Annotated Checklist of Hungarian Bryophytes

Studia bot. hung. 35, pp. 91–149, 2004

ANNOTATED CHECKLIST OF HUNGARIAN BRYOPHYTES

P. ERZBERGER1 and B. PAPP2

1Belziger Str. 37, D-10823 Berlin, Germany; E-mail: [email protected] 2Department of Botany, Hungarian Natural History Museum H-1476 Budapest, Pf. 222, Hungary; E-mail: [email protected]

The checklist presented below contains 629 bryophyte taxa in 199 genera, 2 species of hornworts in 2 genera, 141 species and 2 additional subspecies of liverworts in 50 genera and 483 species and 1 additional subspecies of mosses in 147 genera. It is the first inventory of Hungarian bryophytes in twenty years. Not all problems resulting from changes in taxonomy could be resolved yet, but the fields where further research is necessary are pointed out. Separate lists of excluded (46), doubtful (42) and recently added taxa (30) are provided. The annotations comment on taxonomic and nomenclatural changes, quote specimen details or contain otherwise relevant information.

Key words: bryophytes, checklist, Hungary

INTRODUCTION

This is the first attempt to prepare an up-dated checklist of all bryophyte taxa of Hungary, more than twenty years after the publication of the “Handbook of the Hungarian Bryoflora” (ORBÁN and VAJDA 1983). Since then, partly as a result of fieldwork and partly resulting from herbarium revision, several new species have been added to the Hungarian bryoflora. Other taxa, some reported in ORBÁN and VAJDA (1983) and many in numerous other papers as occurring in Hungary, are now considered to be absent from Hungary, due to the application of better descriptions of taxa or to the amendment of various errors, especially with respect to geography. The species concerned are listed in the appendices. A considerable number of taxonomic and nomenclatural changes have accumulated within the last two decades, and thus the need to replace the somewhat out-of-date ORBÁN and VAJDA (1983) by a modern checklist has been increasingly felt. The application of national and international legislation with respect to the conservation of nature must be founded on an inventory of species that reflects the present state. This kind of inventory is also a prerequisite for the preparation of a red data book of bryophytes satisfying modern criteria (IUCN 1994). One of the most serious problems of an up-dated treatment of Hungarian bryophyte taxa lies in the difficulties of interpretation of old reports, in cases when the concepts of taxa have changed in the meantime. This concerns a considerable proportion of the Hungarian bryoflora. On the other hand, Hungarian bryology, fortunately, is in possession of a large body of herbarium specimens collected by

Studia Botanica Hungarica 35, 2004 Hungarian Natural History Museum, Budapest 92 ERZBERGER, P. and PAPP, B.

BOROS and VAJDA during long decades of the last century, and now to a large part housed in the bryophyte collections of Budapest (BP), Eger (EGR) and Szom- bathely (SZO), with vouchers for almost every one of the records published by these two outstanding personalities of Hungarian 20th century bryology. In princi- ple, therefore, the problems due to progress in taxonomy can be overcome by herbarium revision. However, only few genera or species groups have been studied in this respect up to now (e.g. Tortula Sect. Rurales:TÓTH (1986, 1987), Barbula, Didymodon:GALAMBOS (1992), Grimmia:MAIER,PAPP and ERZBERGER, unpubl., Hedwigia, Dicranum tauricum / viride, Ditrichum crispatissimum / flexicaule, Funaria muhlenbergii / pulchella:ERZBERGER (1996, 1999, 2001, 2002), Pohlia Sect. Pohliella:ERZBERGER (in press), Sphagnum:SZURDOKI (2003)). Especially some large and/or difficult genera like Bryum, Racomitrium and Schistidium are in bad need of revision, and our results with respect to these are far from definite. For- tunately, on the other hand, Hungarian specimens have been included in world- wide or European revisions of some genera, e.g. Aloina (DELGADILLO 1975), Grim- mia (MUÑOZ and PANDO 2000), Schistidium (BLOM 1996), Seligeria (GOS and OCHYRA 1994) and the species of the Calliergon-Scorpidium-Drepanocladus complex (HEDENÄS 2003). This means a substantial contribution to the solution of the problems outlined above. In order to facilitate future research and as a first step towards a specimen- based Hungarian flora, we quote specimen details (herbarium, inventory number, locality, date and collector) for virtually all taxa that are known from a single locality or a few sites only. The taxonomy of hornworts and liverworts follows the recently published checklists of GROLLE and LONG (2000) and SÖDERSTRÖM et al. (2002). For mosses, no comparable comprehensive modern taxonomic treatment is available, therefore, in general, we follow CORLEY et al. (1981) and CORLEY and CRUND- WELL (1991), with the exception of some genera listed below together with the publications on which we rely (Table 1). We have not adopted the more radical changes proposed by ZANDER (1993) in Pottiaceae and HEDENÄS (2003) in Am- blystegiaceae and other pleurocarpous mosses, because, contrary to BLOCKEEL and LONG (1998), we feel that perhaps more research is needed to corroborate these concepts. In the interest of nomenclatural stability, we rather followed the more moderate amendments of nomenclature published in several German reference works (LUDWIG et al. 1996, KOPERSKI et al. 2000, NEBEL and PHILIPPI 2000, 2001). For the preparation of the checklist, published reports on the occurrence of taxa in Hungary were evaluated from the following main sources, with critical reference to specimens where appropriate: BOROS (1968), ORBÁN and VAJDA

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Table 1. Taxonomic references for some moss genera.

Aloina DELGADILLO (1975)

Bryum HODGETTS (2001)

Dichodontium WERNER (2002)

Ditrichum FRISVOLL (1985), ERZBERGER (2001)

Drepanocladus aduncus group HEDENÄS (2003)

Grimmia GREVEN (1995), MAIER (2002a, b), MAIER and GEISSLER (1995), MUÑOZ (1998a, b), MUÑOZ and PANDO (2000)

Mnium KOPONEN (1994)

Orthotrichum LEWINSKY (1993), LEWINSKY-HAAPASAARI (1998), LUDWIG et al. (1996), MAZIMPAKA et al. (2000), VITT (1973)

Plagiothecium MASTRACCI and SAUER (2001)

Pterygoneurum CANO et al. (1994), FREY et al. (1990), PÓCS (1999), SEGARRA et al. (1998)

Racomitrium FRISVOLL (1983), BEDNAREK-OCHYRA (1995)

Schistidium BLOM (1996, 1998)

Tortula NEBEL and HEINRICHS (2000), TÓTH (1986, 1987), WERNER and HÉBRARD (1986)

Ulota SMITH and PROCTOR (1993), ERZBERGER (2003)

(1983), DÜLL (1983, 1984, 1985, 1992), SÖDERSTRÖM et al. (2002). Numerous smaller papers with floristical data on Hungarian bryophytes were evaluated as well and are referenced in the annotations. We have compiled species and – in a few cases – subspecies of bryophytes, but not varieties. Nevertheless, nearly all reports on varieties in BOROS (1968) and ORBÁN and VAJDA (1983) and some other sources are referenced in the annotations. Each entry begins with the accepted name of the taxon (in bold face), and au- thorities in standardised abbreviations (BRUMMIT and POWELL 1992). In brackets follow synonyms, if any. In general, only the most important synonyms are given, to facilitate the use of current bryological works. If names in ORBÁN and VAJDA (1983) and BOROS (1968) differ from the accepted name in this checklist, these differing names are always the first synonyms given. In some cases, additional synonyms used in English and German floras (e.g.SMITH 1978, 1990, FRAHM and FREY 1992, FREY et al. 1995) or even in taxonomic treatments that might become influential in the future (e.g.HEDENÄS 2003, ZANDER 1993) are included to facilitate the use of the checklist. It must be emphasised that synonyms are listed here

Studia bot. hung. 35, 2004 94 ERZBERGER, P. and PAPP, B. for practical purposes; they may not comply with the rules of the International Code of Botanical Nomenclature (GREUTER et al. 2000), and in some but not all cases this is stated in a note. Annotations highlight nomenclatural, taxonomic or other questions of interest and quote the literature. We treat the three groups of bryophytes: Anthocerotae, Hepaticae and Musci separately. After the main list – i.e. the checklist proper – we have compiled some separate lists. Appendix 1 contains the taxa excluded from the Hungarian flora (7 hep., 39 musc.), because some kind of error could be clearly demonstrated. Appen- dix 2 is a list of doubtful species (1 hep., 29 musc.) not included in the main list, because we feel that their inclusion would probably be erroneous, but evidence for this is less clear than for the excluded taxa in App. 1, e.g. doubtful taxa are included here, if no specimen could be located. Taxa in Apps 1 and 2 that appeared in ORBÁN and VAJDA (1983) are marked with an asterisk (3 hep., 16 musc.). Appen- dix 3 contains taxa that are less doubtful, where our reservations do not warrant ex- clusion from the main list (6 hep., 6 musc.). We are confident that a large proportion of these taxa is in fact a member of the Hungarian bryoflora. To facilitate comparison with ORBÁN and VAJDA (1983), in Appendix 4 a list of taxa recorded after 1983 is provided (2 hep., 28 musc.). App. 5 is a table of useful synonyms. The deci- sion to place particular taxa in any one of these lists is based on the evidence available at the time of compilation. It is hoped that in a later edition of the checklist these problems will be solved due to progress in herbarium and field research.

* * * Acknowledgements – We thank the directors and curators of the following herbaria for the ar- rangement of loans of specimens: B, EGR, JE, SZO. We are especially grateful to E. MAIER, Geneva, for revising a large number of Grimmia specimens, to J. KLAWITTER, Berlin, J. KUČERA, České Budejovice and TH.HOMM,ELSFLETH, for the revision of some specimens, and to L. BALOGH,T. PÓCS and E. SZURDOKI for valuable comments. Part of this work was supported by the Institute of Environmental Management, Hungary.

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ORBÁN, S. (1991): A mohakutatás alakulása hazánkban az utóbbi évtizedekben. (Decades in the Hungarian bryology). – Bot. Közlem. 78(Suppl.): 29–32. ORBÁN, S. and PÓCS, T. (1976): Rhodobryum ontariense (Kindb.) Kindb. in Central Europe. – Acta Bot. Acad. Sci. Hung. 22: 437–448. ORBÁN, S. and VAJDA, L. (1983): Magyarország mohaflórájának kézikönyve. (Handbook of the Hungarian Bryoflora). – Akadémiai kiadó, Budapest, 518 pp. PAPP, B. and RAJCZY, M. (1999): Bryophytes of the Kiskunság National Park and of the other parts of the Danube–Tisza Interfluve. – In: LŐKÖS, L. and RAJCZY, M. (eds): The flora of the Kiskunság National Park, Cryptogams, Magyar Természettudományi Múzeum, Budapest, pp. 363–413. PAPP, B. and ERZBERGER, P. (2000): A Villányi-hegység mohaflórája (Bryophyta). (The bryophyte flora of the Villány Hills, South Hungary). – Dunántúli Dolg. Term. Tud. Sorozat 10: 37–45. PAPP, B. and RAJCZY, M. (2000): Contributions to the knowledge of the bryophytes of the forests along the Fekete-Körös, SE Hungary. – Studia bot. hung. 30–31: 57–67. PATON, J. A. (1999): The Liverwort Flora of the British Isles. – Harley Books, Great Horkesley, 626 pp. PÓCS, T. (1999): A löszfalak virágtalan növényzete I. Orografikus sivatag a Kárpát-medencében. (Studies on the cryptogamic vegetation of loess cliffs, I. Orographic desert in the Carpathian Basin). – Kitaibelia 4(1): 143–156. PÓCS, T., GOIA, I., KIS, G., ORBÁN, S., SASS-GYARMATI,A.andVAN ZANTEN, B. O. (2002): Hilpertia velenovskyi (Schiffn.) Zander and other pottoid mosses (Bryophyta) new to Roma- nia. Studies on the cryptogamic vegetation of loess cliffs, IX. – Contributii Botanicae Grad. Bot. A. Borza Cluj-Napoca 37: 13–24. PÓCS, T., SABOVLJEVIĆ, M., PUCHE, F., SEGARRA MORAGUES, J. G., GIMENO, C. and KÜRSCHNER, H. (2004): Crossidium laxefilamentosum Frey and Kürschner (Pottiaceae), new to Europe and to North Africa. (Studies on the cryptogamic vegetation on loess cliffs, VII.) J. Bryol. 26: 113–124. PODPERA, J. (1954): Conspectus muscorum europaeorum. – Nakladatelstvi Ceskoslovenské Akade- mie Ved, Praha, 697 pp. RAJCZY, M. (1990): Mohák – Bryophyta. – In: RAKONCZAI, Z. (ed.): Vörös Könyv. Akadémiai Kiadó, Budapest, pp. 322–325. ROBINSON, H. E. and IGNATOV, M. S. (1997): Brachythecium laetum (Brid.) B. S. G. the correct name for the species commonly known as Brachythecium oxycladon. – The Bryologist 100(3): 359–361. SEGARRA, J.-G., PUCHE, F., FREY, W. and KÜRSCHNER, H. (1998): Pterygoneurum squamosum (Pottiaceae, Musci), a new moss species from Spain. – Nova Hedwigia 67(3–4): 511–515. SILLER, I. (1979): Mannia triandra (Scop.) Grolle in Hungary. – Acta Bot. Hung. 25: 139–142. SMITH, A. J. E. (1978): The Moss Flora of Britain and Ireland. – Cambridge University Press, Cam- bridge, New York, Port Chester, Melbourne, Sidney, 706 pp. SMITH, A. J. E. (1990): The Liverworts of Britain and Ireland. – Cambridge University Press, Cam- bridge, New York, Port Chester, Melbourne, Sidney, 362 pp. SMITH,A.J.E.andPROCTOR, M. C. F. (1993): Further observations on the Ulota crispa complex. – J. Hattori Bot. Lab. 74: 171–182 SÖDERSTRÖM, L., URMI, E. and VÁŇA, J. (2002): Distribution of Hepaticae and Anthocerotae in Eu- rope and Macaronesia. – Lindbergia 27: 3–47. STECH, M. and FRAHM, J.-P. (1999): Rhynchostegiella and Rhynchostegium (Brachytheciaceae, Bryopsida) based on molecular data. – Bryobrothera 5: 203–211. SZURDOKI, E. (2003): Peat mosses of North Hungary. – Studia bot. hung. 34: 55–79.

