October. Fruiting period: It takes about 14 weeks from althesis for nature seeds to develoo:these have beencoliected in mid-December.

Figure 8. ,-4ca./d liginosa. A,-Upper part of branch- B to C-Phyllodes showing shape and sizevariation (gland position arrowed). H Node showingascending slipules (s) and patenl, senritr.ullatephyllode (gland position arrowed). I-Flower. J Bracreole. K- Seed. L-Legume (tereteand longitudinally striaie). A, C and I lrom B. R. Maslin 3778(the type);B from B. R. Maslin 474; D from E. M. Scrymgeour1198; E fron B. R. Maslin2880; F from R. T. Lange257; G-H, J fronl B. R. Maslin3785: K L from B. R. Maslin4018. LgZ 'LLblNtlsaw u I I 1V\\ p;'1li :il \-a/ .----trEZ--"-

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.-=----_-\__=______=-8 -- -- -r.u ul--08- Selecledspecimens: W$rERN AusrRALrA: 3 mi (4 8 km) N of Northcliffe,R. T. Lange 257(PERTH); 187 nri pegon MargaretRiver-Augusta road, B. R. Moslin414 (CAN B, NSW, PERTH); Near Augusta,B. R. Maslin 2880(PERTH); 2 km N of Northcliffetowards MaDjimup,-n. R. Md:lin 3785(AD, BRr, G, K, PERTH,RSA); Marbellup,ca. 15km W of Aibany towardsDenmark, B. R. Maslln4018 (PERTH); 2l n]i (33 5 km) W of Walpole on road to Northcliffe,L. M. Scttngeour1198 (PERTH). Acqcia horridula vat. hqstulqtoidesE. Pritzel is treated here as a taxonomic synonym of .4. uliginosa. Although Pritzel's description is very brief it seems to fit my new speciesquite well. It is noted that the types of both laxa were collected from the same locality, viz. Marbellup. It is doubtful that the type of var. lustulatoides is extant for it is neither at the BotanischesMuseum, Berlin (B) ror among the many fragmentsof Diels and Pritzel specimensheld at the WesternAustralian Herbarium (PERTH). Acacia uliginosa is placed in the A. horridula group (p. 270) and is most closely related to A. semitrullata (seep. 285 for details). This speciesis recog- nisedby a combinationof the following characters:phyllodes patent, normally very narrowly triangular, rounded (or sometimesinsignificantly angled) on the adaxial margin 0 5-l mm above the pulvinus; flowering pedunclesI 3 mm long; flower headsglobular to slightly obloid (alwaysstrictly globular in the other species)with 8 15, pale yellow to cream flowers; petals glabrous and nerveless. Other characters useful in recognising A. uliginosa include its normalJysparsely puberulous branchlets (hairs normally restrictedto branch- let ribs), its relatively small flowers with narrow calyx lobes and its often diffuse,sprawling habit. This speciesis of variable habit. Under definite swanpy conditions (e.g. Nolthcliffe-Marbellup) it is a sprawling shrub with weak and slender brancheswbich scrambleover (and through) the associateddense vegetation, Fufther nofth (e.g. Witchcliffe-Augusta) where less obviously swampy con- ditions prevail and where the associatedvegetation is not as thick, A. ttliginosa grows as an erect shrub with shorter, more rigid brancheswhich do not scramble over the associated species. Irrtermediate forms have been observed. The specific epithet alludes to the species'preference for swampy habitats.

7. Acacia biflora R.Br. in Ait. f., Hort. Kew. ed.2,5:463(1813)-Figure 9. "Mimosa Lectotype: scalenaBay I South Coastin coli . . . . apric: Jany. l802" (BM upper right-hand specimenon sheetbearing Bennettdistribution oum- ber 4309: iso: K). lecto. nov. Mimosa biflora(R.Br.) Poir., Encycl.Meth. (Bot.) Suppl.5:530 (1817) basedon,.lcQcra bifloft R.Br. (1837); Acacia triangulatis Benth.'f:tpe: in Endl. et al., En\n\. . Hueg. 42 Bentham, Flora Altstral.2:351 (1864). Australiit,-Bdu?r s.n., ex Herb- Vind. 1837(holo: K; iso: W). Shrub 0'3-0 6 (l) nr tall, either oper and single-stemmed(or sparsely branched at base) with spreading-erectbranches, or dense round and intricate (forming clumps to l'5 m diam.) and much branchedat basef new shoots light green but tinged red at apex; branchesgreyrshl' branchlets tercte, obscurely nerved, sparselyto denselypuberulous to shortly pilose (indumentum variable- hairs retrorse to antrorse). Sfipales narrowly triangular to very nalrowly triangular, I 2'5 (3) mm long, scarious but thickened at base (basespersistent after phyllodes have fallen, scarcely pungent), glabrous or ciliolate, straight or rarely slightly recurved, dark brown. Phyllodes obliquely obdeltate to obliquely obtriangular, prominently angled on adaxial margin,3 7 (8) mnr long (as measuredalong abaxialmargin), 3 8 (13) mm wide (asmeasured along adaxial distal margin), slightly undulate, nornally slightly ascending (as in- dicated by angle of principal nerve with branch), slightly thickened, moderately puberulous or shortly pilose (hairs + patent, sometimes tubercule-based), 6lJZ rul : (Hldad) allsDl,\t'V g'eJuErodsEspre,rol qctBd sserDJo Lul :(HIUEd) 'us 9 t '9681/0g'^oNZ '.{ueqlv ''IghI ') 'htg)S 9.61't sntaq tr : (lueru8e{ HIUEd € ou llo. pue (HIUgd ,-IE',{ ,) ,t,ta) ou (t puotutun.te'r.^rt qpA\S:(HL\AA) zrDaa1 '8od 'peor Z |o. 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, 3mm I l(Jl l= l= ,11]\f I litilt/fi tH It tl'ttl ls \]F I Bfl Ansttu 1177 FigtLte9. Acacla biflora. A-Upper part of branch. Btoc Phyllodesshowing shape, size and indumentum variability; gland position arrowed. H-Node with insert stlowing gland (s : stipule). I-Inflorescence (b : bracteole). J-Petal showing 5 nerves. K- Seed. l-Legume with broad outer margin and narrow inner margin. A, I J frorn B. R. Maslin 3524A; B from B. R. Maslio 2507; C from J. Drun'rmond2; D from R. Helmss.n.; E from K. Newbey1328j F from B. R. Maslin2589; C from H. Demarz3639; H fron B. R. Maslin 4038; K L from B. R. Maslin 4005, 162 .Io ^ ou{ I 7'rq routqol / Jo eiuer erlt urq4^^lnJJo seop 1l (ZZ 8rJ) pelnqul -srp {1eprli atou sr n.tog[tq]. q8noqllv fueqly ruo{ snrp"r rul 0t lnoqe per" ue ol pelJulseJv aDutqot t'q8noql se srzeddelI suoll?JtpurluosoJd Jo 'aDutqot lj:roJl t ol pel"leJ flesolJ sI uo fq V o^oq" peleclpul sY 's.rolcElEqJocueJsorouuI 8ursn,(1rsreelrnb peqsrn8utlsrpeq uec,{eq1uot}codsut esolJ uodn lnq JsJltuts ,{11urcg.rednsserurleluos pu" elqelr"^ ,{1elt1t1e3o,reJe solceds e^oq" eq} llv '(sl!€tep reqgnJ soSolotadoqapld pun DuDalooul ees) pera,t,tog-3 Sutaq V 7 'ds sp€eq slr fq seroodsoserll qloq ruo4 poqsln8ullslP sr lnq r\ou Dpladoqaltld 'y pur '3z1rg '1\ 'll^ oupa]oour 't, qlr.'!\ seIlIuSB ssolc s€q osle Drcllq DQnV '(,tr.o1equorssncsrp ees) ,rou 'ds aowqot 'f' pu? ^ou ds ttlotldatos'itt1t y eJ? pasnJuoJeq o1 fyaln l,soLLrv p.to q f qtli{,l.r qlr,tt sercedso,u1 aq1 'p3^Jou-/-e pue sLrolrutu 9. t-z sl€led :p€ar{:ed 7 va.tog :8uol ruur 8 €(Z) solcunpeduo euroq pu? eltq,{\ol ruzorc elrd speeq Jo,{\ou :sreJJ".IeqJ3ut,{o11o3 eql Jo uolleulquro3 ? ,{q pesltl -8oie-rsr pu" (0tZ d) dnor8 "rogtcl V,, eqt Jo requelu 2 st D.to4rqDtrD)V 'aoLtt(lo.! 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LrIEllIoc lNg tP u,trotg 1;ego1 ,{q sldlJcsnu"J4l ,re.rorcruno' rJo N (u,) er) *u 0r ',rtrr., ,ror,r,f$fg.;?fl4]"11#J'f'J'{ # :(HIXAd) ullsol^l 'peo.r dnSunLu"J]el-,{uEqlvuo re^Iu dnurllEd aN rul e 9t0i v I 'ssed 'r8ued Jo :(HJUEd) 900t rrsr_t{ y 6r urnD p.U SurlIlS :fHItad) 89Lt ulltDn { g 'so8ueUdnrn8uorod pue SuII-rIlSueo{rJq p"or urProS-^uPqLV:(HJUAd)^VtZStullsel,4{ 'y ',{uEqlv g Jo N url tZ lnoqe:(HIUld')'ANVI) 68SZtttl\nl4l A A onbunuerJofJoS no area whele these speciesgrow syrnpatrically. The most reliable characters for distinguishing the two speciesare those of the inflorescencebut there are also differences in the phyllodes and seeds. In addition, A. biflora flowers rnainlyin the summer-autumnperiod qhile A. robinqeflowers later in the year (late autunn to early spring). ln A. bifora the flower buds are 2 3'5 nrtn Jong and ovoid rvhile in A. robinae they are narrower, slightly longer (.34 mm) and - cylindrical to narrowly ovoid-compare Figures 9 and 10. in addition, the petals irr .4. bifora are 3 7-nerved while in l. robi ae thcy are l-nerved (this characteris best observedin dry specimens). Other characters useful in recognising A. bifora include its phyllode shape (see below) and indumentum(normally puberulousin A. biflora,always glabrous in A. robinae), its longer peduncles((2) 3-8 mm compared with 1 2'5 mm), and its dulJ, curved or once-folded aril (waxy looking and once- or twice-folded in ,4. robinae). Although the phyllodes in both these speciesvary from obliquely obdeltate to obliquely shallowly obtriangular, the curyature and relative lengths of their margins vary thus producing different shapes compare Figures 9 and 10. Also, in A. biflora Ihe apical mucro is straight or slightly recurved while in A. robinqe it is always straight. Finally, in A. bifora a second nerve (in ad- dition to the principal nerve) normally diverges from the base of the phyllode and extendstowards the gland. This nerve is absentor obscurern A. robinae (except in juvenile foliage). For lnuch of their range A. bifora a:ndA. cltrysocephalahave correspond- ing distributionsand in placese.g. the Stirling Ranges,they grow sympatric- ally. In the field A. bifora is easily distinguished from A. chrysocephalaby rts pale cream to white flower heads (not bright medium golden yellow). Flower heads in dried specimensof A. chr),socephalanormally retain their original (fresh state) colour but ir A. bifora they turn orange-brown. Other inflor- escence characters useful in distinguishing A. bifora include its glabrous, 3-7-nerved petals* (l-nerved and glabrous or hispidulous in A. chrysocephala), its flowers always 2 per head (frequently 2 but sometimes 4 per head in l. chrysocephala) and its normally longer peduncles. In addition, A. bifora can often be distinguished by the following characters: branchlets frequently less prominently ribbed, phyllodesfrequently slightly larger and more patent (as indicatedby angle of midrib with branch), legumesnormally longer and Iesshairy (N.B. glabrouslegumes do occur in both species)and seedsslightly larger.

