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Theropod Composition of Early Late Cretaceous Faunas from Central
CORE Metadata, citation and similar papers at core.ac.uk Provided by Repository of the Academy's Library 1 Feeding related characters in basal pterosaurs: implications for jaw mechanism, dental function and diet RH: Feeding related characters in pterosaurs Attila Ősi A comparative study of various feeding related features in basal pterosaurs reveals a significant change in feeding strategies during the early evolutionary history of the group. These features are related to the skull architecture (e.g. quadrate morphology and orientation, jaw joint), dentition (e.g. crown morphology, wear patterns), reconstructed adductor musculature, and postcranium. The most basal pterosaurs (Preondactylus, dimorphodontids and anurognathids) were small bodied animals with a wing span no greater than 1.5 m, a relatively short, lightly constructed skull, straight mandibles with a large gape, sharply pointed teeth and well developed external adductors. The absence of extended tooth wear excludes complex oral food processing and indicates that jaw closure was simply orthal. Features of these basalmost forms indicate a predominantly insectivorous diet. Among stratigraphically older but more derived forms (Eudimorphodon, Carniadactylus, Caviramus) complex, multicusped teeth allowed the consumption of a wider variety of prey via a more effective form of food processing. This is supported by heavy dental wear in all forms with multicusped teeth. Typical piscivorous forms occurred no earlier than the Early Jurassic, and are characterized by widely spaced, enlarged procumbent teeth forming a fish grab and an anteriorly inclined quadrate that permitted only a relatively small gape. In addition, the skull became more elongate and body size 2 increased. Besides the dominance of piscivory, dental morphology and the scarcity of tooth wear reflect accidental dental occlusion that could have been caused by the capturing or seasonal consumption of harder food items. -
SG125 035-140 Veldmeijer 16-01-2007 07:46 Pagina 35
SG125 035-140 veldmeijer 16-01-2007 07:46 Pagina 35 Description of Coloborhynchus spielbergi sp. nov. (Pterodactyloidea) from the Albian (Lower Cretaceous) of Brazil. André J. Veldmeijer Veldmeijer, A.J. Coloborhynchus spielbergi sp. nov. (Pterodactyloidea) from the Albian (Lower Cretaceous) of Brazil. Scripta Geologica 125: 35-139, 22 figs., 16 pls; Leiden, May 2003. André J. Veldmeijer, Mezquitalaan 23, 1064 NS Amsterdam, The Netherlands ([email protected]). A new species of pterosaur, Coloborhynchus spielbergi sp. nov. (Pterodactyloidea), from the Romualdo Member (Albian) of the Santana Formation is described. The type consists of the skull, mandible and many of the post-cranial bones. The specimen displays a high degree of co-ossification indicating that the animal was an adult and likely quite old when it died. The wingspan is reconstructed at nearly 6 m. Among the characteristic features are a large anteriorly positioned premaxillary sagittal crest and a smaller, also anteriorly positioned dentary sagittal crest, a flat anterior aspect of the skull from which two teeth project and a ventrally fused pelvis. Comments on Brazilian pterosaurs are made in connec- tion with the classificiation of the Leiden specimen. Keywords –– Pterosaur, Coloborhynchus, Santana Formation, Lower Cretaceous, Brazil. Contents Introduction ..................................................................................................................................................... 35 Material ............................................................................................................................................................. -
On Two Pterosaur Humeri from the Tendaguru Beds (Upper Jurassic, Tanzania)
“main” — 2009/10/20 — 22:40 — page 813 — #1 Anais da Academia Brasileira de Ciências (2009) 81(4): 813-818 (Annals of the Brazilian Academy of Sciences) ISSN 0001-3765 www.scielo.br/aabc On two pterosaur humeri from the Tendaguru beds (Upper Jurassic, Tanzania) FABIANA R. COSTA and ALEXANDER W.A. KELLNER Museu Nacional, Universidade Federal do Rio de Janeiro, Departamento de Geologia e Paleontologia Quinta da Boa Vista s/n, São Cristóvão, 20940-040 Rio de Janeiro, RJ, Brasil Manuscript received on August 17, 2009; accepted for publication on September 30, 2009; contributed by ALEXANDER W.A. KELLNER* ABSTRACT Jurassic African pterosaur remains are exceptionally rare and only known from the Tendaguru deposits, Upper Jurassic, Tanzania. Here we describe two right humeri of Tendaguru pterosaurs from the Humboldt University of Berlin: specimens MB.R. 2828 (cast MN 6661-V) and