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SZURDOKI, E., TÓTH, Z. and PELLES, G. (2000): The Sphagnum populations of the Zemplén Mts, NE Hungary. – Studia bot. hung. 30–31: 113–125. THERRIEN, J. P., CRANDALL-STOTLER,B.J.andSTOTLER, R. E. (1998): Morphological and genetic variation in Porella platyphylla and P. platyphylloidea and their systematic implications. – The Bryologist 101: 1–19. TÓTH, Z. (1986): A Tortula Hedw. Sect. Rurales De Not. (Musci, Pottiaceae) rendszertani revíziója és elterjedése a Kárpát-medencében (Taxonomic revision of Tortula Hedw. sect. Rurales De Not. (Musci, Pottiaceae) and its distribution in the Carpathian Basin). – Abstracta Botanica 10: 145–185. TÓTH, Z. (1987): A phytogeographic review of Tortula Hedw. Sect. Rurales De Not. (Pottiaceae, Musci) in Hungary. – Acta Bot. Hung. 33: 249–278. VAJDA, L. (1956): Fissidens algarvicus Solms-Laubach aus dem Börzsönygebirge (Mittelungarn). – Annls hist.-nat. Mus. natn. hung. 48 (ser. n. 7): 299–301. VAJDA, L. (1958): Fissidens kosaninii Latzel dans la Montagne Börzsöny en Hongrie. – Rev. Bryol. Lichénol. 27: 49–51. VAJDA, L. (1960): Bryologische Notizen. II. – Annls hist.-nat. Mus. natn. hung. 52: 155–161. VAJDA, L. (1962): Die Verbreitung einiger seltener Leber- und Laubmoose in Ungarn. – Fragm. bot. Mus. hist.-nat. hung. 2: 23–31. VAJDA, L. (1966a): A Coscinodon cribrosus, Magyarország mohaflórájának új tagja. – Fragm. bot. Mus. hist.-nat. hung. 4(1–4): 77–78. VAJDA, L. (1966b): Die Moosflora des Börzsöny-Gebirges. – Fragm. bot. Mus. hist.-nat. hung. 4(1–4): 79–100. VAJDA, L. (1975): Bryologische Notizen IV. – Studia bot. hung. 10: 119–122. VAJDA, L. and ORBÁN, S. (1975): Vorarbeiten zur Moosflora des Semenic-Gebirges. – Studia bot. hung. 10: 115–118. VITT, D. H. (1973): A revision of the genus Orthotrichum in North America, North of Mexico. – Bryophytorum Bibliotheca 1: 1–108. VÖRÖSS,L.ZS. (1969): Mohok és harasztok az Abaligeti barlangban, villanyfényben. (Moose und Farne bei elektrischem Licht in der Höhle von Abaliget (S-Ungarn)). – Bot. Közlem. 56: 176. WERNER, J. (2002): A comparison of Dichodontium flavescens (Dicks.) Lindb. and D. pellucidum (Hedw.) Lindb. (Bryopsida). – J. Bryol. 24(3): 215–221. WERNER, J. and HÉBRARD, J. P. (1986): Tortula caninervis (Mitt.) Broth. subsp. spuria (Amann) W. Kramer var. spuria (Pottiaceae, Musci) à Touet-sur-Var (Alpes maritimes), nouveau pour la bryoflore de France. – Cryptogamie, Bryol. Lichénol. 7(4): 487–493. WILSON, P. and NORRIS, D. H. (1989): Pseudoleskeella in North America and Europe. – The Bryolo- gist 92: 387–396. ZANDER, R. H. (1993): Genera of the Pottiaceae. – Bull. Buffalo Soc. Nat. Sci. 32. 378 pp. ZANTEN,B.O.VAN (1999a): Studies on the cryptogamic vegetation of loess cliffs, II. The genus Bryum Hedw. on loess cliffs in the Pannonian Basin, including Bryum gemmiferum Wilcz. & Demar. and Bryum violaceum Crundw. et Nyh., new to Hungary. – Kitaibelia 4: 157–162. ZANTEN,B.O.VAN (1999b): Studies on the cryptogamic vegetation of loess cliffs, III. Ceratodon conicum (Hampe) Lindb., new to Hungary. – Kitaibelia 4: 163–164. ZANTEN,B.O.VAN (2000): Studies on the cryptogamic vegetation of loess cliffs, IV. Chenia leptophylla (C. Müll.) Zander, new to Hungary. – Kitaibelia 5: 271–274.

(Received: 24 August, 2004)

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CHECKLIST OF HUNGARIAN BRYOPHYTES

ANTHOCEROTAE

Anthoceros agrestis Paton (Anthoceros punctatus auct. non L.; Anthoceros crispulus auct. non (Mont.) Douin) Phaeoceros carolinianus (Michx.) Proskauer (Anthoceros laevis auct.). – Anthoceros laevis L. is not a synonym of P. carolinianus, but was confused with that species earlier.

HEPATICAE

Anastrophyllum hellerianum (Lindenb.) R. M. Schust. –ÓDOR (2000). Anastrophyllum michauxii (F. Weber) H. Buch (Sphenolobus michauxii (F. Weber) Steph.) Anastrophyllum minutum (Schreb.) R. M. Schust. (Sphenolobus minutus (Schreb.) Steph.) Aneura pinguis (L.) Dumort. (Riccardia pinguis (L.) Gray) Apometzgeria pubescens (Schrank) Kuwah. (Metzgeria pubescens (Schrank) Raddi) Asterella saccata (Wahlenb.) A. Evans (Fimbriaria saccata (Wahlenb.) Nees; Fimbriaria fragrans Nees) Athalamia hyalina (Sommerf.) S. Hatt. (Clevea hyalina (Sommerf.) Lindb.) Barbilophozia barbata (Schreb.) Loeske. – BOROS (1968) mentions “var. biloba Schiffn.” (= fo. bilobata Schiffn.), probably a modification without taxonomic value. Barbilophozia floerkei (F. Weber et D. Mohr) Loeske. – Specimens BP 48261/H and SZO. Comit. Vas. Borostyánkő. Nedves lejtőkön, szerpentin talajon. 19.05.1905. leg. and det. V. Piers, rev. A. Latzel. Bazzania trilobata (L.) Gray Blasia pusilla L. Blepharostoma trichophyllum (L.) Dumort. – subsp. trichophyllum. Calypogeia azurea Stotler et Crotz (Calypogeia trichomanis auct.). – The distinction between C. azurea and C. muelleriana has not always been made, therefore old reports or specimens of C. “trichomanis” cannot automatically be named C. azurea; old specimens may be impossible to name, when the blue pigment of oilbodies has disintegrated, because taxa vary a great deal morphologically (PATON 1999); the characters given in the key in ORBÁN and VAJDA (1983) are not always reliable. Calypogeia fissa (L.) Raddi Calypogeia integristipula Steph. (Calypogeia meylanii H. Buch). – known from a single locality; specimen: BP 29710/H Ózdi Dombvidék. Inter pag. Tarnalelesz et Borsodszentgyörgy. Ad terram Fagetorum vallis Futyó, secus vias 21.10.1974 leg. T. Pócs, det. L. Vajda. Calypogeia muelleriana (Schiffn.) Müll. Frib. Calypogeia neesiana (C. Massal. et Carestia) Müll. Frib. Calypogeia suecica (Arnell et J. Perss.) Müll. Frib. Cephalozia bicuspidata (L.) Dumort. (Cephalozia lammersiana (Huebener) F. Lees). – taxonomy controversial, C. bicuspidata var. lammersiana is not recognised in PATON (1999) or BLOCKEEL and LONG (1998), here placed into synonymy with C. bicuspidata due to difficulties in differentiation (KOPERSKI et al. 2000), but recognized as species (BOROS (1968), ORBÁN and VAJDA (1983)), subspecies or variety by other authors (e.g. DAMSHOLT 2002) Cephalozia catenulata (Huebener) Lindb.

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Cephalozia lacinulata J. B. Jack ex Spruce. – DÜLL (1983), Ann. 24: “C. lacinulata is reported from three localities near the Hungarian–Austrian border. One of these is in Hungary now (“oberer Walkgraben”, leg. Latzel). Because the collector was a good taxonomist, Vajda and Rajczy regard this species as a member of the Hungarian flora though no specimen has been seen.” The specimen in BP (9256/H Com. Castriferrei Hámortó: Im unteren Gössbachtal unter Ceph. pleniceps alt. ca. 380 m, Substr. Moderholz, 12.05.1896, leg. A. Latzel) was collected in Austria near the Hungarian border (cf. KIRÁLY 1996). Cephalozia lunulifolia (Dumort.) Dumort. (Cephalozia media Lindb.) Cephalozia macrostachya Kaal. – var. macrostachya; known from a single locality; specimen: BP 9268/H Com. Castriferrei. Im oberen Walkgraben zwischen Kőszeg und Hámortó alt. ca. 590 m, Substr. Moorboden, 18.05.1905, leg. A. Latzel; the collection site is near the Austrian border but still in Hungary. Cephalozia pleniceps (Austin) Lindb. Cephaloziella divaricata (Sm.) Schiffn. (Cephaloziella starkei (Funck) Schiffn.) Cephaloziella hampeana (Nees) Schiffn. – BOROS (1968) mentions var. erosa (Warnst.) Müll. Frib., but this is generally included in the synonymy of C. hampeana. Cephaloziella integerrima (Lindb.) Warnst. Cephaloziella rubella (Nees) Warnst. – var. rubella, var. sullivantii (Austin) Müll. Frib.; infraspecific taxonomy critical. Cephaloziella spinigera (Lindb.) Warnst. (Cephaloziella subdentata Warnst.). – known from a single locality, specimen: BP 21165/H Comit. Abauj-Torna. In piceetum in decl. montis Borindzás cum Leucobryum glaucum, prope pag. Telkibánya, montes Sátorhegyséeg 19.06.1960 leg. L. Vajda. Cephaloziella stellulifera (Spruce) Schiffn. Chiloscyphus pallescens (Hoffm.) Dumort. – var. pallescens, var. fragilis (Roth) Müll. Frib. Chiloscyphus polyanthos (L.) Corda. – var. polyanthos, var. rivularis (Schrad.) Gottsche, Lindenb. et Nees. Cololejeunea calcarea (Lib.) Schiffn. Cololejeunea rossettiana (C. Massal.) Schiffn. Conocephalum conicum (L.) Dumort. Diplophyllum albicans (L.) Dumort. Diplophyllum obtusifolium (Hook.) Dumort. Fossombronia foveolata Lindb. (Fossombronia dumortieri Huebener et Genth ex Lindb.). – Specimens BP 48263/H and EGR. Comit. Esztergom. Ad limites paludum Szarvasszérű II. prope montis Szentlászlóhegy supra pag. Pilisszentlászló. 26.09.1948. leg. L. Vajda. Fossombronia pusilla (L.) Nees. – BOROS (1968), ORBÁN and VAJDA (1983) include var. decipiens Corb., but this taxon is now included in the synonymy of F. pusilla (KOPERSKI et al. 2000). Fossombronia wondraczekii (Corda) Lindb. – BOROS (1968), ORBÁN and VAJDA (1983) include var. loitlesbergeri (Schiffn.) Chalaud, which is recognised or placed into synonymy with F. wondraczekii (PATON 1999). Frullania dilatata (L.) Dumort. – BOROS (1968) includes var. anomala Corb., but this taxon is now included in the synonymy of F. dilatata (KOPERSKI et al. 2000). Frullania fragilifolia (Taylor) Gottsche, Lindenb. et Nees Frullania inflata Gottsche (Frullania cleistostoma Schiffn. et Wollny; Frullania saxicola Austin) Frullania jackii Gottsche. – known from a single locality, specimens: BP 21295/H, BP 21296/H, BP 44329/H Comit. Abauj-Torna. In rupibus andesiticis umbrosis (Kutyaszorító sziklák

Studia bot. hung. 35, 2004 ANNOTATED CHECKLIST OF HUNGARIAN BRYOPHYTES 103 jobbold.) vallis rivi Osvapatak prope pag. Telkibánya, montes Sátorhegység, 17.09.1959, leg. L. Vajda. Frullania tamarisci (L.) Dumort. Gymnocolea inflata (Huds.) Dumort. – known from a single locality; representative specimen: BP 37368/H Comit. Heves. In argillosis humidis ripae lacus fontis alumen continentis “Timsós-tó” dicti in valle “Ilona-völgy” prope balneam Parádfürdő, 16.08.1931, leg. Á. Boros, also collected 19.08.1961 (BP 37369/H, BP 27746/H). Jamesoniella autumnalis (DC.) Steph. Jungermannia atrovirens Dumort. (Solenostoma triste (Nees) Müll. Frib.; Solenostoma riparia (Taylor) Dumort.) Jungermannia caespiticia Lindenb. (Solenostoma caespiticium (Lindenb.) Steph.; Haplozia caespiticia (Lindenb.) Dumort.) Jungermannia gracillima Sm. (Solenostoma crenulatum Mitt.; Haplozia crenulata (Sm.) Dumort.) Jungermannia hyalina Lyell (Plectocolea hyalina (Lyell) Mitt.) Jungermannia leiantha Grolle (Jungermannia lanceolata auct. non L.; Liochlaena lanceolata Nees; Haplozia lanceolata auct. non (L.) Dumort.) Jungermannia pumila With. (Solenostoma pumilum (With.) Müll. Frib.; Haplozia pumila (With.) Dumort.) Jungermannia sphaerocarpa Hook. (Solenostoma sphaerocarpum (Hook.) Steph.; Haplozia sphaerocarpa (Hook.) Dumort.) Jungermannia subulata A. Evans. – BOROS and VAJDA (1970), therefore only in ORBÁN and VAJDA (1983), missing in BOROS (1968). Leiocolea badensis (Gottsche) Jörg. (Lophozia badensis (Gottsche) Schiffn.) Leiocolea collaris (Nees) Schljakov (Lophozia collaris (Nees) Dumort.; Leiocolea muelleri (Lindenb.) Jörg.; Leiocolea alpestris (F. Weber) Isov.; Lophozia muelleri (Lindenb.) Dumort.) Leiocolea heterocolpos (Hartm.) H. Buch (Lophozia heterocolpos (Hartm.) M. Howe) Lejeunea cavifolia (Ehrh.) Lindb. (Lejeunea serpyllifolia Lib.). – BOROS (1968) includes var. loitlesbergeri Schiffn. Lepidozia reptans (L.) Dumort. Lophocolea bidentata (L.) Dumort. – nomenclature confused (see annotations in SÖDERSTRÖM et al. (2002), KOPERSKI et al. (2000)); if taxa are distinguished at varietal level, the correct names are as follows: L. bidentata var. bidentata (for L. cuspidata auct. non (Nees) Limpr., sensu Limpricht, BOROS (1968), ORBÁN and VAJDA (1983)) and L. bidentata var. rivularis (Raddi) Schiffn. (for L. bidentata auct. non (L.) Dumort. sensu Limpricht, BOROS (1968), ORBÁN and VAJDA (1983)); ENGEL and SCHUSTER (1985) transfer Lophocolea into the genus Chiloscyphus, their names are Chiloscyphus coadunatus (Sw.) J. J. Engel et R. M. Schust. and C. latifolius (Nees) J. J. Engel et R. M. Schust. for the above varieties, respectively. Lophocolea alata Mitt. ex Larter, treated as separate species in BOROS (1968), ORBÁN and VAJDA (1983), is today included in the synonymy of L. bidentata var. bidentata. Paton (1999) does not recognise either var. rivularis nor L. alata. Lophocolea heterophylla (Schrad.) Dumort. (Chiloscyphus profundus (Nees) J. J. Engel et R. M. Schust.) Lophocolea minor Nees (Chiloscyphus minor (Nees) J. J. Engel et R. M. Schust.) Lophozia ascendens (Warnst.) R. M. Schust. (Lophozia gracillima H. Buch) Lophozia bicrenata (Hoffm.) Dumort. (Isopaches bicrenatus (Hoffm.) H. Buch) Lophozia excisa (Dicks.) Dumort. – BOROS (1968), ORBÁN and VAJDA (1983) mention f. limprichtii (Lindb.) C. Massal. which is a synyonym of var. excisa (DAMSHOLT 2002).