8. Acaciarobinae Maslin sp. nov.-Figure 10. lAcaciabiflora aoct. non R.Br.: Bentham in Endl.?1 a1., Enum. Plant. Hueg. 41 (1837); Diels andPritzel, Bot. Jb. 35:635 (1905) as to P/'it;e/356.1 lAcacia.livergens jn sched. PERTH.I trlen ad 0.6 nr altus;/-ar?rrlisparsim ad modicepilosi et/vel antrorse puberuli. St?r//dc I 5-2.5 mm longae,basibus persisteniibus et r pungentibus.Phlllodia obhqDe obdeltata ad obliquelate obtriangularia3-7 (8) mm longa,3-5 (8-10)mm lata,paientia, pungentia, glabra,nervo principali excentrico. Pedunculi l-2 5 mm lorgi; capitula+ alba2 floribus. Florcs4i1eri, alabastra: cyclindraceaad angusteobloidea, 3+ mm longa;callcis lobi lriang[lates,petala 3-4 mm longa,glabra, l-nervia. Leguminaang[ste oblonga sed parum curva,ad 60mm longa,3+mm lata, plana. Serrrirain leguminelongitudinalia, 2.5-3 (3.5) mm longa,ca. 2 mm lata,nitida; ar,l/rrs ceraceus. "Low Type.' Mount Manypeak, Western Australia. intricately branched shrub 1-2 ft., liowers white. Granite rocks." 5 Sept. 1935, C. A. Gardner 3324(holo: PERTH; iso: BM, CANB, K, PERTH).

* Best to observeapex ol petals when dry at not lessthan x 10 mag.; sometimesthe flanking nerveson either side of the central nede are poorly developed. t6z ruo4 :99/€urrsel ruo{ : gztoc,{or{ ( r!lo{ : ^;99?z t f i x I J l .V ! D'E,S I8}t .l{trqt;Yj .g E :1801 ullsel,{ { B Luo4 J :82t9 03roo0 S ruo{ I H :T8rt ullsel?'{ !uo{ V Bu,^\oqsreled-r .(soroope.rq : or "",?$1"1"#,irp:?S#r,;"T'#314;";T,X3i1 lJesur qtr^\ opoN D_ pe.&orte uotlrsod puelS :(tpoll,tqd elrue nt-d) uorlBrre,{azrs pue .aDutqot .OI ecleqsbur,roqs sepoll^qd-J o] g qautJq lo tred l:ddn-V DDDV ernfuJ tLbl /vtTsHWa g

! ftl --uEoiz-".-

f il;l dffi,"loN V1 s#':&io ----EurE-----'

V

{hFz- - El j ffiS' Bl +tls I \/ \,/ :tr1 / -i.lr--EE-f-- $tr1 H-l-€M vv # Harsh, spreading,open s/rlub to 0'6 m tall, single-stemmedor much branchedneai base; bark reddish brown towards base of plant, greenishon branches; ev' shootslight green with phyllode margins reddish; btanclllets terete, finely ribbed 1ri6s yellowish). sprrsely to rnoderctelyshortly pilose and/or antiorsely puberulous, greenish. Slirrles I'ery ttarrowly triangular to setaceous,l 5-2 5 l]im long, thickened towards base (persistentstipule bases somewhat pungent), glabrous to sparsely hairy. Phyllodes obliquely ob.leltate to obliquely shallowly obtriangular (shape somewhat-variable but generallv-r,orarher sim11a1 1q A. d1;crgens),prominently angled on adaxialmargin, 5 7 (E) long (as rneasuredalong abaxial margin), 3 5 (8 10) mm wide 1as miasured along adaxial proximal margin), slightly undulate,.patent (as indicated by angle of principal nerve with branch), glabrous, dark green; apical nucrb (teflninatiig principal nerve) pungent, l-l 5 mm long, shaight, brown adaxial proximal niargin convex, ascendentand I parallel to branch, 3 5 (8 10) nm long; adaxial tiistal nargla concare. 3 7 (9) mm long' abaxial matgil corvex, 3 7 (8) mm long Trincipctlll?r'r? excentric..slightly curved, raisJd (when dry), secondarynerve (arising from adarial side of principal nerve near its base) obscure or absent (more obvious on juvenile plalts); zrlr'rirusobscure. Glaarlobliquelysituated at apexofadaxial angleofphyllode' iircutar. 5 0 5 rntn diam., orifice dark coloured. Inforescencessimple, 1 per node; petlurcles l-2 5 mm long, glabrous or som€times .glabrescent; hasal pecfuicular bratls 2, unequal; flover heads - white, with 2 flowers. Brqctioles sessile,ovate, 0 5 mm long (shorter than calyx), concave, minutely ciliolateotherwise glabrous. F/ouets 4-merous; buds I cylindricalto narrowly ovoid, 3 4 mm long, straightor sometimescurved at apex,prominently attenu- ate, minutely pedicillate; call'.r *r length of corolla, divided for j-+ its length illto broadly tiangular ciliolate lobes,tube glabrousand obscurely4-nerved; oerclsnarrowlv oblong but narrowedinvolute and thickenedat apex, 3 4 mm iong, refleredit anthesit,glabrous, 1-nerved (nerve thickenedand prominent at aper of petals when dry): nlcrl' glabrous, minutely stiprtate. Legumes narrowly oblong but slightly curved and sometimesslightly twisted, to 60 mm long, 3-4 mh wide, flat but slightly raisedover seeds(umbo yellow),glab- rous, daik brown, apex acute and slightly uncinate, basal stipe shott: margins thickened(outer edgl broaderthan the inner one), not (or only slightly)con- tracted betweenseeds, pale-coloureC. Seerlslongitudinal in legume, obloid, .5-3 '5) 2 (3 mm long, ca. 2 mm wide, pure dark brown (greyish,brown before maturiiy), glossy plen'ogrant with a narro\Y opening towards the hilum; areule ia. 2 5 rlm long and I lnm wide, not darker than rest of seed; fiirrlclcfiliforrr and 0 5-l tnm lorrg. abruptly expandedinto a once- or lwice- lolded shiny lwary looking) alli which at first is pale yellow but then darker (sometimesdrying orange)towards attachmentto seed.

