Journal of Threatened Taxa

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Journal of Threatened Taxa Building evidence for conservaton globally www.threatenedtaxa.org ISSN 0974-7907 (Online) | ISSN 0974-7893 (Print) Short Communication A new record of the lesser-known butterfly Small Woodbrown Lethe nicetella de Nicéville, 1887 (Lepidoptera: Nymphalidae: Satyrinae) from Khangchendzonga National Park, Sikkim, India

Sailendra Dewan, Bhoj Kumar Acharya & Sudeep Ghatani

26 May 2018 | Vol. 10 | No. 6 | Pages: 11775–11779 10.11609/jot.3987.10.6.11775-11779

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Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2018 | 10(6): 11775–11779

A new record of the lesser-known butterfly Small Woodbrown Lethe nicetella de Nicéville, 1887

(Lepidoptera: Nymphalidae: Satyrinae) from ISSN 0974-7907 (Online) Khangchendzonga National Park, Sikkim, India ISSN 0974-7893 (Print) Communication Short

Sailendra Dewan 1, Bhoj Kumar Acharya 2 & Sudeep Ghatani 3 OPEN ACCESS

1,2,3 Department of Zoology, Sikkim University, Tadong, Gangtok, Sikkim 737102, India 1 [email protected], 2 [email protected] (corresponding author), 3 [email protected]

Abstract: This study reports the recent sightng of Small Woodbrown Lethe Hubner [1819], is a buterfy genus under the Lethe nicetella from Khangchendzonga Natonal Park in West Sikkim subfamily Satyrinae of the family Nymphalidae. The District, India. It was originally described by de Nicéville (1887) based on the collecton of males and one female by Oto Möller from genus is distributed from Borneo through the Sunda Sikkim but the exact type locality was unknown. We also reviewed Islands, Japan, Siberia, Himalaya and peninsular India various historical and contemporary reports on the descripton and distributon of this species. We did not fnd any report of collecton (Mani 1986). Morphologically, the upperpart of these or sightng of the species from India afer Elwes & Möller (1888). buterfies are brown with apical spots on the forewing The occurrence of this species in Sikkim is mentoned in Haribal and spots or ocelli on the hindwing. They also bear (1992) but it is not clear whether the report is based on sightngs or historical records because sightng locaton is not given, indicatng its distnctve ocelli on the under parts of the wings. The descripton based on museum specimens. Hence, we conclude that habitat of most of the species of this genus is bamboo the Small Woodbrown L. nicetella was sighted afer a gap of around forest or grassy patches in the forest. 120 years. Further, we have provided the frst photographic records of a live individual of this species from India. Our fnding indicates a Sikkim is a small land locked Himalayan state in possibility of existence of many cryptc taxa that should be explored India covering an area of 7,096km². It lies in western using morphological and molecular approaches. extremites of the eastern Himalaya, a part of one Keywords: Buterfies, Lethe nicetella, Sikkim, Small Woodbrown. among the 36 biodiversity hotspots of the world (CEPF

DOI: htp://doi.org/10.11609/jot.3987.10.6.11775-11779 | ZooBank: urn:lsid:zoobank.org:pub:AF06A883-FD20-4383-A6F0-D28B3963E1E6

Editor: Sanjay Sondhi Titli Trust, Dehradun, India. Date of publicaton: 26 May 2018 (online & print)

Manuscript details: Ms # 3987 | Received 30 December 2017 | Final received 08 February 2018 | Finally accepted 02 May 2018

Citaton: Dewan. S., B.K. Acharya & S. Ghatani (2018). A new record of the lesser-known buterfy Small Woodbrown Lethe nicetella de Nicéville, 1887 (Lepidop- tera: Nymphalidae: Satyrinae) from Khangchendzonga Natonal Park, Sikkim, India. Journal of Threatened Taxa 10(6): 11775–11779; htp://doi.org/10.11609/ jot.3987.10.6.11775-11779

Copyright: © Dewan. et al. 2018. Creatve Commons Atributon 4.0 Internatonal License. JoTT allows unrestricted use of this artcle in any medium, reproducton and distributon by providing adequate credit to the authors and the source of publicaton.

Funding: This paper is a part of the project “Distributon patern and conservaton of buterfies along the elevatonal gradient in Rangeet Valley, Sikkim, Eastern Himalaya’’ funded by The Ruford Foundaton through Ruford Small Grants, UK (Grant ID:20758-1). SD was supported with non-NET fellowship of University Grants Commission, New Delhi provided through Sikkim University.