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Lophozia incisa (Schrad.) Dumort. Lophozia longidens (Lindb.) Macoun Lophozia longiflora (Nees) Schiffn. (Lophozia guttulata (Lindberg) A. Evans; Lophozia porphyroleuca auct.). – Specimen: BP 47408/H Heves county, Kékes, North Forest Reserve, Mátraháza village, Mátra Mts, on decaying wood in montane beech forest, 24.09.1999, leg. P. Ódor, conf. T. Pócs. Lophozia obtusa (Lindb.) A. Evans (Barbilophozia obtusa (Lindb.) Meyl.). – known from a single locality, specimens: BP 150/H, 158/H, 162/H, 36539/H Comit. Abauj-Torna. In decl. sept. montis Kis Péterménkő supra vall. Vajdavölgy prope pag. Pálháza, 23.06.1953, leg. L. Vajda. Lophozia sudetica (Huebener) Grolle (Lophozia alpestris auct.) Lophozia ventricosa (Dicks.) Dumort. Lophozia wenzelii (Nees) Steph. (Lophozia confertifolia Schiffn.). – synonymy and taxonomy controversial (KOPERSKI et al. 2000, DAMSHOLT 2002). Lunularia cruciata (L.) Dumort. ex Lindb. Mannia fragrans (Balb.) Frye et L. Clark (Grimaldia fragrans (Balb.) Corda; Grimaldia barbifrons Bisch.) Mannia triandra (Scop.) Grolle (Grimaldia rupestris (Nees) Lindenb.; Mannia rupestris (Nees) Frye et L. Clark). – I. Siller in ORBÁN (1978) (Bükk: Ómassa); SILLER (1979); ORBÁN and VAJDA (1983); in BOROS (1968) missing; no specimen could be located in BP and EGR. Marchantia polymorpha L. subsp. polymorpha (Marchantia aquatica (Nees) Burgeff) Marchantia polymorpha L. subsp. ruderalis Bischl. et Boisselier (Marchantia polymorpha auct.) Marsupella emarginata (Ehrh.) Dumort. Marsupella funckii (F. Weber et D. Mohr) Dumort. (Marsupella hungarica Boros et Vajda). – synonymy according to GROLLE (1976), DÜLL (1983). Marsupella sprucei (Limpr.) Bernet. – VAJDA (1960, 1962), DÜLL (1983): ann. 102: “M. sprucei was reported from several places in Mt. Börzsöny in N-Hungary, but Váňa 1978 revised these specimens to M. funckii”; doubtful according to SÖDERSTRÖM et al. (2002). Metzgeria conjugata Lindb. subsp. conjugata Metzgeria conjugata Lindb. subsp. simplex (Müll. Frib.) R. M. Schust. (Metzgeria simplex Lorb. ex Müll. Frib.). – a critical taxon (GROLLE and LONG 2000). Cannot be reliably distinguished from M. conjugata subsp. conjugata by morphological characters (KOPERSKI et al. 2000, AUER et al. 1998), only by chromosome counts. Its occurrence in Hungary is doubtful, since no chromosome counts are known to us. The only specimen in BP: 20710/H Comit. Nógrád. In rupibus umbrosis montis Hangyásbérc prope pag. Kemence, montes Börzsöny, 05.10.1958, leg. L. Vajda. Metzgeria fruticulosa (Dicks.) A. Evans. – BOROS (1968) doubts the record of Szepesfalvi; not in ORBÁN and VAJDA (1983), but recorded in SÖDERSTRÖM et al. (2002); doubtful, because no specimen could be located in BP, EGR and SZO. Metzgeria furcata (L.) Dumort. – var. ulvula Nees is included in the synonymy by most authors. Nardia geoscyphus (De Not.) Lindb. Nardia scalaris Gray (Alicularia scalaris (Gray) Corda) Nowellia curvifolia (Dicks.) Mitt. Oxymitra incrassata (Brot.) Sergio et Sim-Sim (Oxymitra paleacea Lindenb.; Tesselina pyramidata Dumort. p.p.?) Pedinophyllum interruptum (Nees) Kaal. Pellia endiviifolia (Dicks.) Dumort. (Pellia fabbroniana auct.; Pellia calycina Taylor)

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Pellia epiphylla (L.) Corda. – known from a single locality; representative specimen: BP 32526/H Comit. Vas. In paludibus rivi Koponyás ad Jelipuszta prope pag. Kám, 01.06.1924, leg. Á. Boros, several duplicates. Plagiochila asplenioides (L.) Dumort. (Plagiochila major (Nees) S.W. Arnell, nom. illeg.). – P. asplenioides and P. porelloides have not always been separated, e.g.inBOROS (1968) they are treated both under P. asplenioides, but under different varieties. Plagiochila porelloides (Nees) Lindenb. Porella arboris-vitae (With.) Grolle (Madotheca laevigata (Schrad.) Dumort.). – Sometimes spelled “levigata”, e.g.inDAMSHOLT (2002). BOROS (1968) includes var. arboris-vitae (= var. thuja Nees), var. killarniensis (Pearson) M. F. V. Corley, and var. obscura (Nees) M. F. V. Corley; none of these varieties is recognised by PATON (1999). Porella cordaeana (Huebener) Moore (Madotheca cordaeana (Huebener) Dumort.). – BOROS (1968), ORBÁN and VAJDA (1983) mention var. simplicior (J. E. Zetterst.) S. W. Arnell. Porella platyphylla (L.) Pfeiff. (Madotheca platyphylla (L.) Dumort., Porella platyphylloidea (Schwein.) Lindb., Madotheca platyphylloidea (Schwein.) Dumort.). – SÖDERSTRÖM et al. (2002) list P. baueri (Schiffn.) C. E. O. Jensen as species, but according to BOISSELIER-DUBAYLE et al. (1998) it is an allopolyploid (parents: P. platyphylla and P. cordaeana). Other authors treat it as var. of P. platyphylla. P. baueri is recognised in BOROS (1968), ORBÁN and VAJDA (1983); P. platyphylloidea is listed in BOROS (1968), ORBÁN and VAJDA (1983), but according to THERRIEN et al. (1998) it is not specifically distinct from P. platyphylla. Preissia quadrata (Scop.) Nees Ptilidium pulcherrimum (Weber) Vain. Radula complanata (L.) Dumort. Radula lindenbergiana Gottsche ex C. Hartm. – BOROS (1968) estimates the report from Kőszeg Mts by Latzel as reliable, since the plants were collectded with perianthia, whereas reports from other localities are estimated doubtful; these latter reports are omitted in ORBÁN and VAJDA (1983). In BP, only a sterile specimen from Börzsöny Mts could be located (BP 10073/H Comit. Hont. Hung. centr. In cortice Fraxini montis “Hegyes tető” inter pag. Zebegény et Nagymaros, 25.08.1927, leg. Szepesfalvi, teste R. Düll as “probably R. lindenbergiana“). Reboulia hemisphaerica (L.) Raddi Riccardia chamedryfolia (With.) Grolle (Riccardia sinuata (Hook.) Trevis.; Aneura sinuata Dumort.) Riccardia incurvata Lindb. – known from few localities close to Telkibánya in Zemplén Mts, representative specimen: BP 32705/H Comit. Abauj-Torna. In arenoso-argillosis subhumidis ad marg. silv. montis Farkas-hegy prope Telkibánya, 19.10.1959, leg. Á. Boros. Riccardia latifrons (Lindb.) Lindb. (Aneura latifrons Lindb.) Riccardia multifida (L.) Gray (Aneura multifida Dumort.) Riccardia palmata (Hedw.) Carruth. (Aneura palmata Dumort.) Riccia bifurca Hoffm. Riccia canaliculata Hoffm. Riccia cavernosa Hoffm. emend. Raddi (Riccia crystallina auct. non L.). – BOROS (1968), ORBÁN and VAJDA (1983) include var. angustior (Nees) Damsholt. Riccia ciliata Hoffm. (Riccia subtumida Milde). – BOROS (1968), ORBÁN and VAJDA (1983) include var. intumescens Bisch., but this may be the oldest name for what is now called R. crinita Taylor (KOPERSKI et al. 2000, SÖDERSTRÖM et al. 2002), see R. crinita. Riccia ciliifera Link ex Lindenb. (Riccia bischoffii Huebener)

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Riccia crinita Taylor (Riccia intumescens (Bisch.) Underw.; Riccia canescens Steph.; Riccia trichocarpa M. Howe). – nomenclature according to SÖDERSTRÖM et al. (2002); not reported for Hungary in SÖDERSTRÖM et al. (2002) but in MARSTALLER (1993a); specimen: JE, Villanier Geb. Nagy Cser b. Harkany, 29.05.88, leg. R. Marstaller, confirmed P. Erzberger (Erzberger, unpubl.). Riccia crystallina L. emend. Raddi Riccia duplex Lorb. ex Müll.Frib. – according to BOROS (1968), the only specimen was inter- preted controversially by Vajda (R. duplex) and by Jovet-Ast (R. canaliculata); the record of R. duplex is accepted by ORBÁN and VAJDA (1983), DÜLL (1983), RAJCZY (1990) and SÖDERSTRÖM et al. (2002). Specimen: BP 19028/H Comit. Békés. ad margines agrorum (rizsföld) Vizesfás prope Vésztő, 04.09.1954, leg. V. Csapody, det. L. Vajda; rev. S. Jovet-Ast (in litt. 07.07.1959): “21: R. duplex. Je ne crois pas. C‘ est, je crois, R. canaliculata car les cellules des écailles mesurent 40–45 µm × 26–40 µm, les spores 77–83 µm et ont 3–4 alvéoles ayant chacune un mucron”. Riccia fluitans L. emend. Lorb. Riccia frostii Austin Riccia glauca L. Riccia gougetiana Durieu et Mont. Riccia huebeneriana Lindenb. (Riccia pseudo-frostii Schiffn.) Riccia papillosa Moris (Riccia pseudopapillosa Steph.) Riccia rhenana Lorb. ex Müll. Frib. Riccia sorocarpa Bisch. Riccia subbifurca Warnst. ex Croz. (Riccia baumgartneri Schiffn.) Riccia warnstorfii Limpr. ex Warnst. Ricciocarpos natans (L.) Corda Scapania aequiloba (Schwägr.) Dumort. Scapania apiculata Spruce. – known from a single locality; specimens: BP 39243/H, BP 39244/H Comit. Borsod. Ad lignos putresc. in umbrosis vallis Leány-völgy prope Nagyvisnyó, 04.10.1951, 30.09.1950 (resp.), leg. Á. Boros. Scapania aspera M. Bernet et Bernet Scapania calcicola (Arnell et J. Perss.) Ingham Scapania curta (Mart.) Dumort. Scapania irrigua (Nees) Nees Scapania lingulata H. Buch (Scapania microphylla Warnst.) Scapania mucronata H. Buch Scapania nemorea (L.) Grolle (Scapania nemorosa Dumort.) Scapania scandica (Arnell et H. Buch) Macvicar (Scapania parvifolia Warnst.). – according to GROLLE and LONG (2000), S. parvifolia Warnst. is included; it has been treated separately in BOROS (1968), ORBÁN and VAJDA (1983). Scapania umbrosa (Schrad.) Dumort. – known from a single locality; specimens: BP 39980/H, BP 39981/H Comit. Borsod. Ad truncos putresc. in piceeto culto ad Nagymező prope Répáshuta, 05.10.1951, 26.09.1952 (resp.), leg. Á. Boros. Scapania undulata (L.) Dumort. – BOROS (1968), ORBÁN and VAJDA (1983) include var. undulata, var. aequatiformis De Not., var. dentata (Dumort.) Jörg.; but infraspecific taxonomy is controversial (KOPERSKI et al. 2000). Sphaerocarpos texanus Austin (Sphaerocarpos californicus Austin). – BOROS (1968), ORBÁN and VAJDA (1983): a single collection, reported in DÜLL (1983) and accepted in SÖDERSTRÖM et al. (2002); specimens: BP 238/H, BP 239/H, BP 30007/H Comit. Somogy. In agris arenosis ad viam ferream prope Darány, 29.03.1923, leg. Á. Boros, det. V. Schiffner.

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Trichocolea tomentella (Ehrh.) Dumort. Tritomaria exsecta (Schmidel) Loeske (Sphenolobus exsectus (Schmidel) Steph.) Tritomaria exsectiformis (Breidl.) Loeske (Sphenolobus exsectiformis (Breidl.) Steph.) Tritomaria quinquedentata (Huds.) H. Buch (Lophozia quinquedentata (Huds.) Cogn.)

MUSCI

Acaulon muticum (Hedw.) Müll. Hal. Acaulon triquetrum (Spruce) Müll. Hal. Aloina aloides (Schultz) Kindb. – considered doubtful by BOROS (1968) and DÜLL (1984), missing in ORBÁN and VAJDA (1983), but reported by MARSTALLER (1993a); specimens: JE: Villanier Gebirge, Südungarn, Nagyharsányi-hegy, 27.05.88, leg. R. Marstaller, JE: Pecs-Südungarn Kis Tubes in Felsspalten, 08.08.88, leg. R. Marstaller, both confirmed by P. Erzberger (Erzberger, unpubl.); also found by G. Kis in recent collections of T. Pócs and others, specimen: EGR 94205/C pp. Tolna County, N part of Szekszárdi dombság, “Szarvas szurdik”, old branching hollow road 4 km W of Szekszárd town, 46° 20.5’ N, 18° 39’ E, 26.10.1997, leg. S. et T. Pócs and B. O. van Zanten, conf. P. Erzberger. Aloina ambigua (Bruch et Schimp.) Limpr. Aloina brevirostris (Hook. et Grev.) Kindb. Aloina rigida (Hedw.) Limpr. – incl. var. mucronulata (Bruch et Schimp.) Limpr., which was first reported for Hungary by DELGADILLO (1975); specimen: Comit. Komárom, Neszmély, Boros, 1937 (S-PA); the var. was recently synonymised with A. obliquifolia (Müll. Hal.) Broth. by GALLEGO et al. (1999); however, contrary to GALLEGO et al. (1999), we feel that the only relevant diagnostic character – the mucronate leaf apex – does not warrant specific status, since it can vary within an individual plant, as already noted by the authors of Bryologia Europaea (BRUCH et al. 1836–1855, vol. 2, p. 75). Amblyodon dealbatus (Hedw.) Bruch et Schimp. – extinct; according to RAJCZY (1990), last found at a source in Vértes Mts (specimens: BP 89904, BP 119647, BP 119648, BP 119649 Comit. Komárom. In arenosis humidis silvaticis ad fontem Dobai-kút in valle rivi Fekete-ér prope pagum Oroszlány, 05.09.1937, leg. Á. Boros); BOROS (1968) reports a population in an electrically lit cave, see VÖRÖSS (1969). Amblystegium confervoides (Brid.) Schimp. (Amblystegiella confervoides (Brid.) Loeske; Platydictya confervoides (Brid.) H. A. Crum). – Hedenäs (1987) demonstrated that this species and A. subtile are not related to Platydictya jungermannioides and therefore should not be included in that genus. Amblystegium fluviatile (Hedw.) Schimp. (Hygroamblystegium fluviatile (Hedw.) Loeske) Amblystegium humile (P. Beauv.) Crundw. (Leptodictyum kochii (Schimp.) Warnst.; Ambly- stegium kochii Schimp.; Amblystegium trichopodium (Schultz) C. Hartm.; Leptodictyum trichopodium (Schultz) Warnst.) Amblystegium radicale (P. Beauv.) Schimp. (Amblystegium saxatile Schimp.; Campylium radicale (P. Beauv.) Grout; Chrysohypnum radicale (P. Beauv.) Boros; Chrysohypnum hygrophilum (Jur.) Loeske, nom. illeg., nom. superfl.) Amblystegium serpens (Hedw.) Schimp. (Amblystegium juratzkanum Schimp.). – var. serpens and var. juratzkanum (Schimp.) Rau et Herv. Amblystegium subtile (Hedw.) Schimp. (Platydictya subtilis (Hedw.) H. A. Crum; Amblystegiella subtilis (Hedw.) Loeske). – see note on A. confervoides.