MB.R. 2833 (cast MN 6666-V). MB.R. 2828 consists of a three- dimensionally preserved proximal portion. The combination of the morphological features of the deltopectoral crest not observed in other pterosaurs suggests that this specimen belongs to a new dsungaripteroid taxon. MB.R. 2833 is nearly complete, and because of a long and round proximally placed deltopectoral crest it could be referred to the Archaeopterodactyloidea. It is the smallest pterosaur from Africa and one of the smallest flying reptiles ever recorded. These specimens confirm the importance of the Tendaguru deposits for the Jurassic pterosaur record. This potential, however, has to be fully explored with more field work. Key words: Tendaguru, Tanzania, Africa, Upper Jurassic, Pterosauria. INTRODUCTION in providing isolated remains up to now (Kellner and Mader 1997, Wellnhofer and Buffetaut 1999, Mader Africa shows a great potential for pterosaur material and Kellner 1999). -
Analyzing Pterosaur Ontogeny and Sexual Dimorphism with Multivariate Allometry Erick Charles Anderson [email protected]
Marshall University Marshall Digital Scholar Theses, Dissertations and Capstones 2016 Analyzing Pterosaur Ontogeny and Sexual Dimorphism with Multivariate Allometry Erick Charles Anderson [email protected] Follow this and additional works at: http://mds.marshall.edu/etd Part of the Animal Sciences Commons, Ecology and Evolutionary Biology Commons, and the Paleontology Commons Recommended Citation Anderson, Erick Charles, "Analyzing Pterosaur Ontogeny and Sexual Dimorphism with Multivariate Allometry" (2016). Theses, Dissertations and Capstones. 1031. http://mds.marshall.edu/etd/1031 This Thesis is brought to you for free and open access by Marshall Digital Scholar. It has been accepted for inclusion in Theses, Dissertations and Capstones by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected], [email protected]. ANALYZING PTEROSAUR ONTOGENY AND SEXUAL DIMORPHISM WITH MULTIVARIATE ALLOMETRY A thesis submitted to the Graduate College of Marshall University In partial fulfillment of the requirements for the degree of Master of Science in Biological Sciences by Erick Charles Anderson Approved by Dr. Frank R. O’Keefe, Committee Chairperson Dr. Suzanne Strait Dr. Andy Grass Marshall University May 2016 i ii ii Erick Charles Anderson ALL RIGHTS RESERVED iii Acknowledgments I would like to thank Dr. F. Robin O’Keefe for his guidance and advice during my three years at Marshall University. His past research and experience with reptile evolution made this research possible. I would also like to thank Dr. Andy Grass for his advice during the course of the research. I would like to thank my fellow graduate students Donald Morgan and Tiffany Aeling for their support, encouragement, and advice in the lab and bar during our two years working together. -
Klinghardt, F. (1941) Beobachtungen an Flugsauriern. Paläontologische Zeitschrift , 4, 250- 258
Klinghardt, F. (1941) Beobachtungen an Flugsauriern. Paläontologische Zeitschrift , 4, 250- 258. Observations on Pterosaurs by F. Klinghardt, Berlin, 1941 (with 1 text figure and plates 14-16) Preliminary Remarks Interest in pterosaurs has always been great and it has become even greater since BROILI (A Dorygnathus with Skin Fossils, Sitz. Ber. d. Bayr. Akad. d. Wissensch., Jahrg. 1939, p. 129) was able to prove the presence of hair in these animals. DÖDERLEIN (On Rhamphorhynchus etc. On Anurognathus etc. A Pterodactylus with throat pouch and swimming membrane. Ebenda 1929, p. 1-175) gave us information regarding the throat pouch and a large number of particular anatomical points. However, many questions have to be answered and O. KUHN is quite correct when he writes in The Fossil Reptiles p. 119: "In the structure of the skull many close analogies with birds can be observed and these certainly merit further investigation." The conclusion that these most specialised of all reptiles were "warm- blooded" would certainly be rather precipitate. A Pterodactylus scolopaciceps preserved in almost natural position Herm. v. Meyer (Pl. 14, fig. 1 and 2) (Original in Berlin Geological-Palaeontological Institute and Museum) Preliminary Remarks: This specimen is very well preserved. In this paper the organs which seem particularly important to the author are emphasised. The specimen was found in the White Jura of Solnhofen. The Preparation: On the whole, excellent. Above the braincase, level with the occiput, it has unfortunately been scratched; perhaps there are also preparation faults in the neck area, so that the originally present cervical skin was removed accidentally. -
Is Our Understanding of Santana Group Pterosaur Diversity Biased by Poor Biological and Stratigraphic Control?