Distributio : (Figure 23) South-westWestern Australia: Restrictedto the Albcny district from Marbellup (15 km west of Albany) to Mount Manypeaks (aboul40 lm due northeastol'Albanl). Habitat: Most collectionsof this specieshave been made from laterite or sand over laterjte in open-forest dominaledby EtrcalypttLsnargrtnla and species of Casuarina alld Banksiq. Some of the sandy areas where ,1. robinae grows are seasonallywet. At the easternlimit of its known range (i.e. Mount Manypeaks)the speciesis recordedfrom granite rocks

Floh'ering and Ji'uititg perr'orlr Flowers from May to August-September. Legunei with nrature seedshave been collectedin mid-December'

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the adaxial distal margins of its phyllodes are often less obviously concave and its seedsare slightly )arger and darker. Distributionally the two species are well separated(Figure 22). Acacia phlebopetalais rather polymorphic and two varieties are recognised here with the typical one containing two distinct variants. Key to varieties: la. Phyllodes,peduncles and legumesglabrous ... var. phlebopetala b. Phyllodes,peduncles and legumesminutely puberulous .... var. pubescens

9a. var. phlebopetala Figure I l. Shrub 0'4-0'6 (1 3) m tall, either spindly, erect, open (with foliage re- stricted to upper portions of branches)and single-stemmedto moderately branched at base ("typical variant") or dense(foliage I to ground), much branched,spreading (with main branches t prostrate) and forming clumps ca. I 1.5 m diam. ("maritime variant"); branchletspuberulous to strigose with + retrorse hairs ("typical variant") or moderately to densely puberulous to shortly villous with - patent hairs ("maritime variant"). Plryllodes 1 obliquely obdeltate to + obliquely shallowly obtriangular, 6 10 mm long, 6 ll- (18) mm wide, glalxous. Pedunclesglabrous. Legumes to 35 55 (70) mm long, (3) 3 5 5 mm wide. glabrous. Seerls2 5-3 5 mm long, brown to alrnostblack; areole l 5 ! mm )ong. Distribution: (Figure 22) South-west Western Australia: Pallinup River east-northeastto Munglinup (80 km eastof Ravensthorpetowards Esperance). Habitat: As noted below, two variants of var, pl ebopetqla are recognised One is restrictedto coastal and near coastal areasfronr the Pallinup here. "maritime River (the variant") while the other extends River to the Hamersley "typical further inland from East Mt Barren to Munglinup (the variant"). "maritinre The variant" occursin closed-heathor open-heathin shallow sand over quartzite on cliffs or in deep sand on stabiliseddunes. Some of the associated species include Acacia cedroides,A. cocl earis, Pinelea JArruginea, Platysace cbmpressa, Tenpletoniq retusa a\d Westringiq dampie . This been recolded from some coastal creeks where it grows itt variant has al'o "typical sand or silt in cfosed-scrubdomirated by Eucql),Pttrseremopltila. The variant" grows in sand in high-shrublandoften dominated by Eucallptus tetragonq. The understory consistsof low, harsh, sclerophyllscrub which includes species such as D4r'icsl4 terelifolia, Gretilleu hookerqna and G. palentiloba. Flowering and fruiting period: This variety often has two flowering periods. The first is in June and July while the secondoccurs from Decemberto Feb- ruary, Legumes with mature seeds have been collected in November and Decimber.- Sometimesthese legumes are presenton the bushesat the same time as the next season'sflowers reach anthesis. Selectedspecimens: WESTER--AUSTRALIA: East Mount Barren,,'1.S. GedrSe3668 (PERTH); 24 mi (38.5 km) E of Ravensthorpe,A. S. Georye'7616(PERTH); l8 km W of Munglinup { Ra\ ensthorDe-Es pe rance road), -R. R. Ma:lin 255'l (PERTH); Fitzgerald River National Park-B. R. Mdrltr 1882and 4058 (PERTH); About 2 5 km dueNW of No TreeHill, 20 km due NW of Hopetoun. A. R. Marr'rt 4048 (PERTH). Within var. phlebopetalo two variants are recognised and these can generally be distinguished by a combination of branch indumentum and habit (see "typical description above). The variant" extends inland from East Mount "maritime Barren to the Munglinup district (80 km E of Ravensthorpe) while the variant" occurs on (or near) coastal clifs and dunes between the Pallinup and Hamersley Rivers. 298 662 Surpu:.tsu,(1rqBr1s ot iUJted.JlElnpuI {;tqBrls ,{.rar .{urB.rer-u '(UTBJELU ;eurxold ;eD,epe duolE p:ln\€eu se) Jpr\1LU ur (01) g-t 3uo1epalnsEaLu \?) 'urblEru_ lurxrqe,reln8uer.rlqo duol,tutu 0l 9 lErxEpBuo pcl3ue flluoururoJd ,{1,no1eqs ^lenollqoor JlEtlepqoAl,rollEr{s \pnbtlqo sapoll.(tld .{oseqre Llrrnolle()unolq -\noIJeJs sJutnJL!oq.tuJBund (llpruJou.BulpEeld\ .Auol uJUo pue prBU ',snolnreqnd urru 'J"JnBuerJl iluelsrsi-ed t Z ,{J^\orreu.{.re.l o1 elzlnqns saTnd4g",xode ,{lJsrorluE ,{lesrEdsro- rsolrd lroqs ,{151eiepouroi sno.rqeli '(^\ollJ,{ ip:e,ro1 relniup JJql?J sqrJ)peqqrJ,{leuu splq)LtoJq lxedu 1r pa: qll.r po8u$ uJaJ; stoo?,r,41Jr lnq lqb![ :u,rojq lo,{o]S.qlootus 1,/rg :splE,i^u,ropqcJ€ ol tsurpuelALluJnr,0{ puE pJpr^rpun pua 3uo1,{Ie,tr1e1er uego serlJu"tq lnq 'll?l ,]ceJe,asnJJrp lsurulel,qsl€H peqJusJqqcnur u €,2 ol gt.ll/speLuuels_el8urs ,ur|'I]o(] 'lMolrq pnurrlou- J.l D.to )nDdew)oJ t-llut(Isua8Ja^lp o!JDV uor.:nr\rpers'..eln:lrq ^ suJdtJrpeJJcrv.. ,e urruoqtq p:tilouue,{jiJrjro:ur rroq\- .\ou.uou(\ '\tZ6D):oloq) u.tDuts-u)u)oOI y.:1,r1ernri1.,de.l puiang'urttoi rrlf/ tutZ'ZZ6I-lZ6Torooqog .r.S cos trrr6 .uru]oq,/rr.r/r/ .rer iqlueg sua?nttp onotV (^\oloq uorssncsrpoos-EgJ : osl :) : oloq) Io11o14Js.t1,t1 ,re^rx esse^ ..ad,{J 'el .[ , e-rn8rC (ZlSt) tge: l tog uopuo-1 qtueg suaB.rarlpElJBrv TI

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divergent. A. Upper part of branch. B, lo F-Ph)llodes showing Fir:ute 12. Aca.ia - shioe and sizevarration; gland position arro\ ed F Node\ tlh ln\erl .bowlnggland (s stin;lesr. C-Inflorescence. H Flower. I Seed. J Legume valve (dehisced). K- Legume(not dehisced). A from F. M. C. Schock 72; B from S PaLrst154; C from B. R. Maslin 2806; D ftom A. Monison s.n.; E. C lrom B. R. Maslin 3792;F from P. G. Wilson 3725; H ltom A. R. Fairall 622: I-K frorn B. R. Maslin 4014. 300 I0t llbl Ntlsul4/\''a g vm$ {"m ffi\ .--=*- @