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Amblystegium tenax (Hedw.) C. E. O. Jensen (Hygroamblystegium tenax (Hedw.) Jenn.; Hygroamblystegium irriguum Loeske) Amblystegium varium (Hedw.) Lindb. Amphidium mougeotii (Bruch et Schimp.) Schimp. Anacamptodon splachnoides (Brid.) Brid. Andreaea rupestris Hedw. (Andreaea petrophila Fürnr.). – MURRAY (1988) recognises 2 varieties: var. rupestris and var. papillosa (Lindb.) Podp.; according to revisions by B. Papp (unpubl.), both varieties occur in Hungary. Anomodon attenuatus (Hedw.) Huebener Anomodon longifolius (Brid.) Hartm. Anomodon rostratus (Hedw.) Schimp. Anomodon rugelii (Müll. Hal.) Keissl. (Anomodon apiculatus Sull.) Anomodon viticulosus (Hedw.) Hook. et Taylor Antitrichia curtipendula (Hedw.) Brid. Aphanorrhegma patens (Hedw.) Lindb. (Physcomitrella patens (Hedw.) Schimp.) Archidium alternifolium (Hedw.) Schimp. (Phascum alternifolium Dicks. ex Hedw.; Archidium phascoides Brid., nom. illeg., nom. superfl.) Atrichum angustatum (Brid.) Bruch et Schimp. (Polytrichum angustatum Brid.; Catharinea angustata (Brid.) Brid.) Atrichum undulatum (Hedw.) P. Beauv. (Catharinea undulata (Hedw.) F. Weber et D. Mohr). – var. undulatum (= Catharinea undulata var. minor (Hedw.) F. Weber et D. Mohr) and var. gracilisetum Besch. (= Atrichum haussknechtii Jur. et Milde, Catharinea haussknechtii Broth.); in BOROS (1968), ORBÁN and VAJDA (1983) treated as 2 species. Aulacomnium androgynum (Hedw.) Schwägr. Aulacomnium palustre (Hedw.) Schwägr. Barbula convoluta Hedw. (Streblotrichum convolutum (Hedw.) P. Beauv.). – BOROS (1968) qualifies as doubtful reports by Latzel of var. commutata (Jur.) Husn.; infraspecific taxonomy critical; in BLOCKEEL and LONG (1998), var. commutata is no longer recognised, whereas FRAHM and AHMED (2004) recognise it at species level as B. sardoa (Schimp.) J.-P. Frahm. Barbula indica (Hook.) Spreng. – GALAMBOS (1992), DÜLL (1984, 1992), RAJCZY (1990): endangered; specimens conf. J. Kučera. Barbula unguiculata Hedw. Bartramia halleriana Hedw. (Bartramia norvegica Lindb.) Bartramia ithyphylla Brid. Bartramia pomiformis Hedw. – BOROS (1968) includes var. pomiformis and var. crispa (Brid.) Bruch et Schimp., nom illeg., nom. superfl., accepted name: var. elongata Turner. Blindia acuta (Hedw.) Bruch et Schimp. Brachydontium trichodes (F. Weber) Milde. – in BOROS (1968), MÖNKEMEYER (1927) spelled “Brachyodontium”. Brachythecium albicans (Hedw.) Schimp. – var. julaceum Warnst. is placed into synonymy with B. albicans (KOPERSKI et al. 2000). Brachythecium campestre (Müll. Hal.) Schimp. – according to BOROS (1968), ORBÁN and VAJDA (1983) most specimens are doubtful. Brachythecium capillaceum (F. Weber et D. Mohr) Giacom. (Brachythecium salebrosum var. capillaceum (F. Weber et D. Mohr) Lorentz; Brachythecium rotaeanum (De Not.) H. Rob.). – in BOROS (1968), ORBÁN and VAJDA (1983) included in B. salebrosum. Brachythecium geheebii Milde (Homalothecium geheebii (Milde) Wigh)

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Brachythecium glareosum (Spruce) Schimp. Brachythecium mildeanum (Schimp.) Schimp. ex Milde Brachythecium laetum (Brid.) Schimp. (Brachythecium oxycladon auct. non (Brid.) A. Jae- ger). – nomenclature according to ROBINSON and IGNATOV (1997). Brachythecium plumosum (Hedw.) Schimp. Brachythecium populeum (Hedw.) Schimp. – BOROS (1968) includes var. populeum and var. amoenum (Milde) Limpr. Brachythecium reflexum (Starke) Schimp. Brachythecium rivulare Schimp. Brachythecium rutabulum (Hedw.) Schimp. – BOROS (1968) includes var. plumulosum Bruch et Schimp. and var. eurhynchioides Limpr.; no infraspecific taxa are recognised today (KOPERSKI et al. 2000). Brachythecium salebrosum (F. Weber et D. Mohr) Schimp. (Brachythecium lanceolatum Warnst.) Brachythecium velutinum (Hedw.) Schimp. Bryoerythrophyllum recurvirostrum (Hedw.) P. C. Chen (Erythrophyllum rubellum Hilp.; Didymodon rubellus Bruch et Schimp.; Barbula recurvirostra (Hedw.) Dixon) Bryum algovicum Sendtn. ex Müll. Hal. (Bryum pendulum (Hornsch.) Schimp.; Bryum angustirete Kindb. ex Macoun). – incl. f. hazslinszkyanum (Péterfi) Podp., and var. ruppinense (Warnst.) Podp.; the latter is today included in the synonymy of B. algovicum, of which no infraspecific taxa are recognised (KOPERSKI et al. 2000). Bryum alpinum Huds. ex With. – BOROS (1968) includes f. spindleri (Podp. et Stolle) Podp., which is now in the synonymy of B. alpinum (KOPERSKI et al. 2000). Bryum argenteum Hedw. Bryum bicolor Dicks. (Bryum atropurpureum Bruch et Schimp.). – BOROS (1968) includes f. arenarium (Jur.) Podp. and var. bohemicum (Podp.) Podp.; taxonomy of the former is controversial (DEMARET 1993, KOPERSKI et al. 2000); revision of specimens is necessary. Bryum bornholmense Wink. et R. Ruthe. – In ORBÁN and VAJDA (1983) as species, in BOROS (1968) as segregate of B. erythrocarpum Schwägr.; BOROS (1968), ORBÁN and VAJDA (1983) quote the specimen revised by CRUNDWELL and NYHOLM (1964). Other published record in PAPP and RAJCZY (1999). Bryum caespiticium Hedw. – var. caespiticium, var. badium Brid. (= Bryum badium (Brid.) Schimp.) and var. imbricatum Bruch et Schimp. (= B. kunzei Hoppe et Hornsch. = B. caespiticium subsp. kunzei (Hoppe et Hornsch.) Giacom., see BOROS (1968), ORBÁN and VAJDA (1983)); Bryum bakonyense Latzel (incl. var. tettyense Podp.) is now included in the synonymy of B. caespiticium (KOPERSKI et al. 2000). Bryum capillare Hedw. – var. capillare and (according to DÜLL 1985) var. platyloma (Schwägr.) Schimp. Bryum creberrimum Taylor (Bryum affine Lindb. et Arnell) – author of B. affine quoted from DEMARET (1993); according to KOPERSKI et al. (2000), the name B. affine Schultz is a nom. illeg., hom. post. Bryum elegans Nees ex Brid. (Bryum capillare subsp. elegans (Brid.) Podp.). – in BOROS (1968) as subspecies of B. capillare; see also ZANTEN (1999a). Bryum funckii Schwägr. – Specimens BP 169883 and SZO: Comit. Vas. Kőszeg, a meszesvölgy kőfejtőben. Phylliton. leg. and det. A. Latzel. Bryum gemmiferum R. Wilczek et Demaret. – ZANTEN (1999a). Bryum gemmilucens R. Wilczek et Demaret. – ORBÁN and VAJDA (1983) Appendix.

Studia bot. hung. 35, 2004 110 ERZBERGER, P. and PAPP, B.

Bryum imbricatum (Schwägr.) Bruch et Schimp. (Bryum inclinatum (Brid.) Blandow, nom. illeg., hom. post.). – BOROS (1968) includes var. otoeides Podp., var. laubacense (Roth) Podp., var. hageni (Limpr.) Podp. (= B. hagenii Limpr., now included in the synonymy of B. imbricatum), subsp. borosii Podp. ined.; infraspecific taxonomy controversial (KOPERSKI et al. 2000). Bryum intermedium (Brid.) Blandow Bryum klinggraeffii Schimp. – reported by DÜLL (1985). Published records in PAPP and RAJCZY (1999) and PAPP and ERZBERGER (2000). Bryum laevifilum Syed (Bryum flaccidum Brid.). – taxonomy according to HODGETTS (2001): This taxon has been called (probably incorrectly) B. flaccidum Brid. (also as variety of B. capillare) and B. subelegans Kindb., but the latter is a different species not yet recorded from Hungary; see also ZANTEN 1999a. Bryum mildeanum Jur. (Bryum alpinum var. mildeanum (Jur.) Podp.). – in BOROS (1968), ORBÁN and VAJDA (1983) as var. mildeanum (Jur.) Podp. under B. alpinum. Bryum neodamense Itzigs. ex Müll. Hal. Bryum pallens Sw. Bryum pallescens Schleich. ex Schwägr. (Bryum cirrhatum Hoppe et Hornsch.). – In BOROS (1968), ORBÁN and VAJDA (1983) B. pallescens and B. cirrhatum are treated as two separate taxa; BOROS (1968) includes under B. cirrhatum var. praecox Warnst., var. fuscum (Lindb.) Podp. (= B. fuscum Lindb.; this taxon is placed into synonymy with B. intermedium according to KOPERSKI et al. (2000)) and var. affine (Bruch) Podp. (syn. B. creberrimum?); the delimitation of taxa is not clear and herbarium revision desirable. Bryum pseudotriquetrum (Hedw.) P. Gaertn., B. Mey. et Scherb. (Bryum ventricosum Relhan, nom. illeg., nom. superfl.). – var. pseudotriquetrum and var. bimum (Schreb.) Lilj., treated as varieties in BOROS (1968), but as separate species in ORBÁN and VAJDA (1983). Bryum radiculosum Brid. (Bryum murale Wilson ex Hunt) Bryum rubens Mitt. – a widespread species in Hungary, (to our knowledge) first collection in Hungary: SZO Comit. Győr-Sopron, in lapicida ad fontem Deákkut supra opp. Sopron, 29.10.1977, leg. et det. I. Galambos; further references JAKAB (1997), ZANTEN (1999a), PAPP and RAJCZY (1999, 2000). Bryum ruderale Crundw. et Nyholm. – reported in ORBÁN and VAJDA (1983) as species, in BOROS (1968) as segregate under B. erythrocarpum Schwägr.; BOROS (1968) quotes the specimen from Hungary revised by CRUNDWELL and NYHOLM (1964). Other published records in PAPP and RAJCZY (1999). Bryum schleicheri DC.–inBOROS (1968) as subspecies of B. turbinatum; known from a single locality (Specimen: BP 117254 Comit. Borsod. In turfosis rivi Disznóspatak prope Jávorkút. 05.10.1952, leg. Á. Boros), all other specimens from the localities listed in BOROS (1968), ORBÁN and VAJDA (1983) were revised to B. pseudotriquetrum (B. Papp, unpubl.). Bryum stirtonii Schimp. – ORBÁN and VAJDA (1983) Appendix. Bryum torquescens Bruch et Schimp. – in BOROS (1968) as subspecies of B. capillare; ZANTEN (1999a). Bryum turbinatum (Hedw.) Turner. – BOROS (1968) includes subsp. stephani Podp. Bryum uliginosum (Brid.) Bruch et Schimp. (Bryum cernuum Lindb.). – BOROS (1968) includes Bryum danubiale Podp. ined.; no specimen could be located in BP and EGR. Bryum versicolor A. Braun ex Bruch et Schimp. – 1 locality in BOROS (1968), omitted in ORBÁN and VAJDA (1983), reported in DÜLL (1984); no specimen could be located in BP and EGR. Bryum violaceum Crundw. et Nyholm. – ZANTEN (1999a), PAPP and RAJCZY (1999).

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Bryum warneum (Röhl.) Blandow ex Brid. – known from a single locality (Specimen: BP 117001 Comit. Komárom. In arenosis humidis foveae “Nagy-Irtás” prope Bánhida. 08.08.1937, leg. Á. Boros), the only other specimen in BP (from Zamárdi, Lake Balaton, BOROS (1968), ORBÁN and VAJDA (1983)) was revised to B. algovicum (B. Papp, unpubl.). Bryum weigelii Spreng. (Bryum duvalii Voit) Buxbaumia aphylla Hedw. Buxbaumia viridis (Lam. et DC.) Brid. ex Moug. et Nestl. (Buxbaumia indusiata Brid.) Calliergon cordifolium (Hedw.) Kindb. – BOROS (1968) mentions f. phyllorhizans Latzel. Calliergon giganteum (Schimp.) Kindb. Calliergon stramineum (Brid.) Kindb. (Straminergon stramineum (Brid.) Hedenäs) Calliergonella cuspidata (Hedw.) Loeske (Calliergon cuspidatum (Hedw.) Kindb.; Acrocladium cuspidatum (Hedw.) Lindb.). – BOROS (1968) mentions var. pungens (Schimp.) Latzel, a modification of dry habitats. Campylium calcareum Crundw. et Nyholm (Chrysohypnum calcareum (Crundw. et Nyholm) Boros; Chrysohypnum sommerfeltii auct.; Campylophyllum calcareum (Crundw. et Nyholm) Hedenäs) Campylium chrysophyllum (Brid.) Lange (Chrysohypnum chrysophyllum (Brid.) Loeske; Campyliadelphus chrysophyllus (Brid.) Kanda) Campylium elodes (Lindb.) Kindb. (Chrysohypnum helodes (Lindb.) Loeske, orth. var.; Campylium helodes (Lindb.) Broth., orth. var.; Campyliadelphus elodes (Lindb.) Kanda) Campylium polygamum (Schimp.) C. E. O. Jensen (Chrysohypnum polygamum (Schimp.) Loeske; Drepanocladus polygamus (Schimp.) Hedenäs) Campylium stellatum (Hedw.) C. E. O. Jensen (Chrysohypnum stellatum (Hedw.) Loeske). – var. stellatum and var. protensum (Brid.) Bryhn (according to BOROS (1968), DÜLL 1985); both also reported in HEDENÄS (2003). Campylopus pyriformis (Schultz) Brid. (Campylopus torfaceus Bruch et Schimp.). – a single record is reported in BOROS (1968), ORBÁN and VAJDA (1983), but no specimen could be located in BP and EGR. Campylostelium saxicola (F. Weber et D. Mohr) Bruch et Schimp. Ceratodon conicus (Hampe) Lindb. – ZANTEN (1999b) collected a single, sterile specimen that could not be named with absolute certainty; therefore doubts remain. Ceratodon purpureus (Hedw.) Brid. Cinclidotus danubicus Schiffn. et Baumgartner Cinclidotus fontinaloides (Hedw.) P. Beauv. (Cinclidotus minor Lindb.) Cinclidotus riparius (Brid.) Arnell (Cinclidotus nigricans (Brid.) Wijk et Margad.) Cirriphyllum piliferum (Hedw.) Grout Cirriphyllum tommasinii (Boulay) Grout (Cirriphyllum tenuinerve Wijk et Margad., nom. invalid.; Cirriphyllum vaucheri Loeske et M. Fleisch.; Eurhynchium tommasinii (Sendtn.) R. Ruthe) Climacium dendroides (Hedw.) F. Weber et D. Mohr Cnestrum schisti (F. Weber et D. Mohr) I. Hagen (Rhabdoweisia schisti (F. Weber et D. Mohr) Bruch et Schimp.; Cynodontium schisti (F. Weber et D. Mohr) Lindb.). – reported in BOROS (1968), ORBÁN and VAJDA (1983), ORBÁN (1976); but DÜLL (1992): “The record from Hu needs verifica- tion”; it is not clear why Düll doubts these records. Conardia compacta (Müll. Hal.) H. Rob. (Amblystegium compactum (Müll. Hal.) Austin; Rhynchostegiella compacta (Müll. Hal.) Loeske; Amblystegium densum Milde) Cratoneuron filicinum (Hedw.) Spruce (Amblystegium filicinum (Hedw.) De Not.; Hygroamblystegium filicinum (Hedw.) Loeske). – BOROS (1968), ORBÁN and VAJDA (1983) include