Anhanguera taxonomy revisited: is our understanding of Santana Group pterosaur diversity biased by poor biological and stratigraphic control? Felipe L. Pinheiro1 and Taissa Rodrigues2 1 Laboratório de Paleobiologia, Universidade Federal do Pampa, São Gabriel, RS, Brazil 2 Departamento de Ciências Biológicas, Universidade Federal do Espírito Santo, Vitória, ES, Brazil ABSTRACT Background. Anhanguerids comprise an important clade of pterosaurs, mostly known from dozens of three-dimensionally preserved specimens recovered from the Lower Cretaceous Romualdo Formation (northeastern Brazil). They are remarkably diverse in this sedimentary unit, with eight named species, six of them belonging to the genus Anhanguera. However, such diversity is likely overestimated, as these species have been historically diagnosed based on subtle differences, mainly based on the shape and position of the cranial crest. In spite of that, recently discovered pterosaur taxa represented by large numbers of individuals, including juveniles and adults, as well as presumed males and females, have crests of sizes and shapes that are either ontogenetically variable or sexually dimorphic. Methods. We describe in detail the skull of one of the most complete specimens referred to Anhanguera, AMNH 22555, and use it as a case study to review the diversity of anhanguerids from the Romualdo Formation. In order to accomplish that, a geometric morphometric analysis was performed to assess size-dependent characters with respect to the premaxillary crest in the 12 most complete skulls bearing crests that are referred in, or related to, this clade, almost all of them analyzed first hand. Results. Geometric morphometric regression of shape on centroid size was highly Submitted 4 January 2017 statistically significant (p D 0:0091) and showed that allometry accounts for 25.7% Accepted 8 April 2017 Published 4 May 2017 of total shape variation between skulls of different centroid sizes. -
Pterosaur Distribution in Time and Space: an Atlas 61
Zitteliana An International Journal of Palaeontology and Geobiology Series B/Reihe B Abhandlungen der Bayerischen Staatssammlung für Pa lä on to lo gie und Geologie B28 DAVID W. E. HONE & ERIC BUFFETAUT (Eds) Flugsaurier: pterosaur papers in honour of Peter Wellnhofer CONTENTS/INHALT Dedication 3 PETER WELLNHOFER A short history of pterosaur research 7 KEVIN PADIAN Were pterosaur ancestors bipedal or quadrupedal?: Morphometric, functional, and phylogenetic considerations 21 DAVID W. E. HONE & MICHAEL J. BENTON Contrasting supertree and total-evidence methods: the origin of the pterosaurs 35 PAUL M. BARRETT, RICHARD J. BUTLER, NICHOLAS P. EDWARDS & ANDREW R. MILNER Pterosaur distribution in time and space: an atlas 61 LORNA STEEL The palaeohistology of pterosaur bone: an overview 109 S. CHRISTOPHER BENNETT Morphological evolution of the wing of pterosaurs: myology and function 127 MARK P. WITTON A new approach to determining pterosaur body mass and its implications for pterosaur fl ight 143 MICHAEL B. HABIB Comparative evidence for quadrupedal launch in pterosaurs 159 ROSS A. ELGIN, CARLOS A. GRAU, COLIN PALMER, DAVID W. E. HONE, DOUGLAS GREENWELL & MICHAEL J. BENTON Aerodynamic characters of the cranial crest in Pteranodon 167 DAVID M. MARTILL & MARK P. WITTON Catastrophic failure in a pterosaur skull from the Cretaceous Santana Formation of Brazil 175 MARTIN LOCKLEY, JERALD D. HARRIS & LAURA MITCHELL A global overview of pterosaur ichnology: tracksite distribution in space and time 185 DAVID M. UNWIN & D. CHARLES DEEMING Pterosaur eggshell structure and its implications for pterosaur reproductive biology 199 DAVID M. MARTILL, MARK P. WITTON & ANDREW GALE Possible azhdarchoid pterosaur remains from the Coniacian (Late Cretaceous) of England 209 TAISSA RODRIGUES & ALEXANDER W. -