Competng interests: The authors declare no competng interests.

Acknowledgements: We thank Sikkim University for providing facilites to undertake this research. We would like to thank Dr. Peter Smetacek and Mr. Monsoon Jyot Gogoi for helping us with identfcaton of Lethe nicetella and providing valuable historical notes on this species. We thank Dr. Basundhara Chetri for valuable suggeston on taxonomic approaches. We are grateful to the Research Evaluaton and Monitoring Cell (REMC) of the Department of Forest, Environment & Wildlife Management, Government of Sikkim for providing us with permit to study buterfies in Sikkim. We would like to thank Mr. Nawangla Bhuta along with other members of Buterfy and Moths of Sikkim, Nature Conservaton Society (BAMOS) for their contnued support in our feld studies.

11775 Record of Lethe nicetella afer 120 years Dewan et al.

2017). Due to its steep mountainous terrain, Sikkim Table 1. Details of historical records of Lethe nicetella from India experiences rapid changes in climatc conditon from sub- Scientfc Name Locality and elevaton References tropical type in low elevaton to alpine conditon in high Sikkim (based on Oto Lethe nicetella de Nicéville (1887) elevaton. The synergetc efect of elevaton, climate and Möller’s collecton) Collected from Ghoom historical factors (pre-historic tectonic movements and Lethe nicetella and Tonglu (now in West Elwes & Möller (1888) paleoclimate) has made it one of the most biologically Bengal), 2,134–2,744 m Sikkim (Based on de diverse regions in the world, despite its small area (Ali Sinchula nicetella Moore (1892) Nicéville 1887) 1962). Buterfies in Sikkim are represented by 689 Lethe nicetella Sikkim, 2,134m Bingham (1905) species, and genera such as Lethe appear to display high Lethe nicetella Sikkim, 2,134m Antram (1924) diversity (Haribal 1992). Mentoned about museum Eminent naturalists extensively documented specimen collected from Lethe nicetella Tablot (1947) th Gangtok and Karponang in buterfies of Sikkim in the 19 century. Among them Sikkim, 1,524–2,744 m were the noted entomologists de Nicéville (1881, 1882, Lethe nicetella Sikkim Wynter-Blyth (1957) 1883, 1885, 1894) and Elwes (1882, 1887) but most of Lethe nicetella Sikkim Haribal (1992) this literature refers to taxonomy and listng of species. Lethe nicetella Sikkim, 1,800–2,800 m Kehimkar (2008) Haribal et al. (1988) presented a checklist of 103 species of buterfies along with their sightng locatons in Sikkim. Haribal (1992) remains the most exhaustve publicaton made so far on the buterfies of Sikkim. menton of this species is by Elwes & Möller (1888). The With a gap of almost two decades, systematc studies authors recorded this species along the Goompahar and on buterfies in the region have increased in recent the fanks of Tonglo during July and August. “Goompahar” years (Acharya & Vijayan 2011, 2015; Chetri 2015). referred to here is now known as “Ghoom” and “Tonglo” Species protected under Indian Wildlife (Protecton) Act as Tonglu, both lies in Darjeeling District, West Bengal, 1972, such as Symbrenthia silana (Kunte, 2010), Lethe India. Darjeeling was a part of the then Kingdom of margaritae and Nepts nycteus (Rai et al., 2012) have Sikkim, but later managed under the Britsh Empire and recently been rediscovered in the state afer almost 100 now a part of West Bengal State in India (Subba 1992). years. All these studies have indicated the probability of Hence, based on Elwes’s observaton (Elwes & Möller occurrence of many species that awaits rediscovery or 1888), Darjeeling, West Bengal, should be included possibly even new species discovery. in distributon range of the species in the upcoming Here, we report the recent sightng of Small literatures. Bailey (1951) reports the occurrence of the Woodbrown Lethe nicetella from Bakhim in West species in Chandagiri (1,820m) and Godavari (1,520m) in Sikkim District, Sikkim, India. The species is protected Nepal, which could be considered as the western most under Schedule II of Indian Wildlife (Protecton) Act extent of the species. The species has also been known 1972 (Anonymous 1997). We also reviewed various to occur in Bhutan (Singh & Chib 2015). Several authors historical and contemporary reports on descripton and have mentoned the occurrence of this species from distributon of this species. Since very less informaton India but it is not clear whether the authors collected is available on this species (and genus Lethe as a the specimens or referred to the museum specimens whole), our fndings add to the existng informaton on (Table 1). Lepidoptera of the Indian sub-contnent in general and The male buterfy has been reported to be more Eastern Himalayan region in partcular. common than the female (de Nicéville 1887; Elwes & Möller 1888); however, this buterfy has been Distributon, habitat and status based on literature considered to be rare (Evans 1927; Tablot 1947; Haribal Lethe nicetella is one among diverse group of species 1992; Kehimkar 2008). under Lethe genus of subfamily Satyrinae. L. nicetella was originally described by de Nicéville (1887) based on Descripton of the species the collecton of males and one female by Oto Möller Lethe nicetella is one of the smallest species in the from Sikkim but the exact type locality is unknown. Lethe genus with a wingspan of 45–50 mm. Moore Based on the museum specimen housed in the Natural (1892) referred to this species as Sinchula nicetella but History Museum, London, Talbot (1947) reported the successive authors have followed the original name, i.e., occurrence of the buterfy in Gangtok and Karponang Lethe nicetella. in Sikkim (1,524–2,740 m elevaton). The other notable The key morphological features of this species that