Studia bot. hung. 35, 2004 112 ERZBERGER, P. and PAPP, B. var. filicinum, var. atrovirens (Brid.) Ochyra (= var. fallax (Brid.) G. Roth) and f. trichodes (Brid.) Mönk. Crossidium crassinerve (De Not.) Jur. – reported by ORBÁN (1991: found by Galambos), DÜLL (1984), RAJCZY (1990): potentially endangered; recent records in PÓCS (1999). Crossidium laxefilamentosum W. Frey et Kürschner. – reported as new to Hungary (and Eu- rope and North Africa) by PÓCS et al. (in press). Ctenidium molluscum (Hedw.) Mitt. Cynodontium polycarpum (Hedw.) Schimp. – incl. var. polycarpum and var. strumiferum (Hedw.) Schimp.; taxonomy according to KOPERSKI et al. (2000); var. strumiferum according to BOROS (1968) not in Hungary, but it was found in Hungary (Erzberger, unpubl.). Cynodontium tenellum Limpr. – some records doubtful according to BOROS (1968), but reported in ORBÁN and VAJDA (1983) and RAJCZY (1990): endangered; specimens: BP 57648, BP 57975, BP 98268 Comit. Abauj-Torna. In rupibus andesiticis umbrosis Kerekkő montis Nagyhangyás ad Kőkapu prope pag. Pálháza montes Sátor-hegység, 19.07.1957, 17.07.1957, 21.07.1957 (resp.), leg. L. Vajda. Desmatodon cernuus (Huebener) Bruch et Schimp. (Tortula cernua (Huebener) Lindb.) Desmatodon heimii (Hedw.) Mitt. (Pottia heimii (Hedw.) Hampe; Hennediella heimii (Hedw.) R. H. Zander) Dialytrichia mucronata (Brid.) Broth. (Cinclidotus mucronatus (Brid.) Guim.). – Specimen: BP 110580. Com. Sopron. Kéthely. In der Klausen. 25.04.1895. leg. A. Latzel. Dichodontium pellucidum (Hedw.) Schimp. – in Hu only var. pellucidum; var. flavescens is absent, see excluded taxa. Dicranella cerviculata (Hedw.) Schimp. – known from a single locality, specimens: BP 78522, BP 97180, BP 97182 Comit. Heves. In alveo exsicc. alumen. lacus “Timsós-tó” vallis Ilona-völgy pr. Parádfürdő, 18.08.1961, 19.08.1961 (resp.), leg. Á. Boros. Dicranella crispa (Hedw.) Schimp. (Anisothecium vaginale (Dicks.) Loeske; Anisothecium crispum (Hedw.) C. E. O. Jensen). – author citation of Anisothecium crispum must be C. E. O. Jensen, because according to DÜLL (1992), D. crispa (Schreb.) Lindb. stands for D. schreberiana. Dicranella heteromalla (Hedw.) Schimp. Dicranella humilis R. Ruthe (Anisothecium rigidulum (Hedw.) C. E. O. Jensen; Dicranella varia var. tenella Bruch et Schimp.). – in BOROS (1968) as var. tenella under D. rubra (= varia); specimen: BP 97465 Comit. Veszprém. In apertis silv. sub monte Borostyán-hegy prope Márkó, 21.04.1967, leg. Á. Boros. Dicranella rufescens (Dicks.) Schimp. (Anisothecium rufescens (Dicks.) Lindb.) Dicranella schreberiana (Hedw.) Hilf. ex H. A. Crum et L. E. Anderson (Dicranella schreberi Schimp., nom. illeg., nom. superfl.; Anisothecium schreberianum (Hedw.) Dixon). – var. schreberiana and var. robusta (Braithw.) H. A. Crum et L. E. Anderson (corresponds to var. lenta (Wilson) Limpr., included in BOROS (1968)). Dicranella staphylina H. Whitehouse (Anisothecium staphylinum (H. Whitehouse) Sipman, Rubers et Riemann) Dicranella subulata (Hedw.) Schimp. (Dicranella secunda Lindb., nom. illeg., nom. superfl.; Dicranella curvata (Hedw.) Schimp.). – DÜLL (1984) distinguished 2 varieties, var. subulata and var. curvata (Hedw.) Rabenh.; but DÜLL (1992) remarks that the latter has “probably no taxonomic value”. BOROS (1968) includes var. curvata with a single record. Dicranella varia (Hedw.) Schimp. (Dicranella rubra Lindb., nom. illeg., nom. superfl.; Anisothecium varium (Hedw.) Mitt.; Anisothecium rubrum Lindb., nom. illeg., nom. superfl.)

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Dicranodontium denudatum (Brid.) E. Britton (Dicranodontium longirostre (F. Weber et D. Mohr) Bruch et Schimp.) Dicranoweisia cirrata (Hedw.) Lindb. ex Milde Dicranum bonjeanii De Not. (Dicranum palustre Bruch et Schimp., nom. illeg., hom. post.) Dicranum flagellare Hedw. (Orthodicranum flagellare (Hedw.) Loeske) Dicranum fulvum Hook. Dicranum montanum Hedw. (Orthodicranum montanum (Hedw.) Loeske) Dicranum muehlenbeckii Bruch et Schimp. Dicranum polysetum Sw. (Dicranum undulatum Ehrh. ex F. Weber et D. Mohr, nom. illeg., hom. post.; Dicranum rugosum (Funck) Hoffm. ex Brid., nom. illeg., nom. superfl.) Dicranum scoparium Hedw. Dicranum spurium Hedw. Dicranum tauricum Sapjegin (Orthodicranum tauricum (Sapjegin) Z. Smirnova; Dicranum strictum Schleich. ex D. Mohr; Orthodicranum strictum (D. Mohr) Culm.). – in BOROS (1968) doubtful; in ORBÁN and VAJDA (1983) reported, obviously expanding its area, ERZBERGER (1999). Dicranum viride (Sull. et Lesq.) Lindb. – has been confused with D. tauricum (e.g.JAKAB 1997): ERZBERGER (1999). Didymodon acutus (Brid.) K. Saito (Barbula acuta (Brid.) Brid.; Barbula gracilis Schwägr., nom. illeg., hom. post.) Didymodon cordatus Jur. (Barbula cordata (Jur.) Braithw.; Didymodon austriacus Schiffn. et Baumgartner). – BOROS (1968) includes Barbula austriaca (Schiffn. et Baumgartner) Boros et Polgár (= subsp. austriacus (Schiffn. et Baumgartner) Wijk et Margad., ORBÁN and VAJDA (1983)); now included in the synonymy of D. cordatus (KUČERA 2000). Didymodon fallax (Hedw.) R. H. Zander (Barbula fallax Hedw.) Didymodon ferrugineus (Besch.) M. O. Hill (Barbula reflexa (Brid.) Brid.) Didymodon glaucus Ryan (Didymodon rigidulus var. glaucus (Ryan) Wijk et Margad.). – in BOROS (1968) treated as subspecies of Barbula rigidula; no specimen could be located in BP and EGR. Didymodon insulanus (De Not.) M. O. Hill (Barbula cylindrica (Taylor) Schimp.; Barbula vinealis var. cylindrica (Taylor) Boulay; Didymodon vinealis var. flaccidus (Bruch et Schimp.) R. H. Zander). – taxonomic status is controversial, some authors prefer to place this taxon as var. flaccidus in D. vinealis; we follow KUČERA (2000) in retaining D. insulanus as a species of its own. Didymodon luridus Hornsch. ex Spreng. (Barbula lurida (Spreng.) Lindb., nom. illeg., hom. post.; Barbula trifaria auct. non (Hedwig) Mitt.; Didymodon trifarius auct. non (Hedw.) Röhl.) Didymodon rigidulus Hedw. (Barbula rigidula (Hedw.) Mitt.). – BOROS (1968) includes var. rigidulus and var. validus (Limpr.) Düll, the latter is doubtful according to BOROS (1968) and reported in DÜLL (1984), but Düll has not seen any specimen (Düll in litt.). Didymodon sinuosus (Mitt.) Delogne (Barbula sinuosa (Mitt.) Grav.; Oxystegus sinuosus (Mitt.) Hilp.) Didymodon spadiceus (Mitt.) Limpr. (Barbula spadicea (Mitt.) Braithw.) Didymodon tophaceus (Brid.) Lisa (Barbula tophacea (Brid.) Mitt.) Didymodon vinealis (Brid.) R. H. Zander (Barbula vinealis Brid.) Diphyscium foliosum (Hedw.) D. Mohr (Diphyscium sessile Lindb., nom. illeg., nom. superfl.) Distichium capillaceum (Hedw.) Bruch et Schimp. (Distichium montanum I. Hagen; Cynodontium capillaceum Hedw.; Swartzia montana Lindb.) Ditrichum crispatissimum (Müll. Hal.) Paris (Ditrichum giganteum R. S. Williams; Dit- richum gracile (Mitt.) O. Kuntze, synonymy doubtful (see ERZBERGER 2001); Ditrichum flexicaule

Studia bot. hung. 35, 2004 114 ERZBERGER, P. and PAPP, B. var. sterile (De Not.) Limpr.; Ditrichum flexicaule var. longifolium (J. E. Zetterst.) I. Hagen). – For an account of taxonomy and occurrence of this moss in Hungary: ERZBERGER (2001); some authors prefer to treat it as variety of D. flexicaule. Ditrichum cylindricum (Hedw.) Grout (Trichodon cylindricum (Hedw.) Schimp.; Ditrichum tenuifolium Lindb.) Ditrichum flexicaule (Schwägr.) Hampe. – ERZBERGER (2001) describes the variability and distribution of this species in Hungary. Ditrichum heteromallum (Hedw.) E. Britton (Ditrichum homomallum (Hedw.) Hampe) Ditrichum pallidum (Hedw.) Hampe Ditrichum pusillum (Hedw.) Hampe (Ditrichum tortile (Schrad.) Brockm.) Drepanocladus aduncus (Hedw.) Warnst. – The taxonomy of this genus is still controversial. An evaluation of Hungarian herbaria is needed. For taxonomy of D. aduncus and related species, HEDENÄS (2003) is adopted (with reservations). BOROS (1968) includes var. kneiffii (Schimp.) Mönk., var. polycarpus (Voit) G. Roth and forms. Drepanocladus cossonii (Schimp.) Loeske (Drepanocladus intermedius (Lindb.) Warnst.; Scorpidium cossonii (Schimp.) Hedenäs). – HEDENÄS (2003) reports the occurrence of this species in Hungary, which coincides with our own observations; all Hungarian material reported D. revolvens (anon.) Warnst. probably is D. cossonii. Drepanocladus lycopodioides (Brid.) Warnst. (Pseudocalliergon lycopodioides (Brid.) Hedenäs) Drepanocladus sendtneri (H. Müll.) Warnst. Drepanocladus sordidus (Müll. Hal.) Hedenäs. – HEDENÄS (2003) reports the occurrence of this species in Hungary, based on the following specimen: (S) Komitat Pest, Cserhát-Gebirge, am Rande der Tiefebene, am Bach Rákos bei Pécel, 160 m a.s.l., auf torfigen Seggenwiesen des Baches Rákos, 15 May 1960, A. Boros (Crypt. exs. ed. a. Mus. Hist. Nat. Vindob. 4872) Hedenäs, pers. comm.) Encalypta ciliata Hedw. Encalypta streptocarpa Hedw. (Encalypta contorta Hoppe ex Lindb., nom. illeg., nom. superfl.) Encalypta vulgaris Hedw. (Encalypta extinctoria Lindb.) Entosthodon fascicularis (Hedw.) Müll. Hal. (Funaria fascicularis (Hedw.) Lindb.) Entosthodon hungaricus (Boros) Loeske (Funaria hungarica Boros; Steppomitra hungarica (Boros) Vondraček et Hadač) Ephemerum cohaerens (Hedw.) Hampe. – a single record by Förster is reported in BOROS (1968), ORBÁN and VAJDA (1983). Ephemerum minutissimum Lindb. (E. serratum var. minutissimum (Lindb.) Grout). – This taxon has not always been separated from E. serratum; its occurrence in Hungary was recently demonstrated by revision of E. serratum s.l. (B. Papp, unpubl.). Representative specimen: BP 113777, Comit. Abaúj-Torna. In locis subhumidis vallis Vajda-völgy prope Pálháza. 06.10.1953, leg. Á. Boros, rev. B. Papp Ephemerum recurvifolium (Dicks.) Boulay. – NOVOTNY (1988) reports this moss as new for Hungary. Ephemerum serratum (Hedw.) Hampe. – Hungarian floras include var. longifolium (R. Ruthe) Mönk. (BOROS (1968)), probably a nom. superfl. for var. rutheanum (Schimp.) Jur. (ORBÁN and VAJDA (1983)), the taxonomic value of which is not clear, see KOPERSKI et al. (2000). Revision of the material of E. serratum s.l. at BP and EGR by B. Papp resulted in evidence for the presence of E. serratum s. str. and E. minutissimum in Hungary, the latter taxon more frequent (B. Papp, unpubl.).

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Representative specimen: BP 113781, Comit. Heves. In argillosis subhumidis ripae lacus Nagysástó-rét inter Mátrafüred et Mátraháza. 11.10.1953. leg. Á. Boros. Ephemerum sessile (Bruch) Müll. Hal. – This species was found recently in a specimen of E. serratum during revision by B. Papp, new for Hungary (B. Papp, unpubl.). Specimen: BP 169882 Comit. Borsod. In argillosis subhumidis ad marg. silv. vallis sub monte Király-hegy prope Rakaca. 27.10.1957. leg. Boros. Eucladium verticillatum (Brid.) Bruch et Schimp. Eurhynchium angustirete (Broth.) T. J. Kop. (Eurhynchium zetterstedtii Størmer; Eurhyn- chium striatum subsp. zetterstedtii (Størmer) Podp.) Eurhynchium crassinervium (Wilson) Schimp. (Cirriphyllum crassinervium (Wilson) Loeske et M. Fleisch.) Eurhynchium flotowianum (Sendtn.) Kartt. (Cirriphyllum reichenbachianum (Huebener) Wijk et Margad.; Cirriphyllum velutinoides (Schimp.) Loeske et M. Fleisch.; Cirriphyllum filiforme Broth., nom. illeg., nom. superfl.; Eurhynchium velutinoides Schimp.) Eurhynchium hians (Hedw.) Sande Lac. (Eurhynchium swartzii (Turner) Curn.; Oxyrrhyn- chium swartzii (Turner) Warnst.; Oxyrrhynchium hians (Hedw.) Loeske; Eurhynchium praelongum auct. non L.). – This taxon is called E. swartzii in BOROS (1968), but in ORBÁN and VAJDA (1983) two names are treated as if they were separate taxa, although both are synonyms of E. hians as con- ceived here: Oxyrrhynchium swartzii (Turner) Warnst. and Oxyrrhynchium hians (Hedw.) Loeske. The latter obviously relates to the single record of “var. hians” of E. swartzii mentioned in BOROS (1968), therefore this “taxon” was considered “potentially threatened” in RAJCZY (1990). In BOROS (1968), E. schleicheri was placed as var. abbreviatum under E. swartzii. Revision of specimens would be desirable. Eurhynchium praelongum (Hedw.) Schimp. (Oxyrrhynchium praelongum (Hedw.) Warnst. var. stokesii (Turner) Podp.; Eurhynchium stokesii (Turner) Schimp.) Eurhynchium pulchellum (Hedw.) Jenn. – BOROS (1968), ORBÁN and VAJDA (1983) distinguish between var. pulchellum and var. praecox (Hedw.) Dixon. Eurhynchium pumilum (Wilson) Schimp. (Oxyrrhynchium pumilum (Wilson) Loeske; Rhyn- chostegiella pallidirostra (Brid.) Loeske, nom. illeg. (HEDENÄS 1992)) Eurhynchium schleicheri (R. Hedw.) Jur. (Eurhynchium praelongum var. abbreviatum Turner; Oxyrrhynchium schleicheri (R. Hedw.) Roell). – in BOROS (1968) as var. abbreviatum of E. swartzii. Eurhynchium speciosum (Brid.) Jur. (Oxyrrhynchium speciosum (Brid.) Warnst.) Eurhynchium striatulum (Spruce) Schimp. (Isothecium filescens (Brid.) Mönk.; Plasteurhyn- chium striatulum (Spruce) M. Fleisch.). – omitted by mistake in ORBÁN and VAJDA (1983). Eurhynchium striatum (Hedw.) Schimp. – incl. var. magnusii H. Winter = Eurhynchium magnusii (H. Winter) Pilous. Fabronia ciliaris (Brid.) Brid. (Fabronia octoblepharis (Schleich.) Schwägr.) Fabronia pusilla Raddi Fissidens adianthoides Hedw. Fissidens arnoldii R. Ruthe (Fissidens obtusifolius auct. non Wilson) Fissidens bryoides Hedw. Fissidens crassipes Wilson ex Bruch et Schimp. subsp. crassipes (Fissidens crassipes var. rufipes Schimp.; Fissidens crassipes var. mildeanus (Schimp.) Mönk., nom. illeg., nom. superfl.) Fissidens crassipes Wilson ex Bruch et Schimp. subsp. warnstorfii (M. Fleisch.) Brugg.-Nann. (Fissidens crassipes var. submarginatus M. Fleisch. et Warnst.; Fissidens crassipes var. philibertii