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(6 4 kn) w of l)ennark on Manjinluproa.l. ,l ,R Fnitoll.t)22 TPERTH): Blackwood Ri\er.\,\ellerrr Au.traha../. /u.rr'sr t.n. lMfLr; WanganIwongdn] Hill\. Gilbcrr 188 ievll vi.iup, nearcape Naruraltsre. B R i4a'li,t2806 raDw. PLRIHT:8 5 In1S i)] wilchcliffeijnBussel Highway, B. R. Maslin179: (K, PERfHI; S,tirling-Rangc,chestef Fiss, B. n. Maslin4ol4 (PERTHi; EllensPeak [Stirlrns Rcnge]. l^8^oct t902 A Motrisotl ..n.if.pfnfHr:.lu.lnorlhofLllenBrookonCaresRord.se'tofCraccto$n.SParr'tl5'1 IPERTi{):Near Warren Rr!er. a M. C. Stlrcck72 (PERTH); Darling Range' near Vy'aroona i)arn, ca.6 km E of waroona,P. G. lttiron 3725(NSW, PERTH). The designationof a type for A. divrgcns posessome-difficulties. In the "Vasie '. orotolosucit'is cirerrsinrpir a' Rirer, Mrs Mollo5 The description heals rn-ostlvviih veeetaiiuecharacters but some referenceis made to the inflorescencesand legumes. At Kew (K) and Cambridge (CGE) there are specimensof A. direriens which can be regardedas part ofthe type. Theseare libelted VasseRirer.-Mrs. Molloy and are vegetativeexcept for sonlepersistent fruiting peduncles. In addition, the Cambridgespecimen has a.legumevalve on the sheetbut not actuallyattached to the receptacle. Benthamhas annotated "Acacia the Kew specimen divergens"but there is no indicationthat he sawthe Cambridgi one. Although it ii possiblethat flow€ring-type materjal of l' cli|ergens-isextant, thjs is not very probable. It is likglf that the inflorescences describedin the protologuewere vely young (becauseBentham did not describe the individualflower parls ashe normally didl and that thesthave been knocked off the surviving specimens. Therefore,on this basis I have determinedthe Kew specimenis tire holotype of A. dirergensand Cambridgeone an isotype. ln 1923Domin describedA. clitergens\ar. hitsLltafrom materialcollected bv CaotainA. A. Dorrien-Smithfrom"Yallingup and CapeNaturaliste" This ririetj "at distingui'hedfrom the tlpical varietl hy its hir:'ulebranche' rnd (K) hranc'hletr.I haie lookedtor the type of \ar. hihuta al both Kew and Prague (PR), thesebeing the two most ljkely hcrbaria at. which it would be houied.' Nir type was found at Praguebut at Kelv there-is a specimenwhich protologue. This sheet bears the typ€ accords with Domin's very brief "decipiens" annotation exceptthat Domin has written rhe specificepirhet as (instead of "divergens"). As both these specie-soccur i.n the Uninetres- iriangulares,I regaid Donrin's annotationas an unintentionalerror' The Kew speci;en is ther;fore treated as the holotype of A. tlirergens\ar. llitsuta j tiaving now studied the variation in A. tlircrgens,I see I o . ttsti6cationin mainta"iningva(. hirsutaas a distinct taxon. Both hairy and glabrousir.rdivid- uals occur throughout the rangeof this species. Basedon its oYerallgross morphology (includirrgpollen characters Ph' Guinet, pers. comm.) ,4. ditargensis placed in the A. biforu gro-up1p 270)' However, it is noted'that in two infloiercencecharacter\, ie 14) 5 l0 flowers with the A ' truncatq 'srouoDerhead and normally obtttseflower buds, it has afnnities (p. 271\. ,luciq ditergets is distinguishedfrom the other menbers of in 2.'biltoro group by a combination ofthe following characters:heads of t4) 5-l0."creai to pale yellow flowers borne on peduncles3 5-10 mm long; DetalsI-l 7 mm long and obscurely1-nerved: flower buds obtuseor sometimes ibruptty contractedat aper lnol iiSnrficantl)attenulle).. Th^e 'peeie: grow5 tallei rhan the other member.of the A bifttra group.reaching.2 S^m in height Other charactersuseful in recognisingA tlitcrgensinclude its finely ribbed acumrnate branchlets(ribs yellow),'co its often pungent\tipule\. its phl llodes$ ith oungenta pices, n.pic uout gland-biaringangles and concar.'adarirldi'tal liuiEin.. iit relatirel! narro'" legumesl2 I mm widet and it' small 'eed' (2 5-3x15mm). The two speciesto which '4. r/i|etgerts seems most closely.related are '4' phlebopetala aid A. robi Qe. From the former species.it.is distinguished by imallei flowerswhich are normally more numerousin the head and obscurely l-nervedpetals (see p. 297 for further details). In phvllode.shape.l di erge'r nfi"n t"uit a tttitini resemblanceto A. rcbi a,: but is readily distinguishedby t0t -acuo aau€roep^d:o ,tro11e,{ ,(urqs .{11q8r1s lcrql " otul pepuedxe^rrr1jfiIa"Ppl,lJ ,roleq pexouar '8uol ,iuo1 ruIII I eJ puE wroJttu aplunJ:epr^\ unu I gig r.,tu t-9.a J?aro:anssrtelEdJo pueq e {q prrJpJoqurUo .runlrqeqt .pierror'uedo tuDldolnap :Assolb.'AJJrJdutd ,u,rolq punoJ?^eurl I.lBp e.lncsqoue qtr,r {Jep ol u,{lolq tunrp0Lu_'epr^\ruur g.? 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Figure .14. Acacia thry.tocephala. A-Seeds (end and side views). B-Legume. C- tsracteole. D Innorescence.houing 2 flowers wilh pronrinenll)nerved paals 1b bracleole).ttoT-.Phtllodesshouingqhape.sizeandindurnentumrarierion:giandpo.irion arrowed. M \ode \ irh inserrqho\ ing gland(q .ripuler. 4 B_frgm A. R-.-Mains.n.; C D from A. S. Ceorge 3104; E from B. R. Maslin 3916; F from lr_p. f^oyce 102j/1j q llofn C. Perry t72: H from B. R. Maslin 3839; I from B. R. Maslin 17701J ftom B. R. Maslin 2608;K from K. Newbey270; L from K. Newbey41OZ; M from A, Motrison s.n- L0t 'o8uex 'ssed IHIUEd) I/ZOI DioV O V Surlllls runC poU 'plrJo- {\r,rd '0t8I -^oN :(-Igt ) 996ssDrd pueu eerlerlsnv rolrslu' uor8r'Lul :(AN) t96 'r'-niteHi uonLo.l rirU.rr:e,rt(s snqrlurl\ cr:orrelfr-o\oLurlqns ul :(HlUld) ZLI tutdd t '-f Sf.Oil Sli ta'peou(Prrnt\qrr\\uorlJunfl.edpeorlrluluoproDBuole_tull:lHIUld) y'dlue^\s tu{0I:(Hluld)0/z "ir,,,,"N \e[JraLueraJo y' zcjtv,laqnaN dnlE.looJJo,^[ 'urSorr?N )l r].uie-Ztt*E.f :(glufJ) u s rovr.r,/i/ t"t06I ount 9Z :(HJUfd) 9t6t ,rr,/,{ :y j:Lieiril'3 m{ tttlso1{ g 'u11nt g sf1e,[o1 edroqtsu:rey Jo E s 0t:(Hl,Ufd) 6t8t '.sueUv g LUI :(HIUBd) alfoN 'U 'B' 'ssedLunD pou Surlrrls s"pJr-,not:rer elu-I.Jo N tZ OLLI 'llrH :ip1y3.119697raso-7,y y g'IoorquErJJoa LUIt ec ,{6{nS:( HIUSd) u s,?rJ,{ y'y '1.96I ,{lnf 'a8uEd ffollrrs 'ssed relsaqJ :(HIdAd) t ){oux A H seppP.S :lHIUgd) t 'lloLr) lift ai.n;c s ,' Jnl{ Jo eprs N ^\oleg :vrrv{rsllv Nn3Ls?/\A :suaw!)adspa,)al2s 'l3qtu0^oN-pItu ur papelloc ueeq e^€r{ spees oJnletr^l dola^ep fllnJ 01 spees loJ sqluolll llloJ lnoq? sal8l ll JeqolJO pu€ Jeqtu3ldos uI pelcollo. ueeq e^"q suotulJeos nulje,^Aou^\eJ r, rnq t6n8nV o1 ,{e11 ruo.tl .I3,t,iog sqnrqs 6o1,t1 :pot.tatl 3ut11n.tJpuo 3tr1ta'to17 'rueo1 (eltuer8 lo ouolspu"s) ,{lcol tuo4 peproJJr uJJq oslE seq lnq "(PlJ le^o pues ,{11e'ter8Jo pu"s ,\\oll?qs poul?Jp i1:m ur ,{lureru smor8 t; q}eaq-uodo lo qnrJs-uedo ul SurJ.lnoco,{llauots?JJo ,{1uo'iuulqnrqs-uedo qBrqro puelqnrqs-qBrq'lseroJ-uedouI,sJncco ,{lluanbe{ 'sietlqeq tf ;o .{1aue,r " uo4 pep.rocor ueeq s"q serleds stql TzTtgtrg '(ecu"JeosE Jo rluou ru1 97) uupprcg ot }s?e ueql u"{l€O pue utSoj.ruNol q ou l0IlBg lunohtr uroU crp"JodS :"rlurlsnv ulo:lseiA lsa,\\-qtnoS([Z elndrC) :uo!Jnq!.tlstO '7r.r2.,(urqs{pq8rls qsr,rlollef peuolrrr.ll peploj-ecuo ro (pa,r.rnc^llq8Ils ol) lqBterls 't].tJoJtIV '.Pees c olur popuedxe,{1ldnrq" reql€r )ptunJ Jo Ipoq uleu u"ql,le)lep 'epr^,\ 'eu$ oq,(sLu ulru tutu 9.1 "J apatD "LLrttlt\leql spl",lAoluedo ',{reqdrJod16'3uo1 '(u,,|\olq tuotSotnald'. f,wqs punore SurpuelxepuBq u,,\\oJq{lBp ? tlll,t\ 'apr^\ '8uol 'ptosdtlle runrpelu)u^\orq-,{ej8 urru /. I ruur g-9 .7 flltlSrls ol p_rolqo 'euin?a1 'spees ur leurpnlr8uolspars u,tolq lq8rl uea^uaqlelcel:]uoc ^le:€q '(auo rauui JLlluptll JJpl?orq{1tq?t1s e8pa :e1no) f,rr8.irur:u,toJq Peualctql 'spue {.lcp 'snojq"l8 salurlJLuoslo aso;td {I1roqs o1 snolnreqnd ,{leltrapour qloq 'sp;as 'ell]uq lE pJ,rorrtsu^lldnrqp [ Je^o posr€J,{llqBtTS lnq 1€g pu? pl"tl 'api*" u r-ug 3uo1tu tu {0s)0t ol pJ\JnJ (llqttl\ lnq Suolqo (l,\\oJJPur,/i/t?3d7 3 '({lp 'snorqrl8 ,f.iu.ro:esEq pe,to.u"u ELLoroJ uJtl,^ \edt le peue{clq} 1? '(qrrpru ll2}JdJo ,(lruauniord.(11crurou e^rou) pel.reu- 1 o1pelcl:1sel srttq) snolnptdstq:o 'xede 'Jseq sno:qe13 1eelnlo,f.ur ueuo pur pJ^\oJ.rEu ]E pf,llun {llloqs'slseqlu" lE pJyJUf,rtBq,rJuros'3uol r.! u t 9.2 slttltLl:.nolqE16 puE ptueu-t \lJlnJ'qoeqnl :."qo'i"tnlolllr ,{lsnonridsuo.rot tua:.orqel3ralnBuetrt otur qtBual-:tt{-f ro;'p'epr,rip;L11oioc jo rlt6uey','et xrllo:t :01?nue113pu" el?llerlpod,{lo}nurtu 'ellsses'8uol spirci isno:eu-y s./J.rlo1J JIE^o.a"unu€l lutu 9.0 sapaPorg 'iLa,nog rlt!/'r\ 'rPlnqotB '(uea:o tou) no1laf ueplo8 lq3!q runtpeut 1y) Z 'z spDatl.t)t\olf :(Juo lcuJrtttl Jq] u€q} leJlElusLlJnLu lcEJq l"ulolxe) I€nD3ul1 ipo.iq nlnJunptJ ruru (t) saPunpad:epou laszg:sno.rqe13,{lleuuou'3uol Z-E.0 ',lrolJe,{, red {11eur6u teldutrs sacaatsatolful tcLrIlslp pu" lelnlJlJ ecuuo 'jpollkqdi - dr1 :13ue xod? lE PuolD luasqe st1,1r,t/rr1 1o 1tr'

.. .The speciesappears to-be nrost closelyallied to,4. biflora but is readily distioguishedby its goldenflower headsand its 1-nervedpetals (see p. 292 fot fufther details). ln its dwarf habit and golden flower headsA. t:itrysocephalais similar to A. incras.\atabLrt is readilydistinguished byils fewerflowers per head. The specilicephithet refersto the golden flower heads,a characterwhich errablesthis speciesto be distirrguishedfron.r most other mentbersof lhe /. otnota srouo.