11776 Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2018 | 10(6): 11775–11779 Record of Lethe nicetella afer 120 years Dewan et al.

Image 1. Distributon map of Lethe nicetella

a b

Image 2. (a) Temperate forest in Bakhim, Sikkim (KNP); (b) Habitat of Lethe nicetella

distnguishes it from other similar species (Lethe sidonis Michelia, Ilex, Cinnamomum, Betula can also be found and Lethe nicetas) are given in Table 2. here. Secondary growth consists of Debregeasia sp., Utrica sp., Viburnum sp., Osbeckia sp., etc. Recent sightngs Six individuals of Lethe nicetella were seen at around We sighted Lethe nicetella in Bakhim (27.910N & 11:00hr on a sunny morning on 18 June 2017. The 88.190E) at approximately 2,700m elevaton in West species was initally mistaken as Common Woodbrown District of Sikkim, India (Image 1). Bakhim is one of the Lethe sidonis, a closely resembling species; however, restng places along the Yuksom-Dzongri trek, a famous the absence of ocelli on its forewing provided a clue of trail for expediton to Mt. Khangchendzonga. The area it being a diferent species. We quickly photographed falls under Khangchendzonga Natonal Park (KNP), a the species (both underwing and upperwing; Image recently designated UNESCO World Heritage site. The 3) and matched the characteristc features of the mean annual temperature of Bakhim is around 11.58° photographed individual to that of the descripton in C, while mean annual precipitaton is 1,827mm. Bakhim literature (de Nicéville 1887; Evans 1927; Haribal 1992; is characterized by temperate mixed broadleaved Kehimkar 2008). On careful examinaton and with forest. Trees such as Castanopsis sp., Quercus sp. and the help of experts we identfed the species as Lethe Rhododendron arboreum are common in the area nicetella. The buterfies were feeding on faeces of catle (Image 2). Various species belonging to genus Magnolia, and horses and some individuals were basking exposing

Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2018 | 10(6): 11775–11779 11777 Record of Lethe nicetella afer 120 years Dewan et al.

Table 2. Key morphological diferences to distnguish Lethe nicetella from Lethe sidonis and Lethe nicetas (de Nicéville 1887; Elwes & Möller 1888; Bingham 1905; Evans 1927; Wynter-Blyth 1957) Common Woodbrown Yellow Woodbrown Small Woodbrown Features Lethe sidonis Lethe nicetas Lethe nicetella Wingspan 45–60 mm 48–55 mm 45–50 mm Upper part Deep bronzy brown Golden brown Golden to reddish-brown Underside sub-apical ocelli in Present Present Absent forewing Subequal with ocelli in 3 and 4 Underpart ocelli in the hindwing All ocelli clear Similar to Lethe sidonis blurred