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Besch.). – known from a single locality, specimen: BP 10453 Comit. Abauj-Torna. In muris molae prope pag. Jósvafő, 05.04.1952, leg. L. Vajda, det. Potier de la Varde. Fissidens curnovii Mitt. – Specimen: BP 63066. Comit. Abaúj-Torna. Ad margines sylvarum in monte Nagy Farkashegy, prope pag. Telkibánya, mtes Sátorhegység. 19.06.1960. leg. L. Vajda. Fissidens curvatus Hornsch. (Fissidens algarvicus Solms). – It was found only once in Börzsöny Mts, VAJDA (1956), BOROS (1968), ORBÁN and VAJDA (1983).; nomenclature according to BRUGGEMANN-NANNENGA and PURSELL (1996). Specimen: BP 46459. Comit. Nógrád. In argillosis ad margines rivi Kemencepatak prope pag. Diósjenő, montes Börzsöny. 25.05.1955. leg. Vajda. Fissidens dubius P. Beauv. (Fissidens cristatus Wilson ex Mitt.; Fissidens decipiens De Not.) Fissidens exiguus Sull. – The specimen of Börzsöny Mts that Vajda had originally named F. kosaninii Latzel was revised by Pilous to F. exiguus (VAJDA 1975). Fissidens exilis Hedw. Fissidens gracilifolius Brugg.-Nann. et Nyholm (Fissidens minutulus Sull. var. tenuifolius (Boulay) Norkett; Fissidens pusillus var. minutulus (Sull.) Husn., nom. illeg.). – In the older literature, this taxon was partly included in F. pusillus Wilson and F. minutulus Sull. Fissidens gymnandrus Büse. – In BOROS (1968) this taxon is treated as variety of F. bryoides, in ORBÁN and VAJDA (1983) it is missing; doubtful according to DÜLL (1992). Fissidens incurvus Starke ex Röhl. (Fissidens tamarindifolius (Turner) Brid.). – BOROS (1968) includes var. tamarindifolius (Turner) Braithw. as a form of F. bryoides,inORBÁN and VAJDA (1983) as subsp. of F. bryoides. F. incurvus is estimated doubtful by DÜLL (1992), but we do not agree with this opinion. Fissidens pusillus (Wilson) Milde. – In BOROS (1968), this name comprises both F. gracilifolius and F. pusillus;inORBÁN and VAJDA (1983), F. pusillus is called F. minutulus Sull.; however, this is not an approved synonym of F. pusillus (KOPERSKI et al. 2000). Fissidens taxifolius Hedw. – F. taxifolius subsp. taxifolius, excluding subsp. pallidicaulis (Mitt.) Corb. Fissidens viridulus (Sw.) Wahlenb. – In BOROS (1968), ORBÁN and VAJDA (1983) as var. or subsp. of F. bryoides, resp.; also reported by MARSTALLER (1993a). Fontinalis antipyretica Hedw. Fontinalis hypnoides Hartm. – In BOROS (1968), F. antipyretica var. thermalis Boros is mentioned as synonym of F. hypnoides. Funaria hygrometrica Hedw. Funaria muhlenbergii Turner (Funaria dentata Crome; Funaria calcarea Wahlenb.; Funaria hibernica Hook.; F. mediterranea Lindb., nom. illeg.). – For an account of the taxonomy and Hun- garian distribution of this and the following species see CRUNDWELL and NYHOLM (1974), ERZBERGER (2002). Funaria pulchella H. Philib. – This species has for a long time been misunderstood in Hungary (ERZBERGER 2002). Grimmia anodon Bruch et Schimp. Grimmia dissimulata E. Maier. – This species has been discovered recently as new for the Hungarian bryoflora by unpublished herbarium studies of E. Maier, in the following specimens BP 39067 (sub Grimmia montana Br. eur.): Comit. Veszprém. In rupibus basalticis siccis montis Apátihegy prope pag. Tihany, 21.06.1955, leg. L. Vajda, and in herb. Erzberger, Berlin: 6847, 6848 (sub Grimmia cf. pulvinata): Pilisborosjenő, Nagy-Kevély, 14.04.2001, leg. Erzberger and 7077 (sub Grimmia cf. pulvinata): Vértes, Ugró-völgy bei Csákberény, 29.04.2001, leg. Erzberger, all det. E. Maier.

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Grimmia funalis (Schwägr.) Bruch et Schimp. – known from a single locality, representative specimen: BP 112800 Crypt. exs. ed. a Mus. Hist. Nat. Vind. 3958. Com. Heves, an Andesitfelsen des Berges Disznókő bei Parád, 730 m; 25.03.1951, leg. Á. Boros; numerous dupl., some conf. by J. Munoz and E. Maier. Grimmia hartmanii Schimp. (Dryptodon hartmanii (Schimp.) Limpr.) Grimmia laevigata (Brid.) Brid. (Grimmia campestris Burchell ex Hook.; Grimmia leucophaea Grev.) Grimmia longirostris Hook. (Grimmia affinis Hornsch., nom. illeg., hom. post.; Grimmia ovalis auct. non (Hedw.) Lindb.). – In BOROS (1968) and ORBÁN and VAJDA (1983) for G. longirostris the (incorrect) name G. ovalis is used. Nomenclature after MUÑOZ (1998a); in MUÑOZ and PANDO (2000), Hungary is missing from the world distribution, most of the specimens in BP that were referred to G. longirostris belong to G. muhlenbeckii or other species, but a single specimen was confirmed as G. longirostris by E. Maier (unpubl.): BP 46525 (sub Grimmia ovalis Web. et Mohr) Comit. Abauj-Torna. In rupibus andesiticis siccis montis Kakasbérc prope pag. Füzér, montes Sátorhegység, 09.09.1955, leg. L. Vajda. Grimmia montana Bruch et Schimp. – This taxon is meant by “Grimmia donniana Sm.” in ORBÁN and VAJDA (1983); synonymy erroneous. G. donniana is excluded from the bryoflora of Hungary (see Appendix 1). Hungary is missing from the world distribution of G. montana in MUÑOZ and PANDO (2000), but the specimens from Mecsek Mts in BP were confirmed by E. Maier. Grimmia muehlenbeckii Schimp. (Grimmia trichophylla var. muehlenbeckii (Schimp.) Husn.; Grimmia trichophylla var. tenuis (Wahlenb.) Wijk et Margad.). – In BOROS (1968) and ORBÁN and VAJDA (1983) G. muehlenbeckii is treated as a variety of G. trichophylla, viz. var. muehlenbeckii and var. tenuis, respectively. One of the most widespread species of the genus in Hungary according to recent revisions (E. Maier unpubl.). Grimmia orbicularis Bruch ex Wilson. – The alleged synonymy with G. pulvinata var. africana (Hedw.) Hook. f. et Wilson in ORBÁN and VAJDA (1983) is erroneous (e.g. GREVEN 1995). Grimmia ovalis (Hedw.) Lindb. (Grimmia commutata Huebener). – In BOROS (1968) and ORBÁN and VAJDA (1983) the name G. ovalis stands for G. longirostris! Grimmia plagiopodia Hedw. – According to Győrffy (quoted in BOROS (1968)) part of the records are var. arvernica (H. Philib.) Boulay, a taxon not evaluated (type not seen) by MUÑOZ and PANDO (2000). Grimmia pulvinata (Hedw.) Sm. – The significance of var. africana (Hedw.) Hook. f. et Wil- son in ORBÁN and VAJDA (1983) is not clear, since it has been erroneously synonymised with G. orbicularis. Revision of herbarium specimens is necessary. Grimmia teretinervis Limpr. – This species is not in BOROS (1968), but listed in ORBÁN and VAJDA (1983) with two occurrences, one of which has been confirmed by Muñoz and E. Maier (unpubl.): Specimens BP 75600 and BP 111992 Comit. Borsod. In rupibus calc. silvat. adv. montem Kerek-hegy supra vallem Szinva-völgy pr. Lillafüred, 28.03.1959, leg. Á. Boros. Grimmia tergestina Tomm. ex Bruch et Schimp. – var. tergestina occurs in the Bükk Mts according to NOWACK and POELT (1979), who quote a specimen collected by Boros: Magoskő pr. Ómassa; var. tergestinoides (Culm.) Podp. is probably more generally distributed in Hungary. How- ever, the infraspecific taxonomy is controversial (KOPERSKI et al. 2000, GREVEN 1995, MAIER 2002b). Grimmia trichophylla Grev. – For the current taxonomy and an improved description of this species see MAIER (2002a). Gymnostomum aeruginosum Sm. (Gymnostomum rupestre Schwägr.) Gymnostomum calcareum Nees et Hornsch. Gyroweisia tenuis (Hedw.) Schimp.

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Hamatocaulis vernicosus (Mitt.) Hedenäs (Drepanocladus vernicosus (Mitt.) Warnst.) Hedwigia ciliata (Hedw.) Ehrh. ex P. Beauv. (Hedwigia albicans Lindb. p.p., nom. illeg. nom. superfl.). – This species includes 2 varieties, var. ciliata and var. leucophaea Bruch et Schimp., both are widespread in Hungary. Taxonomy according to HEDENÄS (1994), ERZBERGER (1996). Hedwigia stellata Hedenäs (Hedwigia albicans Lindb. pp., nom. illeg. nom. superfl.). – For an account of this species see ERZBERGER (1996), where also data on Hungarian specimens are published. Helodium blandowii (F. Weber et D. Mohr) Warnst. (Thuidium lanatum (Brid.) I. Hagen, nom. illeg. nom. superfl.) Herzogiella seligeri (Brid.) Z. Iwats. (Dolichotheca seligeri (Brid.) Loeske; Dolichotheca silesiaca (F. Weber et D. Mohr) M. Fleisch., nom. illeg., nom. superfl.; Plagiothecium silesiacum (F. Weber et D. Mohr) Schimp., nom. illeg., nom. superfl.; Isopterygium seligeri (Brid.) Dixon; Sharpiella seligeri (Brid.) Z. Iwats.) Heterocladium dimorphum (Brid.) Schimp. (Heterocladium squarrulosum Lindb.; Hetero- cladium squarrosulum Voit, nom. inval., hom. post.) Hilpertia velenovskyi (Schiffn.) R. H. Zander (Tortula velenovskyi Schiffn.) Homalia besseri Lobarz. (Neckera besseri (Lobarz.) Jur.; Neckera sendtneriana Schimp.) Homalia trichomanoides (Hedw.) Schimp. Homalothecium lutescens (Hedw.) H. Rob. (Camptothecium lutescens (Hedw.) Schimp.). – incl. var. lutescens and var. fallax (H. Philib.) Hedenäs et L. Söderström. Homalothecium philippeanum (Spruce) Schimp. (Camptothecium philippeanum (Spruce) Kindb.) Homalothecium sericeum (Hedw.) Schimp. (Camptothecium sericeum (Hedw.) Kindb.) Homomallium incurvatum (Brid.) Loeske (Hypnum incurvatum Schrad. ex Brid.) Hygrohypnum luridum (Hedw.) Jenn. (Hygrohypnum palustre Loeske, nom. illeg., nom. superfl.). – The forms recorded in BOROS (1968) (f. subenerve (Schimp.) Mönk. and f. hamulosa (Schimp.) Mönk.) are probably only modifications (BOROS (1968)). Hylocomium brevirostre (Brid.) Schimp. – known from a single locality, specimens: BP 150314, BP 150315, BP 150316, BP 150320 Comit. Baranya. In silvis vallis “Csatorna-völgy” montis Hármasbükk prope pagum Somogy, 26.07.1932, leg. Á. Boros et A. Visnya, det. A. Latzel. Hylocomium splendens (Hedw.) Schimp. (Hylocomium proliferum (Brid.) Lindb., nom. illeg., nom. superfl.) Hymenostylium recurvirostrum (Hedw.) Dixon (Hymenostylium curvirostre Mitt.). – incl. var. scabrum (Lindb.) Podp. Hypnum cupressiforme Hedw. – Several varieties besides var. cupressiforme are mentioned in BOROS (1968) and ORBÁN and VAJDA (1983), among them var. lacunosum Brid. (syn. var. tectorum Funck ex Brid.), var. subjulaceum Molendo (syn. var. cuspidatum Jur.); DÜLL (1985) reports var. lacunosum, var. resupinatum (Düll in litt.: doubtful) and var. subjulaceum (Düll in litt.: doubtful). Hypnum lindbergii Mitt. (Hypnum arcuatum Lindb., nom. illeg., hom. post.; Calliergonella lindbergii (Mitt.) Hedenäs) Hypnum pallescens (Hedw.) P. Beauv. (Hypnum reptile Michx.). – This taxon is also treated as var. reptile of H. pallescens. Hypnum vaucheri Lesq. Isopterygiopsis pulchella (Hedw.) Z. Iwats. (Isopterygium pulchellum (Hedw.) A. Jaeger et Sauerb.; Plagiothecium pulchellum (Hedw.) Schimp.)