13. Acacia incrassataHook., Icon. Plant. 4:1.370(1841)-Figure 15. Lectotlpe: Swan River, N. Holland. DruntmonJ. (K upper "Herbarinrn rjght hand specimenon sheetstamped l-{ookerjanuml8O7''; iso:k|, Iecto. O\).

Acaciadetipietts (C. Koen.)R.Br. var.incrassata (Hook ) Benth..Lond. J. Bot. l:330 (1842). Acaclakr icosa "MidlandW. V. Fitzg., J. W. Austr.al.Nat. Hist. Soc.No. l:ll (1904),ry on. o.r. Tlpe citatiotr: Junction and vicinity, fl. July [K, pERTH], fr. Oct. tiOl [n.v.]. Serpentine,fl. Aug. l9O2 [n.v.]. ly.V.F.". LAceciabillotu aoct. non R.Br.: Paxton'sMag. Bot. 9, t.221 (1842).1 fAucia.t]ecipiens(C. Koen.) R.Br. yar elongataauct. non Benth.t Meisner in Lehm., plant Preiss.l:9 (l844) as to 9?/:$ 955.1 lAcacia clecipiensauct. _non (C. Koen.) R.Br.: Benthan, Flora Austral. 2t351 (1864), pro parte-as to Drummond296 and P/e[r 955.] Dwarf, erect shrub 20 30 cm tall, apparently spreading by subterranean runners, single- or multi-stemmed brancltes prominently ribbed, sparsely to moderately puberulous (hairs patent or antrorse, confined to rtbs). Stipules very narrowly triangular, 1.5 3 (4) mm long, persistent, indurate, scarcely pungent, ascending, - straight, dark brown. Phyllocles obdeltate to widely obdeltate (broadest above the middle and obliquely tnrncate at apex), outlini 308 60r 'N 'le^fu uE,l^S,,:snql JejooH ^q polBlouue e:e sueutcadsDlDSSoDat 'otossD,Dut V "qI y ere roddn o,\\t aql olnl,ri\(,ro1eq ees) ^ou.ds oalotJtl .y ue :laeqssrql uo sueurcedsJe,{\o[ o/rdt e{f ()) ^\s) lB l$qs elSutse uo pe}unour inojJo ouo st DLosso.t)u!tzJo edFlot.rolaqt s" Jreq palralosu:Lurc:ds aq1

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, 3mm , , 2mm , t-,ffi d M# J\ \sFZ W EF

BR /lnsuu 1977

310 :

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Distribution: (Figure 24) South-west Western Austraiia: Coastal resions from Forest Beach (betweenBurbury and Busselton)to Brener Bay llOd km northeast of Albany). lt also occurs on Rottnest Island which representsan interestingdisjunct distribution.

Habitqt: The speciesgrows on coastaldunes in deep,well drained sand and is quite common_in places. lt occursin open-heath,low open-heath,closed_ scrub, open-scrubor sometimeslow open-forestand is commonlv ajsociated with Agonisflexuosa, Acacia cochleqris and Ac. clclops. Acqcia littorea is nol limestone specific like its closest relative, L truncqts. The species(ulder the \ame A. cuneata)is illustratedirr its coastalenvjronment by Baeiinand Mullins (1968,pp. 14and 15). The speciesis sometimesused in sandrirll]e stabilisation programmes,

Flowering andfruiting period: This speciesflowers from August to November but a few flowers may persist until December. Legurnes with mature seeds have been collected in December and January. SeleLtedspecimens: WESTERN AusrRALrA: In collibussaxosis, Bald Head . . . . : propelittorlrm PortiusRegis ceorgii III, Dec.1801, R. Browns.n. (BM, K-sheer wirh Benndttdjstributjon number4310); ln collibussterilibus ad l(tora PorrusRegis Ceorgii I jn ora australiNova Hollandia,R 3/o,rri{BM, K, E-sheetwirh Bennettdisrribution nunber 4Jl t); CapeAugusra, A. R. Faira 809(PERTH); Cape Leeuq;r), ,.-R Mdrlin 1619(pERTHr; AjLJdIly,wesrern sideof PrincessRoyal Harbour, -8. R. Matlin 4025( PERTH);Vasse Ri\ er, M/:r.taollut ;.". lK);In colliculosisarenosis ad litus Daris prope Vasse,2j Dec. 1839, Z. p*rS SetfC, l\tj; Rgtlnes!Islnrdr Parler Point area,,E. C. Snith 2384(PERTH); Brenrer Bay, p. G. llilsii 4307(AD, E, K, L, MEL,PERTH),

In the past the nameA. clecipiens(C. Koen.) R.Br. has beenapplied 10 this new species. However, ,4. decipie s is feated here as a nomenclatr.rralsvnonvm of A. truncatq. The history of the application of thesetwo namesjja liitle involved and is firfly discussedon p. 318. Acacia Ii ored and A. truncarqare closelyrelated and future work may irdicate they should be treated as infra_ specific taxa of the one variable species- Acaciq littorcq can usually be distinguishedfrom A. truncatq by its caducousstipules but there are also diffelen_cesin the phyllodes,seeds, distribution and floweringtimes (seep. 319 and Table 1).

Although often only one gland-bearrngangle occuls irlons the adaxial margins ofthe phyllodesin A. Iittorca,it is_rot unusualto find,-especiallyon juvenileplants, so_n]e phyllodesvith two (rrrely more.)glrnds. Acaciaclecipien.r yar. tr.apezoide.a^DC.and. A. decipiens-var..praeizorsa R. Graham (see below) weredescribed from specimenswith phyllodescontaining more than one gland. These variants are treated here as part of the natural range of variation-of L littorea. ,It is interestirgto note that De Caldolle (1825)arroreously cited the type locality_of vat. trqpezoideeas eastern Australia, whereas in fait the type was collected in Western Australia. This error was repeated by Don (lg5i). The same is true of l. coriacea DC., A. pyriJblia DC, A. biyenoia DC. ind ,i. hispidissimaDC. (: A. pulchella var. glaberrima Meisn.-see Maslin, 1975). 314 'orcnru prloulr(t :?uol Luur (gg) 97-6 lvcrde aqt ltolaq lsuf xede 1? e,\€cuoc etrnb saullauros lnq lqAIEJls = atS,towlDlxDqo:sur8jetrt lsrx€q€ pue ltuttxord uer{lJe oqs '3uo1urru(g1) gT 'proruStsd1,to1leqs ro e.lecuoc,(lmol1eqs I?rx?p€ E '.Buo1 'lrl8leJls seurr:leruosro lq8ruls atBrDLu ruu (0t {t) SZ 6 'Dalolltl lolstp lotxopo I t, ur se qcu?Jq o1 l:1ered + 8ur^t ro^u lu;puJJsu (,{ler"J ,{Jo^ Jo) roqlreuu!8.tDw o} (3utlo^ uaqn) lao11e,{'3uo1 'lue8undlDutxo.td lDlxopD:t^\oJq {rep urur S.0 ,{le.r€qlnq elllJ€ (o^lau ltdrcutrd SurleutruJrl)oDnla loJldo :seuo oq] uEql ropeorq f11q311ssauo lelx€q" eql qlr.{\ Pue qsl\\olle^ l€rxEp" '(qss.g 'snoJqslS uelgo sur8reu uaq,'n)uoeJi lr"p dlo^IleleJ ol uaer8 utntperu '(qcue.rq qlrm e.treu ledtcutrd jo e13ue,(<1 pelectpur su) 3ulpu33s" o1 Surpuerds '^lp rroq,rpequrr,t ,{pq8rlss3ull]ouros'elplnpun ^pq8rls ^tJ{'pal.rnc,4lare; ro '(ur8J"r.u tqBr"Jls 1e1srplerxrpe 3uo1EpJrnsEaLu sn) eptmruu (91) fl-S'(ul8Juttr 3uo[? pernseeuse) 3uo1 urut (Ot Og)EZ 6'G)9.2 9. I - g/'I 'serrdElvrurxord lerxepe elecuul ,(lenbqqo F qlra .l?ln8u?ulqo 01 al€eunc .tapo11[tg spreudn 'Fuol 'snoecElrs 'eseq pe,unc ,{ll"ru:ou ru|-rlg. f S. I o1 ttln8utul ,{l,torleu Jeeu fylurcedsopouolrlql .{llqBrls'snoJnpeJ,(lluenbe{ul'uollP.Je^eq sepo11,{r1d;etle poo^\ plo uo osle uolJo pue serlf,ut.Iq reddn uo luolslslad ^lleLuJou sal,ldl,/S 'esoird ,{l oqs ,{lelerepolu o1 snorqelS 'paqqrr ,{1:}ueutuordo1 ,{1eugslalqzuD.tq 'sixe :et"uls ,{llBruJorr'SurpuoJs" ,tr{rrr,/g : ueeJBtqSrl eluec?r lrotls ^te^ eql uo lcEJq" u.ro{ Sutsrteslootls rau i,4er81q3q'qloours :eseqlE Luclp Jo lrx" ',{qsnq'esueP l,rrg rrlo9 eJ ot lurlrt :llul ur (t) t.Z 9.0 qnlLlspautop ,{lleuuoN '[epn3ls [ (IZ8l) Z log uIoN |rg U ( uoot J) uou llrtg suadlraPDDDVI paltaqDl^ilrerro ; 'a "1aN'))'AN-9S6 ss/r.r,'orr, {ltot rsaullod '(tt8l) HJxad'tupld Ji"jf,r'f.l3lrt ol se)euEd ord 6:I ssrard Lurlol Lr|usrrr\l ,rqrlJ rP\.qlurg nlndutDupDrv ',o?,r''(zr8r) 0€r: r 'ros r uopuo-r' ,,r,It,;fi:;:1lp '{\";.i,!if'trfii'3iii,""'ii!,"!i'i :uorDlpad,(r '^ou'rk)u,{s '(LtBt) zt ienH }ueldLunus '/, " il*'tli?if,fl} ",;i.#*1.:",#,tl '8ellr'Lrlou'(lt8t) ')itn.I Z€L:Z Irf qpuEH ( ueo) )) suatdDaphopollhld '8eli|ruou'(€l8l) C9t:I'Z pe ^\et uoH'JIV ul r{d (ueo) 3) slaldbapu)o)V '(ru,{oou,{s" sE paltx sE,(ltu1tD)rol,tt tur,r//?rrtH ertlell oql :snongjedns) '8ell '.sop '[rxo 'u.o) Luou (to8l) 8l99€:I puo'] tog uuY J ntaldlrapDsaaiAI 'GLLI) '€I 'sne"uurl O6L p3 3oA lsfs ul .'("rrnwV I:(^u) .."rpul ur le;Jq]eliltbqulatY C €^3[ xe,, :uot1o1oadr(1 '(ssrr)r: ssrsez 'puJ rolC 'JLrJJrlgLanp)un.tt (,,tunqruDtpY) Lu11tuotpY '/l oInAIJ (tz8l) 'S8esuuturlloH 'SI tt lln{gd zre^ xc lroH (J uurg) Bl€cunrl slrErY