their golden-brown upperwing. On being disturbed, the Conclusion buterfies few short distances and kept close to the The rediscovery of Lethe nicetella afer a gap of ground. 120 years in Sikkim has indicated the occurrence of its We did not fnd any reports of collecton or sightngs habitat and host plants in the region. No literatures, of the species from India afer Elwes & Möller (1888). however, on host plants or ecology of this species are While Haribal (1992) mentoned the occurrence in available. We also presume that more such species Sikkim but it is not clear whether the report is based on occur in the region and need further exploraton. There sightngs or historical records. The author also mentons is a possibility of existence of many cryptc taxa that that out of 689 species listed, only 250 species were should be explored using morphological and molecular primarily sighted, and locaton of sightngs has been approach. The conservaton focus is more biased provided for all of these species. Since the locality towards large and charismatc species but buterfies are of occurrence of Lethe nicetella is not mentoned in also a signifcant component of the forest ecosystem. Haribal (1992), we believe that the descripton was Hence, conservaton atenton is necessary for protecton based on museum specimens. Hence, we conclude and long-term survival of underrepresented taxa such as that this is the frst sight record from India afer a gap buterfies. of around 120 years. Further, we have provided the frst photographic records of the occurrence of this species References in India. Photographic record of this species is also not available in exhaustve online sources such as htp:// Acharya, B.K. & L. Vijayan (2011). Buterfies of Sikkim with reference to elevatonal gradient in species, abundance, compositon, similarity www.ifoundbuterfies (Kunte et al. 2018). and range size distributon, pp. 207–220. In: Arawata, M.L., S. Based on our feld study, the species is not very rare Tambe (eds.). Biodiversity of Sikkim: Exploring and conserving a global hotspot. IPR Department, Government of Sikkim, Gangtok. in its distributon range, more specifcally in the area of Acharya, B.K. & L. Vijayan (2015). Buterfy diversity along the recent sightngs. Because of high similarity, researchers elevaton gradient of eastern Himalaya, India. Ecological Research and naturalists could easily confuse Lethe nicetella with 30(5): 909–919; htp://doi.org/10.1007/s11284-015-1292-0 Antram, C.B. (1924). Buterfies of India. Thacker, Spink & Co., Calcuta Lethe sidonis. Hence, the occurrence and sightngs of (Kolkata), 226pp. this species may have gone unnotced. Anonymous (1997). The Wildlife (Protecton) Act, 1972 (as amended

11778 Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2018 | 10(6): 11775–11779 Record of Lethe nicetella afer 120 years Dewan et al.