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Isothecium alopecuroides (Dubois) Isov. (Isothecium viviparum Lindb.; Isothecium myurum Brid.) Isothecium myosuroides Brid. (Pseudisothecium myosuroides (Brid.) Grout) Leptobryum pyriforme (Hedw.) Wilson Leptodictyum riparium (Hedw.) Warnst. (Amblystegium riparium (Hedw.) Schimp.). – The forms recorded in BOROS (1968) (f. longifolia,f.leptophylla) are only modifications (BOROS (1968)). Leptodon smithii (Hedw.) F. Weber et D. Mohr Leskea polycarpa Ehrh. ex Hedw. Leucobryum glaucum (Hedw.) Ångstr. Leucobryum juniperoideum (Brid.) Müll. Hal. Leucodon sciuroides (Hedw.) Schwägr. Meesia triquetra (Richt.) Ångstr. – known from a single locality, representative specimen: BP 119635 Comit. Zala. In turfosis versus Batyk prope Türje, 19.09.1953, leg. Á. Boros, numerous duplicates. Mnium hornum Hedw. Mnium lycopodioides Schwägr. (Mnium ambiguum H. Müll.; Mnium orthorrhynchum var. lycopodioides (Schwägr.) Husn.). – Taxonomy according to KOPONEN (1994) and KOPERSKI et al. (2000); the taxonomic position of “var. dioicum“ (var. riparium (Mitt.) Husn., mentioned under M. marginatum in BOROS (1968)) is unresolved as yet. Mnium marginatum (Dicks.) P. Beauv. – BOROS (1968) includes reports by Latzel of “var. riparium (Mitt.) Husn.”, but the taxonomy of this and its relation to M. marginatum var. dioicum (H. Müll.) Crundw., placed into synonymy with M. lycopodioides by KOPONEN (1980), needs further study. Mnium spinulosum Bruch et Schimp. – Reported from a single record (Bakony Mts) in BOROS (1968), ORBÁN and VAJDA (1983). Specimen: BP 91988. Comit. Veszprém. In faginetis sept. montis Miklóspál-hegy ad Bánd prope Szentgál. 06.06.1949. leg. Á. Boros. Mnium stellare Hedw. Mnium thomsonii Schimp. (Mnium orthorrhynchum auct. non Brid., under this name in BOROS (1968)) Myurella julacea (Schwägr.) Schimp. (Myurella julacea var. scabrifolia Lindb. ex Limpr.) Neckera complanata (Hedw.) Huebener Neckera crispa Hedw. Neckera pennata Hedw. Neckera pumila Hedw. – BOROS (1968), ORBÁN and VAJDA (1983) mention var. philippeana (Bruch et Schimp.) Milde and var. pilifera Jur., but these are no longer recognised (NYHOLM 1960). Orthothecium intricatum (Hartm.) Schimp. Orthotrichum affine Schrad. ex Brid. (Orthotrichum octoblepharum Brid.; Orthotrichum fastigiatum Bruch ex Brid.). – BOROS (1968), ORBÁN and VAJDA (1983) include var. fastigiatum Huebener; but according to modern taxonomists, this taxon is no longer recognised (LUDWIG et al. 1996, VITT 1973, LEWINSKY 1993). Orthotrichum anomalum Hedw. (Orthotrichum saxatile Brid., nom. illeg., nom. superfl.). – BOROS (1968), ORBÁN and VAJDA (1983) include var. saxatile (Schimp.) Milde; according to NYHOLM (1960) probably only a modification. Orthotrichum cupulatum Brid. (Orthotrichum sardagnanum Venturi). – In Hungary var. cupulatum and var. riparium Huebener (= var. nudum (Dicks.) Lindb.) occur according to BOROS (1968); it is not clear, what is meant with var. sardagnanum Venturi.

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Orthotrichum diaphanum Schrad. ex Brid. Orthotrichum gymnostomum Bruch ex Brid. – known from a single locality, specimens: BP 25476, BP 156188 Comit. Pest. In truncos populi in populetis ad Móricgáti tanyák, 14.06.1953, leg. L. Vajda. Orthotrichum lyellii Hook. et Taylor Orthotrichum obtusifolium Brid. Orthotrichum pallens Bruch ex Brid. Orthotrichum patens Bruch ex Brid. – in BOROS (1968) under O. stramineum var. patens (Brid.) Venturi. Orthotrichum pumilum Sw. (Orthotrichum fallax Bruch ex Brid.; Orthotrichum schimperi Hammar). – contrary to BOROS (1968), ORBÁN and VAJDA (1983) modern taxonomists no longer distinguish between varieties of this species (LUDWIG et al. 1996, VITT 1973, LEWINSKY 1993). Orthotrichum rogeri Brid. Orthotrichum rupestre Schleich. ex Schwägr. (Orthotrichum sturmii Hoppe et Hornsch.). – BOROS (1968), ORBÁN and VAJDA (1983) include var. sturmii (Hoppe et Hornsch.) Jur., but this taxon is no longer distinguished (LUDWIG et al. 1996, VITT 1973, LEWINSKY 1993). Orthotrichum scanicum Grönvall (Orthotrichum leucomitrium Bruch et Schimp.) Orthotrichum speciosum Nees Orthotrichum sprucei Mont. – Until its discovery by ERZBERGER and PAPP (2000) in Kiskunság, this moss was only known from oceanic Western Europe, but in the meantime it has been reported from Turkey (ERDAĞ and KÜRSCHNER 2000) and even Kasakhstan (GOFFINET 2002). Orthotrichum stellatum Brid. (Orthotrichum braunii Bruch et Schimp.) Orthotrichum stramineum Hornsch. ex Brid. Orthotrichum striatum Hedw. (Orthotrichum leiocarpum Bruch et Schimp., nom. illeg., nom. superfl.). – BOROS (1968) distinguishes subsp. shawii Wilson, a taxon that is considered by MAZIMPAKA et al. (2000) as a good species. Revision of the specimens is necessary. Orthotrichum tenellum Bruch ex Brid. Orthotrichum urnigerum Myrin Oxystegus tenuirostris (Hook. et Taylor) A. J. E. Sm. (Oxystegus cylindricus (Brid.) Hilp.; Tortella cylindrica (Brid.) Loeske; Trichostomum cylindricum (Brid.) Müll. Hal., nom. illeg., hom. post.; Trichostomum tenuirostre (Hook. et Taylor) Lindb.). – incl. var. gemmiparus (Schimp.) R. H. Zander; GALAMBOS (1986). Palustriella commutata (Hedw.) Ochyra (Cratoneuron commutatum (Hedw.) G. Roth). – Infraspecific taxonomy is controversial. BOROS (1968), ORBÁN and VAJDA (1983) mention 3 additional varieties (var. falcatum, var. ptychodioides and var. sulcatum), which Boros considers to be mere habitat modifications. However, var. commutata and var. falcata (Brid.) Ochyra are treated as separate species by HEDENÄS (2003), the latter he calls Palustriella falcata (Brid.) Hedenäs. On the other hand, KOPERSKI et al. (2000) recognise 4 varieties: var. commutata, var. falcata (Brid.) Ochyra, var. fluctuans (Schimp.) Ochyra, (not reported from Hungary), and var. sulcata (Lindb.) Ochyra. The var. ptychodioides (G. Roth) Mönk. (sub Cratoneuron) mentioned in BOROS (1968), ORBÁN and VAJDA (1983) is included in P. commutata var. commutata by KOPERSKI et al. (2000). Whereas var. falcata and var. sulcata are kept separate by KOPERSKI et al. (2000), HEDENÄS (2003) places var. sulcata into synonymy with Palustriella falcata. Paraleucobryum longifolium (Hedw.) Loeske (Dicranum longifolium Ehrh. ex Hedw.). – BOROS (1968) mentions a record by Latzel of var. hamatum Jur. (= f. hamatum (Jur.) Demaret), but not many authors recognise this form. Phascum curvicolle Hedw. (Microbryum curvicolle (Hedw.) R. H. Zander)

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Phascum cuspidatum Hedw. (Phascum acaulon L. ex With.; Phascum halophilum Šmarda; Tortula acaulon (With.) R. H. Zander). – Whereas no infraspecific taxa are mentioned in ORBÁN and VAJDA (1983), BOROS (1968) includes, besides var. cuspidatum, var. piliferum (Hedw.) Hook. et Taylor (as Phascum acaulon L. forma piliferum (Schreb.) Boros) and Phascum halophilum Šmarda (as f. halophilum Šmarda) Boros et Vajda); the latter is not recognised in KOPERSKI et al. (2000), who treat two additional varieties: var. mitraeforme Limpr. and var. papillosum (Lindb.) C. Hartm. (not reported from Hungary). Herbarium revision or fieldwork is necessary to establish the situation of infraspecific taxa in Hungary. Phascum floerkeanum F. Weber et D. Mohr (Microbryum floerkeanum (F. Weber et D. Mohr) Schimp.) Phascum leptophyllum Müll. Hal. (Chenia rhizophylla (Sakurai) R. H. Zander; Chenia leptophylla (Müll. Hal.) R. H. Zander; Tortula rhizophylla (Sakurai) Z. Iwats. et K. Saito). – ZANTEN (2000) reports this species as new for Hungary. Philonotis arnellii Husn. (Philonotis capillaris auct. ). – for synonymy, see KOPERSKI et al. (2000). Philonotis caespitosa Jur. Philonotis calcarea (Bruch et Schimp.) Schimp. Philonotis fontana (Hedw.) Brid. – KOPERSKI et al. (2000) propose to include P. tomentella Molendo as var. pumila Turner in P. fontana. If this treatment is followed, the typical taxon must be called P. fontana var. fontana. We do not know of any records of P. tomentella in Hungary. Philonotis marchica (Hedw.) Brid. Physcomitrium eurystomum Sendtn. (Physcomitrium acuminatum Bruch et Schimp.). – BOROS (1968) includes f. acuminata (Schleich.) Loeske. Physcomitrium pyriforme (Hedw.) Brid. Physcomitrium sphaericum (C. F. Ludw.) Brid. Plagiobryum zierii (Hedw.) Lindb. Plagiomnium affine (Blandow) T. J. Kop. (Mnium affine Blandow; Mnium cuspidatum auct. p.p. ). – BOROS (1968) includes var. integrifolium Lindb. and var. ciliare Lindb., the taxonomic value of which is not clear. Plagiomnium cuspidatum (Hedw.) T. J. Kop. (Mnium cuspidatum Hedw.) Plagiomnium elatum (Bruch et Schimp.) T. J. Kop. (Mnium seligeri auct.; Mnium affine var. elatum Bruch et Schimp.) Plagiomnium ellipticum (Brid.) T. J. Kop. (Mnium rugicum Laurer). – This species is missing from BOROS (1968), but contained in ORBÁN and VAJDA (1983). Plagiomnium medium (Bruch et Schimp.) T. J. Kop. (Mnium medium Bruch et Schimp.). – subsp. medium. Plagiomnium rostratum (anon.) T. J. Kop. (Mnium rostratum anon.; Mnium longirostre Brid.) Plagiomnium undulatum (Hedw.) T. J. Kop. (Mnium undulatum Hedw.) Plagiopus oederianus (Sw.) H. A. Crum et L. E. Anderson (Plagiopus oederi (Brid.) Limpr.; Bartramia oederi Brid.) Plagiothecium cavifolium (Brid.) Z. Iwats. (Plagiothecium roeseanum Schimp.) Plagiothecium denticulatum (Hedw.) Schimp. – BOROS (1968) includes var. majus (Boulay) Limpr. Plagiothecium laetum Schimp. (Plagiothecium curvifolium Limpr.). – We follow KOPERSKI et al. (2000) in placing P. curvifolium Limpr. as var. curvifolium (Limpr.) Mastracci et M. Sauer within P. laetum Schimp.

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Plagiothecium nemorale (Mitt.) A. Jaeger (Plagiothecium neglectum Mönk.; Plagiothecium sylvaticum auct.) Plagiothecium platyphyllum Mönk. Plagiothecium ruthei Limpr. – KOPERSKI et al. (2000) place this taxon at varietal level within P. denticulatum, but we follow MASTRACCI and SAUER (2001) in recognising it as species. Plagiothecium succulentum (Wilson) Lindb. Plagiothecium undulatum (Hedw.) Schimp. – Specimen: BP 71215. Comit. Vas. Ad margines rivulorum prope pag. Jeli. 12.05.1965. leg. L. Vajda. Platydictya jungermannioides (Brid.) H. A. Crum Platygyrium repens (Brid.) Schimp. Platyhypnidium riparioides (Hedw.) Dixon (Eurhynchium rusciforme Milde, nom. illeg., nom. superfl.; Rhynchostegium riparioides (Hedw.) Cardot). – STECH and FRAHM (1999) and HEDENÄS (1990) suggest placement of this taxon in a genus of its own (KOPERSKI et al. 2000). Pleuridium acuminatum Lindb. (Pleuridium subulatum auct. non (Hedw.) Rabenh., BOROS (1968) uses this name for P. acuminatum Lindb.) Pleuridium subulatum (Hedw.) Rabenh. (Pleuridium alternifolium auct. non (Hedw.) Brid., BOROS (1968) uses this name for P. subulatum (Hedw.) Rabenh.) Pleurochaete squarrosa (Brid.) Lindb. Pleurozium schreberi (Brid.) Mitt. (Entodon schreberi (Brid.) Mönk.) Pogonatum aloides (Hedw.) P. Beauv. Pogonatum nanum (Hedw.) P. Beauv. Pogonatum urnigerum (Hedw.) P. Beauv. Pohlia andalusica (Höhn.) Broth. – Revision of herbarium specimens (Erzberger, in press) showed that this is the most frequent among Hungarian bulbiferous Pohlia species. It is new for Hun- gary. Pohlia annotina (Hedw.) Lindb. (Pohlia grandiflora H. Lindb.). – Synonymy according to KOPERSKI et al. (2000). Synonymies given in ORBÁN and VAJDA (1983) are erroneous. Revision of herbarium specimens (Erzberger, in press) showed that part of the specimens labelled P. grandiflora belong to P. andalusica, P. proligera and P. annotina. Pohlia camptotrachela (Renauld et Cardot) Broth. – Synonymy in ORBÁN and VAJDA (1983) is erroneous. Recently found in a single specimen labelled P. bulbifera (Erzberger, in press): BP 116206: Comit. Abauj-Torna. In arenoso-argillosis subhumidis ad marg. silv. sept. montis Farkas-hegy prope Telkibánya, 19.10.1959, leg. Á. Boros. Pohlia cruda (Hedw.) Lindb. Pohlia elongata Hedw. Pohlia lutescens (Limpr.) H. Lindb. (Leptobryum lutescens (Limpr.) Mönk.). – In BOROS (1968) as doubtful, but confirmed in BOROS and VAJDA (1970), reported in ORBÁN and VAJDA (1983). Pohlia melanodon (Brid.) A. J. Shaw (Mniobryum delicatulum (Hedw.) Dixon; Mniobryum carneum (F. Weber et D. Mohr) Limpr., nom. illeg., nom. superfl.; Pohlia carnea (Schimp.) Lindb.) Pohlia nutans (Hedw.) Lindb. Pohlia proligera (Breidl.) Lindb. ex Arnell. – reported as doubtful in DÜLL (1985), erroneously synonymised in ORBÁN and VAJDA (1983), but some specimens were confirmed (Erzberger, in press); representative specimen: BP 42500: Comit. Veszprém. In locis humidis montis Kabhegy prope pag. Padrag, 05.10.1955, leg. L. Vajda sub Mniobryum albicans, rev. G. Nordhorn-Richter, 1981.