New Information on the Tapejaridae (Pterosauria, Pterodactyloidea) and Discussion of the Relationships of This Clade
AMEGHINIANA (Rev. Asoc. Paleontol. Argent.) - 41 (4): 521-534. Buenos Aires, 30-12-2004 ISSN 0002-7014 New information on the Tapejaridae (Pterosauria, Pterodactyloidea) and discussion of the relationships of this clade Alexander Wilhelm Armin KELLNER1 Abstract. A phylogenetic analysis indicates that the Tapejaridae is a monophyletic group of pterodactyloid pterosaurs, diagnosed by the following synapomorphies: premaxillary sagittal crest that starts at the anterior tip of the premaxilla and extends posteriorly after the occipital region, large nasoantorbital fenestra that reaches over 45% of the length between premaxilla and squamosal, lacrimal process of the jugal thin, distinct small pear- shaped orbit with lower portion narrow, and broad tubercle at the ventroposterior margin of the coracoid. Several cranial and postcranial characters indicate that the Tapejaridae are well nested within the Tapejaroidea, in sister group relationship with the Azhdarchidae. A preliminary study of the ingroup relationships within the Tapejaridae shows that Tupuxuara is more closely related to Thalassodromeus relative to Tapejara. At present tape- jarid remains have been found in the following deposits: Crato and Romualdo members of the Santana Formation (Aptian-Albian), Araripe Basin, Brazil; Jiufotang Formation (Aptian), Jehol Group of western Liaoning, China; and in the redbeds (Cenomanian) of the Kem Kem region, Morocco. An incomplete skull found in the Javelina Formation (Maastrichtian), Texas also shows several tapejarid features and might be a member of this clade. Although information is still limited, the present distribution of the Tapejaridae indicates that this clade of pterosaurs was not exclusive of Gondwana, and was more widespread than previously known. Resumen. NUEVA INFORMACIÓN SOBRE LOS TAPEJARIDAE (PTEROSAURIA, PTERODACTYLOIDEA) Y DISCUSIÓN SOBRE LAS RELACIONES DE ESTE CLADO. -
On the Osteology of Tapejara Wellnhoferi KELLNER 1989 and the first Occurrence of a Multiple Specimen Assemblage from the Santana Formation, Araripe Basin, NE-Brazil
Swiss J Palaeontol (2011) 130:277–296 DOI 10.1007/s13358-011-0024-5 On the osteology of Tapejara wellnhoferi KELLNER 1989 and the first occurrence of a multiple specimen assemblage from the Santana Formation, Araripe Basin, NE-Brazil Kristina Eck • Ross A. Elgin • Eberhard Frey Received: 28 May 2011 / Accepted: 9 August 2011 / Published online: 26 August 2011 Ó Akademie der Naturwissenschaften Schweiz (SCNAT) 2011 Abstract The postcranial elements of two similar sized ocular lobes indicate that Tapejara possessed both excel- and juvenile individuals, along with a partial skull, are lent balancing and visual systems as a consequence of its attributed to the Early Cretaceous pterosaur Tapejara