up to 1993). 4th updated editon. Natraj Publishers, Dehradun, Elwes, H.J. & O. Möller (1888). A catalogue of the Lepidoptera of 158pp. Sikkim; with additons, correctons, and notes on seasonal and Bailey, F.M. (1951). Notes on buterfies form Nepal. Journal of the local distributon. Transactons of the Royal Entomological Society, Bombay Natural History Society 50(1): 63–298. London 36: 269–465; htp://doi.org/10.1111/j.1365-2311.1888. Bingham, C.T. (1905). The Fauna of Britsh India, including Ceylon and tb01313.x Burma (Buterfies), Vol. 1. Taylor and Francis, London, 537pp. Haribal, M. (1992). The Buterfies of Sikkim Himalaya and their CEPF (2017). Critcal Ecosystem Partnership Fund. htp://www.cepf. Natural History. Sikkim Nature Conservaton Foundaton, Gangtok, net. Date accessed 17 December 2017. 217pp. Chetri, N. (2015). Distributon of buterfies along a trekking corridor Haribal, M., N.D. Mulla & N. Chaturvedi (1988). The buterfies of in the Khangchendzonga Biosphere Reserve, Sikkim, Eastern Sikkim. Journal of the Bombay Natural History Society 85: 271–280 Himalayas. Conservaton Science 3(1): 1–10; htp://doi.org/10.3126/ Kehimkar, I. (2008). The Book of Indian Buterfies. Bombay Natural cs.v3i1.13767 History Society & Oxford University Press, Mumbai, 497pp. de Nicéville, L. (1881). A list of buterfies taken from Sikkim in October. Kunte, K. (2010). Rediscovery of the federally protected Scarce Jester Journal of Asiatc Society of Bengal 50(2): 49–60. Buterfy Symbrenthia silana de Nicéville, 1885 (Nymphalidae: de Nicéville, L. (1882). Second list of buterfies taken from Sikkim in Nymphalinae) from the Eastern Himalaya and Garo Hills, October. Journal of Asiatc Society of Bengal 51(2): 54–66. northeastern India. Journal of Threatened Taxa 2(5): 858–866; de Nicéville, L.(1883). Third list of buterfies taken in Sikkim. Journal htp://doi.org/10.11609/JoTT.o2371.858-66 of Asiatc Society of Bengal 52(2): 90–100. Kunte, K., S. Sondhi & P. Roy (2018). Buterfies of India, v. 2.35. Indian de Nicéville, L. (1885). Fourth list of buterfies taken in Sikkim. Journal Foundaton for Buterfies. htp://www.ifoundbuterfies.org. of Asiatc Society of Bengal 54(2): 1–5. Mani, M.S. (1986). Buterfies of the Himalaya. Oxford & IBH Publishing de Nicéville, L. (1887). Descriptons of some new or litle-known Co. New Delhi, 181pp. buterfies from India, with some notes on the seasonal Moore, F. (1892). Lepidoptera Indica, Family Nymphalidae. Subfamilies dimorphism obtaining in the Genus Melanits. Proceedings Euploeinae and Satyrinae, Vol. 1, Lovell and Reeve, London, 277– of the Zoological Society of London 55: 448–467; htp://doi. 317pp. org/10.1111/j.1096-3642.1887.tb02988.x Rai, S., K.D. Bhuta & K. Kunte (2012). Recent sightngs of two very de Nicéville, L. (1894). A list of buterfies of Sikhim, pp. 116–176. In: rare buterfies, Lethe margaritae Elwes, 1882 and Nepts nycteus Risley, H.H. (ed.). Gazeter of Sikhim. Low Price Publicatons, Delhi. de Nicéville, 1890, from Sikkim, eastern Himalaya, India. Journal of Evans, W.H. (1927). The Identfcaton of Indian Buterfies. Bombay Threatened Taxa 4(14): 3319–3326; htp://doi.org/10.11609/JoTT. Natural History Society, Mumbai, 454pp. o2965.3319-26 Elwes, H.J. (1882). On the collecton of buterfies from Sikkim. Singh, I.J. & M.S. Chib (2015). Checklist of buterfies of Bhutan. Proceedings of the Zoological Society of London 50: 398–407; htp:// Proceedings of Bhutan Ecological Society 2: 22–58. doi.org/10.1111/j.1096-3642.1882.tb02746.x Subba, T.B. (1992). Ethnicity, State, and Development: A Case Study of Elwes, H.J. (1887). Descripton of some new Lepidoptera from Sikkim. the Gorkhaland Movement in Darjeeling. Vikas Publicaton House, Proceedings of the Zoological Society of London 55: 444–447; htp:// New Delhi, 269pp. doi.org/10.1111/j.1096-3642.1887.tb02987.x Talbot, G. (1947). The Fauna of Britsh India, including Ceylon and Burma: Buterfies, Vol. 2. Taylor and Francis, London, 506pp. Wynter-Blyth, M.A. (1957). Buterfies of the Indian Region. Bombay Natural History Society, Mumbai, 523pp.

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Journal of Threatened Taxa | www.threatenedtaxa.org | 26 May 2018 | 10(6): 11775–11779 11779 OPEN ACCESS The Journal of Threatened Taxa is dedicated to building evidence for conservaton globally by publishing peer-reviewed artcles online every month at a reasonably rapid rate at www.threatenedtaxa.org. All artcles published in JoTT are registered under Creatve Commons Atributon 4.0 Internatonal License unless otherwise mentoned. JoTT allows unrestricted use of artcles in any medium, reproducton, and distributon by providing adequate credit to the authors and the source of publicaton.

ISSN 0974-7907 (Online); ISSN 0974-7893 (Print)

May 2018 | Vol. 10 | No. 6 | Pages: 11703–11830 Date of Publicaton: 26 May 2018 (Online & Print) www.threatenedtaxa.org DOI: 10.11609/jot.2018.10.6.11703-11830