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D. Tinrlole3917 (PERT|]|. accordingto informationon thisspecinen duplicates atc alsolcdged at A, AD, B, BRI, CANB,K, L, MEL, NSW,RSA, U, UC,US), Although a thorough searchwas made at Gerreva(G), the type of Acacia trLolcat.twas not located. As poirrtedout below, this specieswas originally described$'afetn (/dialtLull truncatum)bythe Dutcb botanist,N. L. Burman. ln the same paper Burman described Polypodiun sltinulosunt ( Synaphea spinulo.sa(Burm.f .) Merr. seeMerrill, J92l) the type of which is still preserved at G. It is tfrereforea little surprisingthat the type of A. truncatais not also presentin that herbarium. The history of the name A. truncata and its nomenclaturalsynonyn ,!. decipiensis a little involved yet quite fascinating(see Koenig 1804,Sims 1815, Smith l8l8 and Merrill l92l). Burman (1768)using only vegetativematerial collectedsupposedly by D. Kleinhof from Java,described this speciesas a new fern, Adiantun truncqtun. Jonas Dryander consideredthat Kleinhof's plant was a speciesof Mintosa and usingthJs information Koenig (1.c.)pubtished the new name M. decipiensfor 'tt citing tl . truncatuffi in synonymy. (ln the original text Koenig erroneouslygave the provenanceofthis plant asnorthwest America, a locality which he later apparently corrected to southwest Western Australia see Sims, l.c.) The name M. decipiensC. Koen. is illegitimate,being super- fluous. lronically, the specimenused by Koenig to compilehis descriptiondid not representthe sametaxol that Burman described. Koenig had therefore misapplied his own illegitirnatename. Robert Brown (1813) published the combination Acacia decipiens,based on ,41.decipiens, but this name was again misapplied and also illegitirlate since he should have DSedthe epithet tru cata. It is unfortunate that most subsequentauthors have foJlowed Brown, thus perpetuatingthe error. Interestingly,Colla (1826) provided the new rame A. dolabrifornis to replacaA. decipiensalrct., but unfortunatelythis name is also ilJegitimatebeing a later homonyn. The taxon to which the name.,1. decipiens has been wrongly applied is described above as .,1. littorea sp. nov. The combination Acacia truncatawas publishedby Hoffmannsegg(1824) and, by indirect reference,was basedon //rantum truncqtum Burm.f. Hoffmannsegg correctlyrealised that this taxon differedfrom the one describedby Koelig as Mintosa decipiens. Hoffnrannsegg's interpretation was never adopted by subsequentauthors wlto usedthe name A. cuneataBe\1th. (1837) for the species which should correctly have been called A. truncatq (Burm.f.) Hofi. ex Hoffnrannsegg. Although Merrill (1.c.)\!rs aware of Hoffmannsegg'scom- bilation l]e misapplied the name A. truncatq to the speciesA. tlecipiens atct, non (C. Koen.) R.Br. ( A. littorea). Until now /1. truncatahas generallybeen known as A. cuneatqand rl ts unfortul'ratethat this well-usedname nlust go into syoonymy. Although I have not seen the type of A. cuneata,viz. Swan River, Hiigel, the original descriptionis conrprehensiveand leavesno doubt as to the application of the name. No specimenpositively identifiable as the type of A. decipiensvar. elongata has been seenby me. The original descriptionof this taxor is very brief and "Swan the type is merelycited as River, Drummond". For the presentI have treatedthis name as a synonymof l. truncqta. Howeyer,it could possiblybe a synonym of A. incrassata(see p. 308). Acaciq cuneata t,ar, glabra Meisner (1844) was based on two specimens viz. Preiss 954 (collected on Rottnest Island) and Preiss 956 (collected from the adjacentmainland near Fremantle). Thesespecimens respectively repre- sent A. littorea sp. nov. and the sand dune variant of A. truncqta rcfefied to below. In a number of herbaria,(K, MEL, P, PERTH) thesecollections are mixed, with the label for Preiss954 accompanyingthe specimensof Preiss 956 and vice versa. 3l ll 6rt '3uoy ruur (Ot) (Og OZ) sepo11,{qde3.re1 ,{1e,tr1z1e.r s1r ,(q pesruSocersr lu?u?^ 'selndrts 'Da.loittl'y srql Jo )cal slr 01 uortrpp" LrI sl (V96ssrerd zr^) ,{teuE^ srql;o ed,(1u,{sreqto eqt :}rr€ue^ s[]1 sr (996 ssroJd zr^) usrel,\ln.tqo18 rcr, 'uo.rJ 'ol oJDaunJv Jo sod^lu,{seqlJo ouo seunpoloJoqt pu"Frr pue lue3plpr ouolsetllrlJe^o ptles ,\\ollsqs s€eJ"ol palculsoJosrMroq]o st oto)untJDt)ocY 'qcrog Jo dn1e,,{1,r1pue ellu€ttlelC uee,^.\leqsounpoJoJ lelseoc eql urqlr,{\ pues doep ur,{llsJrptlods sjntco lu€IJe^1s.tg oq1 selndrlssnocnpec rreq} ur,{gorqc serceds eq1 se,rrleluese.rder eql oro{ regrp oseqt pu" pesruSocaJ ;o 'V l€crd^tr ueaq e^eq olo)untt Jo slueu"A o.\u JlluEUeJ{ pue {Jnqun8 u:o.$lag 'oa.toJJ!l'V ejoJeqsqluolu o,4l'oulrf ur uoseosSuue,rog slr sur8eqoiD)LmJt DpoJy tZ ern8lq (ooqnqulsrp lcunfsrp Surlserelurue sluesardorqcrq,r puBlsllsautloU uo s,ror8 osl€lI'g N),{"9 reruejg ol {rnqung Jo qtnos srnJco oaro ll y otrq,lt (qlrad Jo qllou ru1 0gZ) u€ureef ol .{Jnqung q ou sJ[aao Dio)an.tt DDDJV /\\oleq ees (g-Z : g/1 'epr,,\\ruu (91) Jo '3uo1 6y 9 ruru (0t 0e) gZ Sl) sepoll,(rld,rorr?rr pup SLrolstr ^q pesruSocer sr sercedsoql ses€Jeseqtr ur pu" otnan.ll k, ur snocnp€Josle er€ selndrls eq:lu3qA\'Je^e,roq'seturt ers e;eq1 (s;ooqs,reu8uno,(,fue,t uo fluo luesojd) 't/ Datollll t/ uI luesq" f|.l DlD)untl ur slelqcu"iq eJnl"ul lsoul uo luelslsred or? q3rq,r selndrts orll sr ruaql eltrrdos o1 pasn eal u"c l"Ll; .relJ".r"qc auo arJl secuElsurlsoul uI I e]qeJ ur pelseserqeJr serceqso,r1 0q1 u3a,\\laq sa3u3leJlp ledrcuud aq1 uorl"cgrtuepr ol spre lnJasn oslE ere se{rrl Suria,rou pu? uortnqulsrp lnq sJetceleqcporSoloqdror-u Jo rrorteurquol e ,{q pel"r?des eq vel oa.rolqtl pun De)un.tt nnoty l8oloqd,tou pees puu erun8el crseq pue k, 'spnq 'sre,$og s1e1edpelreu-1 ,{lorncsqo ]o,uog esn]qo 9l-El o] dn q]l^\ sp"eq Je,rou'smuoJsaroUUresoruer"r pecnper fleruerlxe 'odeqsopoll,{qd crs€q lreq} epnlcursorcods lelseoc o,{1 esaqt,(q peieqs sJelJeJerJJ eqJ sercedselq€u?^ ouo eql Jo ?x?l cgrcedse.rJl[sB pe]"ell aq pFoqs ,{eql l"ql el"Jrpur ,{eur IJo,r ernlnJ pue pel"leJ,{1eso1c,(re,r oJr o,rl oql ?€qtpolou sr :}rlnq ereqpeldopr qceoJdde eLIl osl" sr srqJ sorcedsel"r"des sE pele€Jl ueeq e^1:t\I(sLtal(hap t/ eu.I3ueqt Jepun) oato tl y p\te (.DtDunJ f. otueu erJt Jepun) DtDunJt F/ ,{ll?uorllp?rl