aerial lifestyle. wellnhoferi. The remains, recovered from a single con- cretion of the Romualdo Member, Santana Formation, Keywords Brazil Á Lower Cretaceous Á Santana NE-Brazil, represent the first account of multiple specimens Formation Á Pterosauria Á Tapejaridae Á Osteology having settled together and allow for a complete review of postcranial osteology in tapejarid pterosaurs. A comparison Abbreviations of long bone morphometrics indicates that all specimens BSP Bayerische Staatammlung fu¨r Pala¨ontologie und attributed to the Tapejaridae for which these elements are historische Geologie, Munich, Germany known (i.e. Huaxiapterus, Sinopterus, Tapejara) display D Dalian Natural History Museum, Dalian, China similar bivariate ratios, suggesting that Chinese and Bra- IMNH Iwaki City Museum of Coal and Fossils, Iwaki, zilian taxa must have exhibited similar growth patterns. An Japan unusual pneumatic configuration, whereby the humerus is IVPP Institute for Vertebrate Palaeontology and pierced by both dorsally and ventrally located foramina, is Palaeoanthropology Beijing, P. -
New Ctenochasmatid Pterosaur Found from the Lower Cretaceous of China 16 December 2011
New Ctenochasmatid Pterosaur found from the lower cretaceous of China 16 December 2011 unearthed. Five genera and species of ctenochasmatids has been reported before in China: Huanhepterus quingyangensis, Beipiaopterus chenianus, Cathayopterus grabaui, Gegepterus changae and Elanodactylus prolatus. Beipiaopterus and Elanodactylus are known only postcranial skeletons, whereas Huanhepterus, Cathayopterus and Gegepterus have partial or nearly complete skulls. Pterofiltrus qiui differs from other ctenochasmatid Fig. 1: Pterofiltrus qiui gen. et sp. nov., IVPP V12339; pterosaurs by the following combination of photograph (A); line drawing (B) showing the general characters: about 112 teeth in total (including the position of the elements; their relationship (C), blue, upper and lower jaws); the dentition occupies more rostrum; green, mandible; red, posterior part of the skull; than 50% of the skull length; the anterior teeth vary white, indeterminate. Credit: JIANG Shunxing in size; the mandibular symphysis is longer than half of the whole mandible length; in ventral view, an apparent symphyseal trough in the median part of the symphysis. Drs. JIANG Shunxing and WANG Xiaolin from Institute of Vertebrate Paleontology and "Pterofiltrus qiui is assigned to the Paleoanthropology (IVPP), Chinese Academy of Ctenochasmatidae based on the extremely Sciences, described a new ctenochasmatid elongated rostrum, a large number of slender teeth, pterosaur, Pterofiltus qiui gen. et sp. nov., from the and the upper anterior teeth inclined Lower Cretaceous deposits of Liaoning, China, as ventroanteriorly", said WANG Xiaolin, coauthor and reported in Annals of the Brazilian Academy of research designer, "The Jehol Biota comprises Sciences (2011) 83(4): 1243-1249, providing many pterosaurs and ctenochasmatids play a very further information on the global distribution of the important role. -
Redalyc.On Two Pterosaur Humeri from the Tendaguru Beds (Upper Jurassic, Tanzania)