Communicatons Early stages and larval host plants of some northeastern Indian buterfies -- Tarun Karmakar, R. Nitn, Vivek Sarkar, Sarika Baidya, Subhajit Mazumder, Home range and spatal organizaton by the Hoary Fox Lycalopex vetulus V.K. Chandrasekharan, Rudraprasad Das, G.S. Girish Kumar, Swapnil (Mammalia: Carnivora: Canidae): response to social disrupton of two Lokhande, Joyce Veino, Lightson Veino, Rakoveine Veino, Zeeshan Mirza, neighboring pairs Rajesh V. Sanap, Bimal Sarkar & Krushnamegh Kunte, Pp. 11780–11799 -- Julio C. Dalponte, Herson S. Lima, Stuart Klorfne & Nelton C. da Luz, Pp. 11703–11709 Inventory of teloganodid mayfies (Ephemeroptera: Teloganodidae) from southern India with records of endemic taxa People’s attude towards wild elephants, forest conservaton and Human- -- C. Selvakumar, K.G. Sivaramakrishnan, T. Kubendran & Kailash Chandra, Elephant confict in Nilambur, southern Western Ghats of Kerala, India Pp. 11800–11805 -- C.K. Rohini, T. Aravindan, K.S. Anoop Das & P.A. Vinayan, Pp. 11710–11716

Analysis of regurgitated pellets of Spoted Owlet Athene brama Notes (Temminck, 1821) (Aves: Strigiformes: Strigidae) from Punjab, India -- Renuka Malhotra & Neena Singla, Pp. 11717–11724 Durga Das’s Leaf-nosed Bat Hipposideros durgadasi Khajuria, 1970 (Mammalia: Chiroptera: Hipposideridae): a new distributon record in Species diversity and abundance of birds on Bharathiar University Campus, northern India hidden in the Natonal Zoological Collectons Tamil Nadu, India -- M. Kamalakannan, Tauseef Hamid Dar & C. Venkatraman, Pp. 11806– -- L. Arul Pragasan & M. Madesh, Pp. 11725–11731 11811

On the taxonomy of the frst record of rare deep-water rough shark species A new range record of noctuid moth Owadaglaea elongata (Lepidoptera: of Oxynotdae (Chondrichthyes: Squaliformes) in the western Indian Ocean Noctuidae: Xyleninae) from India -- Sarah Viana & Mark W. Lisher, Pp. 11732–11742 -- P.R. Shashank & Balázs Benedek, Pp. 11812–11814

Forest evergreenness and tree endemism in the central Western Ghats, Natural history of Large Cabbage White Pieris brassicae nepalensis southern India Gray, 1846 (Lepidoptera: Pieridae) on Nasturtum, Tropaeolum majus -- Divakar K. Mesta & Ganesh R. Hegde, Pp. 11743–11752 (Tropaeolaceae) in Utarakhand, India -- Bhawana Kapkot Negi & Ravindra K. Joshi, Pp. 11815–11817 Distributon of Rhododendron falconeri Hook. F. (Ericales: Ericaceae) in Yuksam-Dzongri trekking corridor of Khangchendzonga Natonal Park, An account of the occurrence of Wedge Sea Hare Dolabella auricularia Sikkim, India (Lightoot, 1786) (Gastropoda: Aplysiidae) from Andaman Islands, India -- Aseesh Pandey & Hemant K. Badola, Pp. 11753–11759 -- Vikas Pandey, Ganesh Thiruchitrambalam, M. Savurirajan, Raj Kiran Lakra, Jawed Equbal, Kunal Satyam, P. Shanmukha Sainath & Rokkarukala Samson, Pp. 11818–11821 Peer Commentary New pteridophytc records from Mizoram, northeastern India The characteristcs, representatveness, functon and conservaton -- Sachin Sharma, Amit Kumar, Bhupendra Singh Kholia & Surendra Singh importance of tropical dry evergreen forest on India’s Coromandel Coast Bargali, Pp. 11822–11826 -- Mark Everard, Pp. 11760–11769 Clarke’s Morning Glory Ipomoea clarkei Hook.f. (Convolvulaceae): additon to the fora of Eastern Ghats Short Communicatons -- L. Rasingam, J. Swamy & M. Sankara Rao, Pp. 11827–11829

Mugger Crocodile Crocodylus palustris Lesson, 1831 (Reptlia: Crocodilia: Crocodylidae) in river Saberi of Godavari system in southern Odisha, India: Miscellaneous conservaton implicatons -- Subrat Debata, Swetashree Purohit, Anirban Mahata, Sudheer Kumar Jena Natonal Biodiversity Authority & Sharat Kumar Palita, Pp. 11770–11774 Member A new record of the lesser-known buterfy Small Woodbrown Lethe nicetella de Nicéville, 1887 (Lepidoptera: Nymphalidae: Satyrinae) from Khangchendzonga Natonal Park, Sikkim, India -- Sailendra Dewan, Bhoj Kumar Acharya & Sudeep Ghatani, Pp. 11775–11779

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