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17. Acaciadelphina Maslin sp. nov.-Figure 19. F/'rI?n0 5-l(1 6) m altus,rigidus; /amrli teretes,hispiduli vel interdum puberuli,denrum glab(i. Stipulaealiquantum persistentes. Ph:,llodiaplerullqre delphinata,interdun obde!- tala, 8 14 (16)mm longa,4-10mm lata,glabra, pungentia. 6/a/rrobscura, ntucrone levltet pungentisuperata.Pcd nc li4 7 n1mlongi, glabri ; .apilrl.laurea, sLrb an{hesi 4 5mn]longa, l8-32 froribus. Flure: 4-meri; calycrslobi + oblongi; petala l-1 5 rnm longa, glabra. Leguminaad45 fi1, longa, 3 4 mm lata, + rnoniliformia. Senlr?/rin leguminelongitudinalia, obioideaad ellipsoidea.(2 5) 3 4 mm longa, 2-2 5 rnrr lata; arl/rs conicus,albus. ta€ 'V6ttt urlset{ U g uro4 C :660t Iaq^\oN ) tuo{ C-g :828 ,{aqr,\oN) ruo{ V '(3) puelSoqt Surddotro^o(LU) orJr)ttl fftll^\oqs llrsur qll,\\epollrqd 'rr8r1 C eLunBe'I J peas il rjcuuiq 1o lred rJcldn V FttlqdtaPot)L'tv 6l LLblNtlsaw a B

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Surr:nccoo:cnru lueSund,{11q8r1s (urur 8.0 9.0) leuorls puooes"'u,,!\oJq'pl3!l '1q8rrr1s'3uo1ruu '1uef,trnd(e^leu Sulltulrurel) onnru p)ldD:fJp 7-1 'snorqtl3lsdrauud 'saqcu?rq uoq,tlu:e-r8 1q8r1 of u:or8 e'tr1o Jo spuospl€,t\ol polse8uoc '(lulod lsop?otqlE) epr,t\turu 0l-t'(-u!31€tu lErxEqESuol€ pelns"etu leq,treruos 'e1ri1lepqo rel auol *oi 1911y1-g srulleuros'utSrerulutxepe uo rnds rsln8u?ul snonctdsuoce-rir,^ rnq el€urtunc€pue p:deqs-luarselJssJl lo JoLu ,(lleruJou '3uo1 'rr-1n?uur4 sapc,11it14'luat.sisrsd tutu (9.I) salnd4g ieq.teiuos 'f,;r3I S.0 '(iorie ipru,niorsiuueptde- pernssg {1;eutpnlr8rrol lqBr; E {q urelre^o}nq 'snolqe18 'snolnraqnd r.i^o:q-per ,tlruenbo4l re:t Suruoroq uoos ,{losro.rlu? \Jr!rlJurosro .nolnpid'rq'1sse1qu ,{1itr:-redde ,{lluenbJ4l pJqqrJ ^lJJn$qo ,{lJ^ -\atB 'arait'ts.tt1,rn.'q :ipaLni.! tlrilnar.rrr) qBnor lDq,Mruro' 4ro,7: P-rururrls -rllr,rrro:alSuis'iier u, (q.j) l-s 0 {r'rll par1.rtte.rq,{1uadorrqlEr'pl8tu '(IN'X'gNVJ:osr '996I :Hll{gd :oloq) laq,wN y {lnf tZ ,,'pu9suI tlel'uI ZI qnrqs egt9l 'unoledoH auliei?s,, ri.tterjviv u:'e1se16 Jo rluou (utt t g) \rt z : ad'(J at apex of the.adaxial proxinal margirr and overtopping tlte gland; aclaxial rtraighr froxinul lnqrgin . to slighrlyconcare. 7 ll tlll rnni long 1a lirrle shortel lhan .the abarial nargint: alaxial di,ral rnargiuobliquell- -lottg;concare (sintrsnormally a little angular). rarely + straight, 5 j5 mm abaxial n1aryin.) straight to concave,8-14 (16) nlm long; principal ncni cenlraly .ituatedor sonretirnese\ccntric {nonrally ccntial hut appearingexcentrii becauseof phyll_odeasymnetry), secondarylerves absent;pulr,lzai obscure, \rnoolh or .lighrly.wrinkled.slighll) dilaredar ba.e. Clad obscure.solirary 2. 9r.rlr9l),. situatedon a co_nspicuonsangle(s) along adaxialmargin of phyllod! / || {r4r mm abovet.he put\rnu\. o\erlopped b} a rhofl (0.5 0.8 mm) slighrly pungent mucro, Inflorescencessimple, I per node; peduncles4 7 nim long, glabrous; basal peduncularDracr solitary; receptacleglabrous: flower heq;1i golden yellow, obloid, 4-5 mm long just prior to anrheiis,wirh I b 32 flowers. Brqct&)les.ca..l mm 1ong.cllw' narrowlyoblong bur :lighrlydilaled tonards apex. puberutousabartally: laminae Leeled,colcave, ciliolate. Flowerc ,l-rnerousl j.length .a7rx + of petalsnornrally divided for ca. ?its length into ,l _oblongcjliolate light brown lobes which are puberulous a.ttaxrally:,'lcgrl/respetals I l,'5 mnr long^.glabrou.._ very ob.curely l-nerved: orarl sessile, to 45 rnrnlong.3 4 mnrwide. \traigil or slighrllcurved. firmly chanaceous and sorne\hal britrle.unilormly rai.ed o\er seedst rnoniliform).glabrou. or graDrescent,lnedrum to dark brown; r??af8lalsomewhat contractedbetween .eed'.bareiy thickened. pale coloured. Scr/, longiLudinalin legunre,obloid ro ellipsoid.(2.5) 3 4 mnr long-2 2.5 mm uide. ruigid, medium-brownro dark broun. with a darL lineexterrding around the periphery.dull: pleurogron open rowaro( tne n'|umi areole z J mln long, ca. I mm uide: funicle short and fililorm.erpanded rnro a frirl; largeurinkled : conical{nor ionvoluredlnhire slgnuy snrnyafl/ ca. I mm tong.

Dislribution: (Figure 24) Sourh-west Western Australia: South coastal regions from _the-Pallinup River (150 krrr southwestof Ravelsthorpe) east to IsraeliteBay (120km eastof Esperance).

Habitttt: Accord U to Mr Ken Newbey (pers. comm.) this speciesnormally grows o[ expos€dflats (rarely on slopes)in well drairredsand or loam; it ii rare in loamy clay. It occursin 1ow open-woodlandand lall open-shru6land or sotr-tetimestall shrubland.

Flowering periotl: July to September-October.

FrLtitingpetio.l: Legunes with mature seedshave been collected between late Novcmber and late December.

Selecteclspccimens;WESTLRN AustR LIA: 5 mi(8 kll1)N ofnouth ofthe FitzgeraidRiver, K. M Alk ,3l3,IPERTH);fitzgerald Riverarea, ca. I12.7 kln ESE of Ongerup,Z E i./. Apt,ut,_t.,Lcthhrtdseancl. R.. Cnvuty 3-214(NSW, PERTH); Corner ol East Spaddiilup Road and_Hlgh$a],i0 mi (48 Lm) N of Esperance,I. B. A,nitage 551 (PERTH); Condingup, T. q: Daniells.n. tMEL 502977J1Lower reachesof Fitzgeraidnivei. B. R.'li,taslin3fr9'l (8 IPE:\lH")i.) nI km)-NW of Mounr Maxwe|, K. Newbe! 828 (PERTH);2 mi (3.4 Hopetoun, K. Nrw6.7 t6t6 (PERTH); 2 km S flll_rygJ jl9,of Mounr Maiwe . l(. Ne,r6er4099 (PERTH); FitzgeraldRiver Nationatpark,34" 05,S, 32'E,A. S. t/"r/o;6365 (pEitTH); Israelite Bay, 27 Nov. 1950,J. H. Willis s.n. (MEL 502989).