Anais da Academia Brasileira de Ciências ISSN: 0001-3765 [email protected] Academia Brasileira de Ciências Brasil COSTA, FABIANA R.; KELLNER, ALEXANDER W.A. On two pterosaur humeri from the Tendaguru beds (Upper Jurassic, Tanzania) Anais da Academia Brasileira de Ciências, vol. 81, núm. 4, diciembre, 2009, pp. 813-818 Academia Brasileira de Ciências Rio de Janeiro, Brasil Available in: http://www.redalyc.org/articulo.oa?id=32713481017 How to cite Complete issue Scientific Information System More information about this article Network of Scientific Journals from Latin America, the Caribbean, Spain and Portugal Journal's homepage in redalyc.org Non-profit academic project, developed under the open access initiative “main” — 2009/10/20 — 22:40 — page 813 — #1 Anais da Academia Brasileira de Ciências (2009) 81(4): 813-818 (Annals of the Brazilian Academy of Sciences) ISSN 0001-3765 www.scielo.br/aabc On two pterosaur humeri from the Tendaguru beds (Upper Jurassic, Tanzania) FABIANA R. COSTA and ALEXANDER W.A. KELLNER Museu Nacional, Universidade Federal do Rio de Janeiro, Departamento de Geologia e Paleontologia Quinta da Boa Vista s/n, São Cristóvão, 20940-040 Rio de Janeiro, RJ, Brasil Manuscript received on August 17, 2009; accepted for publication on September 30, 2009; contributed by ALEXANDER W.A. KELLNER* ABSTRACT Jurassic African pterosaur remains are exceptionally rare and only known from the Tendaguru deposits, Upper Jurassic, Tanzania. Here we describe two right humeri of Tendaguru pterosaurs from the Humboldt University of Berlin: specimens MB.R. 2828 (cast MN 6661-V) and MB.R. 2833 (cast MN 6666-V). MB.R. -
Universidade Federal Do Rio Grande Do Sul Instituto De Geociências Programa De Pós-Graduação Em Geociências Contribuição
UNIVERSIDADE FEDERAL DO RIO GRANDE DO SUL INSTITUTO DE GEOCIÊNCIAS PROGRAMA DE PÓS-GRADUAÇÃO EM GEOCIÊNCIAS CONTRIBUIÇÃO AO CONHECIMENTO DOS PTEROSSAUROS DO GRUPO SANTANA (CRETÁCEO INFERIOR) DA BACIA DO ARARIPE, NORDESTE DO BRASIL FELIPE LIMA PINHEIRO ORIENTADOR – Prof. Dr. Cesar Leandro Schultz Porto Alegre - 2014 UNIVERSIDADE FEDERAL DO RIO GRANDE DO SUL INSTITUTO DE GEOCIÊNCIAS PROGRAMA DE PÓS-GRADUAÇÃO EM GEOCIÊNCIAS CONTRIBUIÇÃO AO CONHECIMENTO DOS PTEROSSAUROS DO GRUPO SANTANA (CRETÁCEO INFERIOR) DA BACIA DO ARARIPE, NORDESTE DO BRASIL FELIPE LIMA PINHEIRO ORIENTADOR – Prof. Dr. Cesar Leandro Schultz BANCA EXAMINADORA Prof. Dr. Marco Brandalise de Andrade – Faculdade de Biociências, PUC, RS Profa. Dra. Marina Bento Soares – Departamento de Paleontologia e Estratigrafia, UFRGS Profa. Dra. Taissa Rodrigues – Departamento de Biologia, UFES, ES Tese de Doutorado apresentada ao Programa de Pós-Graduação em Geociências como requisito parcial para a obtenção do título de Doutor em Ciências. Porto Alegre – 2014 “Ao ser destampado pelo gigante, o cofre deixou escapar um hálito glacial. Dentro havia apenas um enorme bloco transparente, com infinitas agulhas internas nas quais se despedaçava em estrelas de cores a claridade do crepúsculo. Desconcertado, sabendo que os meninos esperavam uma explicação imediata, José Arcadio Buendía atreveu-se a murmurar: – É o maior diamante do mundo.” Gabriel García Marquez AGRADECIMENTOS Um trabalho como esse não é feito apenas a duas mãos. Durante o percurso de meu mestrado e doutorado, tive o privilégio de contar com o apoio (por vezes, praticamente incondicional) de diversas pessoas. Em primeiro lugar, pelo apoio irrestrito em todos os momentos, agradeço a minha família, em especial a meus pais, Sandra e Valmiro e a meus irmãos, Fernando e Sacha.