Although A. delphina has both 4-merous flowers and phvllodes which are basically triangular in outline (although nornrally very asyrnrnetricy,it does not appear to be very closely related to the other members of the (Jnineryes_ Triangulare.s. It is at once recognised by the small mucro which ovenops rne gland on the.ildaxial nrargin of the phyllode (Figure l9D). This charaiter is otherwrse unknown to ne rn Acacia. Another cltaracter which distinguishes A. delphina,rs.its obloid flower heads with l8-32 closely arranged flowerslheads normally globular and flowers not exceeding 16 in the othei members of the Uninetrcs-Triqngularar). The f noniliform, firmJy chartaceous legumes arg 324 szc '8 .99tr ,log .suadnap .(t0g .) ,clNao) 1 T puo.I trrJV ,sr&/4,1r l) 'tt:I .(|ZBI).J .rcasNNvN,r.roH rlqopN rln)lgd.216^ 1jlrnv I Luu,rlrid,(qp,rr.,lsny urrlse^\-rruoN urporr;i6l pt#',13llt;fi[ ]il"YrAl:i".$Yi\l*, 'tzt .,,,-n rov:z sluurdsnoap,tlu.J.Ircl oq] Jo ^rorsrHINreuaD v arriil'$]*oo 'srlDrucpr..o errr'rlsnvorrqdu,aor,{qd rrururir^r ryGo roo,) a I]lttfi jti"ollot-,o '6lt:ltouuoJ.rrs pE.rV.(aW DDDV .OZgl).V 'tLb-ggtt. 'srlDrnlr]N 1,y11

sru,(nou,{scqo1 e,ro,a rnnnluan" 11,^,[l,fill";,t;?jr';:ilifr,I",i,il,i::?i{,"5.:l: aq1 ur ex?l ol seuruueql JeJf,J,{lelrlrsodlolruec I paLlrrll"xeuoiq e,\"q sod,{l erll lrtun ^ slu"ld polr^rtlnJ LUoJJ,{Jq"unsiid poquJsJpe;em Burpooerdeql qtt,.\\reqteSol ,{1rr;r,rsrql cJ'eiur.les D,tob[iJpLu.tet'suatdt.taltor.to:ty '(6t81) ,e8uueg l8t :t uofl plBf rolg st.!Dlt1?uDt,tt.JE^ suatdlJap DtrDrV saurEusnorqno 'trr qdlop3u rBrrrr 1d' r: selquasol,(ll"nrJou curllno esoLl,r \Jpo11,{qd:q1 o1 s.raga.r lrrllrdo rfrj:Os iq1 '( |1eny1g DllltldoliSuo,lts y 3'e) aosoutatny-sal Jaulun oqlJo slaqtuou ourosqlr,n flidge u" eltJrpur uellod cq] sJelcpr?rlc ourxe eqJ .DUItllJoq .y pue DlDln$oq 'tr Jo lsereeu s\lees ouitf\ap t dno]8 srqt u[llr,r lnq saro\Buo!U-salautun oy^t Jo sla,qLuauJorllo 3rll lll!,\\ drqsuorlelelosolo " elEJrpur1ou seopsercods srql jo .dno:3 uellod 3rlt ( ururolsred) leurnD q(t ot Surprorrv srql ro1 1eord.{1ebqe Mnsr-rr. B. R. (1975). Studies in the genus Acacia (Mimosaccae),1. A. revision of the SeriesPulchellae. Nuytsia l(5):388 494. METSNER.C. F. (1844). Acacia. In J. G. C. Lehmann, PlantaePreissianae l:3 23. Ham- burg. METSNER,C. F. (1848). Acaci1. In J. G. C. Lehmann,Plantae Preissianae 2:199-206. Hamburg. MERRILL.E. D. (1921). A reviewof the new speciesol plautsproposed by N. L. Burmanin his Flora lndica. Phillip.J. Sc'.19(3):347 353 (extract). MUELLER.F. (1859). Contributionesad AcaciarunAustraliae Cognelionen]. J. Proc.Linn. Soc.Bot. 3:l 14-148. RADToRD,A. E- et al(19'74). VascularPlant Systemalics. New York. SEEM^NN.B. C. (1846). Acacilt. VerhandlLLngender Kaiserlich-KoniglichenGartenbauge- sellschaftin Wien, pp. 5-77. SrMs,J. (1815). AcaciaLlccipie s. Bot. Mag.41. 1.1745. Surrn. J. E. (1818). ,4rccia. In Rees,Cyclopaedia vol. 39: Supplement. "Vegetation" SpEcHr,R. L. (1970). chapterin TheAustralian Environment- €d. 4. C S I R.O. Melb. Uni. Press. VAssAL,J. (1972). Apport des RecherchesOntogeniques et S€nlinologiquesa L'Etude Morphologique, Taxononique et PhylogeniqLLedu Cenre Acacia. Tav. Lab. Forest. loulou.eVol.8, Arr. 17,pp. | 128.

Figure 20. Distribution ol Acocia horrid a, A. inttps,A. p1'crocephala,A- senitrttllatd an(l

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perpnls suarupadspalaqunu ol xapul TLF), 3766(8), 3768(7),3769 (ro-CANB, PERrH), 3770(12), 3112 (1\,3771 (7), 3775 (l+Type),3778 (6 Type: CANB, K, MEL, NSw, NY, PERTH),3781and 3781a(8- MEL, NY, PERTH), 3784(1 MEL, PERTH),3785 (GAD, BRI, K, C, PERTH, RSA), 3786(l AD, PERTH),3788(11),3789 (11 K, PERTH),3790(6),3792 (10-K, PERTH), 3'793(6\,3796 (5), 1797 (15), 3798(15 PERTH, TLF), 3799(15-K, PERTH), 3800 (15-MEL, PERTH),3801(ls-AD, PERrH), 3803(1s),3809 (16),3839 (12), 38s4 (12), 3879(9b-MEL, PERTH), 3882(9a), 3883(9a), 3885 (9b), 3890(9a), 3916(12), 3986 (15),3987 (15),3988 (5),3990 (lr),3992 (4),400s(7),4006 02),4014 (r0),4016 (7), 4018(6), 4019(8),4022 0 CANB, PERTH),4025(14),4036 (7), 4038(7), 4048(9a), 4057 (9b-Type:CANB, K, PERTH),4058 (9a CANB, K, PERTH),4063 (4),4082 (15),410?(16),4lll (16-Type:CANB, K, MEL, NSW, NY, P, PERTH),4154(16), 4175(3), 4187 (13),4189 (15),4189a (15), 4190 (15),4l9l (15),4192 (15 CANB,PERTH), 4193(5 Type: CANB, K, MEL, PERTH), 4193A(5-G, PERTH), 4194(5),4195 (14), .1r96(5), 4359(15-CANB, K, PERTH), 4360(15), 4361 (15), 4364 (5), 436s(3), 4401 ( r4),4404 (r5). Mccutcheon.C. S- 327(10). Melville,R. 4417(8 K),4335 (15-K). Nelson,E. C. (l4-PERTH: ANU sheetno. 16885). Newbey,K. 252(12\,21O (12r,682 (7), 828(17), 1328 (7-MEL, PERTH), ls07 (14),1616 (17), l6l6D (j7-Type: CANB, K, NY, PERTH), 3483(7),4083 (9a),4099(17), 4136 (7), 4162(12), 4196Gr, 4897(9a), 4929(9^r, 4930(17). NewmaD.I. V. 750(14-K). Oldfield,A. 463(1 MEL, PERTH). Olsen.I. 618(5 K). Orchard. A. E. 4391(7\. Paust,S. 154(10),178(14),201 (t4),233 (6),314(14),489 (.1),569 (1). Perry, C. 172(12). Phillips,M. E. WA,/684047 (10). P.eiss,L. 954 (l+Syntype of l. cuneatavat. glabtu: P, STR, w; sphal . "956" K, MEL, PERTH, W),955 (13 FI, G. GOET, HBG, K, MEL, NY, P, STR, w),956 (15-Syntype of A. cuneatavar. glab,?. NY, W; sphalm. "954" K, MEL, P, PERTH.),957 (15-FI, G, GOET, HBG, K, L, MEL, PERTH, STR, W),959 (I-G, GOET, HBG. K, L, MEL, NY, STR),960 (10-NY), 963(12 NY), 965(4-syntype of A. horridula:NY), 966 (12- MEL),968 (14-C, NY), I l5l (3 lrctolype: G, MEL. NY, P, PERTH, W). Pritzel,E. 341(12-Type: B, BM, E, G, K, L, PR, US, W),356 (8 B, BM, E, C, K, L, PR), 598(15 BM, G, K, L, PR, W). Roserson,w. 93 (14),336 (12). Royce,R. D. 2816(2), 3603(7), 3638a(12), 3725 (8 NSw, PERTH), 3839(13), 3848 (s), 3913(2), 3935(14),4261 (8), 461s(2'),462s (2), 4830 (5),4908 (2), 8078(7), 8341(16), 8480(16), 8926 (12), 8939 (17), 9013 (7),9419 (16),9194 (1s), 10271 (12). Schock,F. M. C. 60 (t+K, PERTH), 72 (10). Scrymseour,E. M. 179(15), 1065 (14), 1086 (l), 1097(14), ll98 (6), 1264(14). Skinner.P. 252( l). Smith,F. C. 1934(15),2384 (14), 2385 (14), 2386 (14), 2388 (14). Souster.J. 680 (l K). Stewart,W. S. 335 (l). Storr.G. 296(10). Stoward,F. "136 or 938" (4),710(1 K). Tindale,M. D. 3789(4-K),3917 (15). Tindale,M. D. et Maslin, B. R. 273 (12 K, PERTH),292 (7 K, PERTH),298 (10 K), 346(r4-K). Trudgen,M. E. 1708(9-Type: CANB, K, MEL, NSw, NY, PERTH). went, F. W. 58 (14),75 (11),212(16). weston,A. S. 6365(17),6999 (16). Willis,J. H. s.n.(17 MEL,502989). Wilson, P. G. 3725(10 NSW, PERTH),4195 (12 MEL, PERTH),4307 (1,fAD, E, K, L, MEL, PERTH), 5455(9a CANB, PERTH),6279(GK, PERTH),7780(12), 10235 (I+K, MEL, NSW, PERTH). Wittwer,E. 261(l). wrisley, J. w. wA/68 4047(10), WA/68 4580(10), wA/68 4983(7). f32 ttt

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