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Pohlia wahlenbergii (F. Weber et D. Mohr) A. L. Andrews (Mniobryum wahlenbergii (F. Weber et D. Mohr) Jenn.; Mniobryum albicans (Wahlenb.) Limpr., nom. illeg., nom. superfl.). – BOROS (1968) mentions var. calcareum, but according to KOPERSKI et al. (2000), intraspecific taxa are critical. Polytrichum alpinum Hedw. (Polytrichastrum alpinum (Hedw.) G. L. Sm.) Polytrichum commune Hedw. – In BOROS (1968) and ORBÁN and VAJDA (1983) var. commune and var. perigoniale (Michx.) Hampe are treated as subspecies. Polytrichum formosum Hedw. (Polytrichastrum formosum (Hedw.) G. L. Sm.; Polytrichum attenuatum Menzies ex Brid.) Polytrichum juniperinum Hedw. Polytrichum longisetum Sw. ex Brid. (Polytrichastrum longisetum (Brid.) G. L. Sm.; Polytrichum gracile Dicks.) Polytrichum piliferum Schreb. ex Hedw. Polytrichum strictum Menzies ex Brid. (Polytrichum alpestre Hoppe). – In ORBÁN and VAJDA (1983) treated as subspecies of P. juniperinum. Pottia bryoides (Dicks.) Mitt. (Tortula protobryoides R. H. Zander) Pottia davalliana (Sm.) C. E. O. Jensen (Microbryum davallianum (Sm.) R. H. Zander) Pottia intermedia (Turner) Fürnr. (Tortula modica R. H. Zander) Pottia lanceolata (Hedw.) Müll. Hal. (Tortula lanceola R. H. Zander). – BOROS (1968) includes var. gasilieni (Venturi) Corb., but this taxon is placed in Tortula atrovirens in ORBÁN and VAJDA (1983) and also in ZANDER (1993); taxonomic value and position require further studies. Pottia mutica Venturi (Microbryum starckeanum var. brachyodus (Bruch et Schimp.) R. H. Zander). – BOROS (1968), ORBÁN and VAJDA (1983) probably treat this taxon under the name P. starckeana;BOROS (1968) includes var. mutica, but only for a single record by Latzel. In our opinion, however, P. mutica is widespread in open calcareous habitats. Other infraspecific taxa of P. starckeana are not mentioned, in ORBÁN and VAJDA (1983) even var. mutica is omitted. Pottia starckeana (Hedw.) Müll. Hal. (Microbryum starckeanum R. H. Zander). – The presence of this taxon in Hungary could be shown recently (Erzberger, unpubl.). Specimen: Fejér county. Vértes Mts. Open calcareous grassland around Bucka-hill between Csákvár and Csákberény, 12.04.2004, leg. P. Erzberger 9985 (herb. Erzberger, Berlin). See also note under the preceding species. Pottia truncata (Hedw.) Bruch et Schimp. ( Pottia truncatula (With.) Büse, nom. illeg., nom. superfl.; Tortula truncata (Hedw.) Mitt.) Pseudephemerum nitidum (Hedw.) Reimers (Pseudephemerum axillare (Lindb.) I. Hagen, nom. illeg., nom. superfl.) Pseudocrossidium hornschuchianum (Schultz) R. H. Zander (Barbula hornschuchiana Schultz). – BOROS (1968) includes var. pseudorevoluta Reimers = var. obtusula (Lindb.) Podp. Ac- cording to NYHOLM (1990), this variety is better placed under P. revolutum, as var. obtusula (Lindb.) Mönk. Pseudocrossidium revolutum (Brid.) R. H. Zander (Barbula revoluta Brid.). – see note under the preceding species. Pseudoleskea saviana (De Not.) Latzel (Pseudoleskea illyrica Glowacki; Lescuraea saviana (De Not.) E. Lawton). – known from a single locality, representative specimen: BP 83964 Crypt. exs. ed. a Mus. Hist. Nat. Vind. 3993. Com. Heves, an bewaldeten Andesitfelsen des Berges Kékes bei Parád, 950–1000 m; 27.04.1952, leg. Á. Boros; numerous dupl. Pseudoleskeella catenulata (Schrad.) Kindb. (Leskea catenulata (Schrad.) Mitt.)

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Pseudoleskeella nervosa (Brid.) Nyholm (Leskea nervosa (Brid.) Myrin; Leskeella nervosa (Brid.) Loeske) Pseudotaxiphyllum elegans (Brid.) Z. Iwats. (Isopterygium elegans (Brid.) Lindb.; Plagio- thecium elegans (Brid.) Sull.) Pterigynandrum filiforme Hedw. Pterogonium gracile (Hedw.) Sm. (Pterogonium ornithopodioides (F. Weber et D. Mohr) Lindb., nom. illeg., nom. superfl.). – known from a single locality, representative specimen: BP 45022, BP 122886 Basaltberg St. György bei Tapolca am Platten-See, an Felsen, 24.03.1901, leg. et det. J. Baumgartner, later (01.05.1955) also collected by Boros and Vajda (BP 39247, BP 122887). Pterygoneurum compactum M.J. Cano, J. Guerra et Ros. – PÓCS (1999). Pterygoneurum crossidioides W. Frey, Herrnst. et Kürschner. – PÓCS (1999); according to PÓCS et al. (2002), the records need confirmation since European specimens are not typical. Pterygoneurum lamellatum (Lindb.) Jur. Pterygoneurum ovatum (Hedw.) Dixon (Pterygoneurum pusillum (Lindb.) C. E. O. Jensen; Pterygoneurum cavifolium Jur., nom. illeg., nom. superfl.) Pterygoneurum squamosum Segarra et Kürschner. – PÓCS (1999). Pterygoneurum subsessile (Brid.) Jur. Ptilium crista-castrensis (Hedw.) De Not. Pylaisia polyantha (Hedw.) Schimp. – spelled “Pylaiea” in BOROS (1968). Pyramidula tetragona (Brid.) Brid. Racomitrium affine (F. Weber et D. Mohr) Lindb. – spelled “Rhacomitrium“inBOROS (1968), now considered an invalid orthographic variant (KOPERSKI et al. 2000); in BOROS (1968) and ORBÁN and VAJDA (1983) as “subsp. affine (Web. et Mohr) Amann”; specimens should be checked because of change in definition of taxa (KOPERSKI et al. 2000). Racomitrium aquaticum (Schrad.) Brid. (Racomitrium protensum (A. Braun) Huebener) Racomitrium canescens (Hedw.) Brid. – Since the specimens have not been revised, it is not clear, which taxa of the R. canescens group (FRISVOLL 1983) are present in Hungary, apart from R. canescens s. str., confirmed by the first author (Erzberger, unpubl.). Racomitrium heterostichum (Hedw.) Brid. – BOROS (1968) includes var. affine (F. Weber et D. Mohr) Lesqu. [nom. illeg., nom. superfl.] and f. obtusa (see following species), but specimens should be checked. Racomitrium obtusum (Brid.) Brid. – In BOROS (1968), ORBÁN and VAJDA (1983) included as f. obtusa under R. heterostichum; herbarium specimens should be checked. Rhabdoweisia fugax (Hedw.) Bruch et Schimp. (Rhabdoweisia striata (Schrad.) Lindb.) Rhizomnium punctatum (Hedw.) T. J. Kop. (Mnium punctatum Hedw.) Rhodobryum ontariense (Kindb.) Kindb. (Rhodobryum spathulatum auct.). – not in BOROS (1968); for an account of this species in Hungary see ORBÁN and PÓCS (1976). Rhodobryum roseum (Hedw.) Limpr. Rhynchostegiella curviseta (Brid.) Limpr. Rhynchostegiella tenella (Dicks.) Limpr. (Rhynchostegiella algiriana (P. Beauv.) Warnst.) Rhynchostegiella teneriffae (Mont.) Dirkse et Bouman (Rhynchostegiella jacquinii (Garov.) Limpr.) Rhynchostegiella tenuicaulis (Spruce) Kartt. (Cirriphyllum germanicum (Grebe) Loeske et M. Fleisch.) Rhynchostegium confertum (Dicks.) Schimp. Rhynchostegium megapolitanum (F. Weber et D. Mohr) Schimp.

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Rhynchostegium murale (Hedw.) Schimp. – BOROS (1968) mentions var. julaceum Schimp., but no infraspecific taxa are recognised today (KOPERSKI et al. 2000). Rhynchostegium rotundifolium (Brid.) Schimp. Rhytidiadelphus squarrosus (Hedw.) Warnst. (Hylocomium squarrosum (Hedw.) Schimp.) Rhytidiadelphus triquetrus (Hedw.) Warnst. (Hylocomium triquetrum (Hedw.) Schimp.) Rhytidium rugosum (Hedw.) Kindb. Saelania glaucescens (Hedw.) Broth. (Saelania caesia (P. Beauv.) Lindb.) Sanionia uncinata (Hedw.) Loeske (Drepanocladus uncinatus (Hedw.) Warnst.) Schistidium apocarpum (Hedw.) Bruch et Schimp. (Grimmia apocarpa Hedw.). – Taxonomy according to BLOM (1996, 1998); the material of the genus should be revised. The name S. apocarpum in BOROS (1968), ORBÁN and VAJDA (1983) is understood sensu lato. Schistidium brunnescens Limpr. subsp. brunnescens (Schistidium apocarpum subsp. brunnescens (Limpr.) Loeske). – BLOM (1996): Budakalász, “Mt. Monalovác”; widespread in calcareous habitats (Erzberger, unpubl.), see also BOROS (1968), ORBÁN and VAJDA (1983). Schistidium crassipilum H. H. Blom. – BLOM (1996): CBiatorbágy by Budapest, Sziklaperem, VII 1979 Marstaller (JE), Bükk Mts., Leányvölgy near Szilvásvárad Vitt 25526 (ALTA)”. Many records of S. apocarpum s.l. are probably S. crassipilum. Schistidium flaccidum (De Not.) Ochyra (Grimmia flaccida (De Not.) Lindb.; Schistidium pulvinatum var. flaccidum (De Not.) De Not.; Schistidium pulvinatum auct. non (Hedw.) Brid.). – BLOM (1996): Representative specimen: BP 5587, Flora Hungarica exsiccata VIII. 729, Pomáz, comit. Pest Hungariae centralis, in rupium fissuris montis “Kis Csikóvár”. 04.1924. leg. and det. Á. Degen. Several other specimens exist from the same locality. We could still find the species in 2000 (Erzberger and Papp, unpubl.). Schistidium singarense (Schiffn.) Lazarenko. – BLOM (1996): Villányi Mts., Tenkeshegy by Máriagyűd, 4.6.1986, Marstaller (JE). Scleropodium purum (Hedw.) Limpr. (Pseudoscleropodium purum (Hedw.) M. Fleisch. ex Broth.) Scorpidium scorpioides (Hedw.) Limpr. Seligeria calcarea (Hedw.) Bruch et Schimp. – missing in BOROS (1968), only ORBÁN and VAJDA (1983). Seligeria campylopoda Kindb. – GOS and OCHYRA (1994), specimens: BP 96768 leg. Á. Boros, rev. L. Gos 1991 and BP 68281 leg. L. Vajda. rev. L. Gos 1991. Comit. Veszprém. In rupibus dolomiticis umbrosis montis Csókakő prope Keszthely. 06.05.1963. Seligeria donniana (Sm.) Müll. Hal. – missing in BOROS (1968), only ORBÁN and VAJDA (1983). Seligeria pusilla (Hedw.) Bruch et Schimp. Seligeria recurvata (Hedw.) Bruch et Schimp. (Seligeria setacea Lindb., nom. illeg., nom. superfl.) Seligeria trifaria (Brid.) Lindb. var. longifolia (Broth.) Ochyra et Gos. – GOS and OCHYRA (1994), specimen: BP 75532 Comit. Heves. In muris humidis in pag. Eger. 31.08.1962. leg. L. Vajda, rev. L. Gos 1991. Sphagnum angustifolium (Warnst.) C. E. O. Jensen (Sphagnum parvifolium (Warnst.) Warnst.). – This species is treated under S. recurvum in BOROS (1968) as f. parvifolium,itis implicitely included in S. recurvum in ORBÁN and VAJDA (1983). Sphagnum capillifolium (Ehrh.) Hedw. (Sphagnum nemoreum auct.; Sphagnum acutifolium Ehrh. ex Schrad.) Sphagnum centrale C. E. O. Jensen (Sphagnum subbicolor auct. non Hampe)

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Sphagnum compactum Lam. et DC. – Most authors do not recognise infraspecific taxa (BOROS (1968) mentions var. squarrosum Russow, f. laxum Cardot and var. subsquarrosum Warnst.). Sphagnum contortum Schultz Sphagnum cuspidatum Ehrh. ex Hoffm. – SZURDOKI (2003). Sphagnum denticulatum Brid. – In spite of occasional difficulties in separating S. inundatum and S. denticulatum, we follow DIERSSEN (1996) in recognising two species. By earlier authors, these taxa were not always strictly separated from one another and from S. subsecundum. This is also evident from the descriptions in ORBÁN and VAJDA (1983). Therefore, old reports are sometimes in- terpreted with difficultiy. For recent records consult SZURDOKI (2003). Sphagnum fallax (H. Klinggräff) H. Klinggräff (Sphagnum recurvum var. mucronatum (Russow) Warnst.). – in ORBÁN and VAJDA (1983) reported as S. recurvum P. Beauv., a wider taxon; in BOROS (1968) as S. fallax and S. recurvum var. mucronatum (Russow) Warnst. Sphagnum fimbriatum Wilson Sphagnum flexuosum Dozy et Molk. (Sphagnum recurvum var. amblyphyllum (Russow) Warnst.) Sphagnum girgensohnii Russow. – only in ORBÁN and VAJDA (1983), missing in BOROS (1968). Sphagnum inundatum Russow. – In spite of occasional difficulties in separating S. inundatum and S. denticulatum, we follow DIERSSEN (1996) in recognising two species. For recent records consult SZURDOKI (2003). Sphagnum magellanicum Brid. Sphagnum obtusum Warnst. Sphagnum palustre L. Sphagnum platyphyllum (Braithw.) Sull. ex Warnst. Sphagnum quinquefarium (Braithw.) Warnst. Sphagnum riparium Ångstr. – reported as new for Hungary by SZURDOKI et al. (2000). Sphagnum russowii Warnst. (Sphagnum robustum (Warnst.) Cardot) Sphagnum squarrosum Crome Sphagnum subnitens Russow et Warnst. (Sphagnum plumulosum Roell) Sphagnum subsecundum Nees. – possibly not always separated from S. denticulatum or S. inundatum, see note under the former species. Sphagnum teres (Schimp.) Ångstr. Sphagnum warnstorfii Russow. – Extinct according to BOROS (1968) (specimen: BP 5518 Comit. Zala. In paludosis turfosis ad piscinas prope Lesenceistvánd, 25.08.1923, leg. Boros, conf. P. Erzberger); a second locality (specimen: BP 5526 Comit. Vas. In sphagnetis vallis Gössbach prope pagum Hámortó, 18.08.1924, leg. Á. Boros, det. P. Fürst, conf. P. Erzberger; 2 specimens in herb. Piers, SZO: Hungaria. Districtus-transdanubialis. In uliginosis vallis Goos solo phyllitico, 30.04.1897, leg. Piers, rev. A. Latzel) is situated in Austria near the Hungarian border (cf. KIRÁLY 1996). Splachnobryum obtusum (Brid.) Müll. Hal. (Splachnobryum wrightii Müll. Hal.). – Synon- ymy according to VAJDA (1960) and DÜLL (1985); known from a single locality, representative specimen: BP 164959 Crypt. exs. ed. a Mus. Hist. Nat. Vind. 4496. Com. Heves, an berieselten, von Kalktuff durchsetzten Stellen der Thermalquelle in der Stadt Eger, ca. 180 m, 15.09.1959, leg. Á. Boros; numerous duplicates. Taxiphyllum densifolium (Broth.) Reimers (Plagiothecium densifolium (Broth.) Limpr.) Taxiphyllum wissgrillii (Garov.) Wijk et Margad. (Taxiphyllum depressum (Brid.) Reimers)

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Tetraphis pellucida Hedw. (Georgia pellucida (Hedw.) Rabenh.) Thamnobryum alopecurum (Hedw.) Gangulee (Arbuscula alopecura (Hedw.) H. A. Crum, Steere et L. E. Anderson, nom. illeg., nom. superfl.; Thamnium alopecurum (Hedw.) Schimp.) Thuidium abietinum (Hedw.) Schimp. (Abietinella abietina (Hedw.) M. Fleisch.). – It is not clear, to which of two varieties (var. abietinum and var. hystricosum (Mitt.) Loeske) the Hungarian records belong; DÜLL (1985) reports var. hystricosum as doubtful; he has not seen specimens (Düll in litt.). Thuidium delicatulum (Hedw.) Schimp. (Thuidium erectum Duby) Thuidium philibertii Limpr. – BOROS (1968) includes var. pseudotamarisci Limpr., but this is generally placed into synonymy with T. philibertii. Thuidium recognitum (Hedw.) Lindb. Thuidium tamariscinum (Hedw.) Schimp. (Thuidium tamariscifolium Lindb., nom. illeg., nom. superfl.) Timmia austriaca Hedw. – known from a single locality, specimens: BP 8544, BP 8545 Comit. Borsod. In rupibus calcareis montis Hollókő prope pag. Nagyvisnyó, 04.10.1951, leg. L. Vajda, and BP 121385, 07.08.1953, leg. Á. Boros. Timmia bavarica Hessl. Tomentypnum nitens (Hedw.) Loeske (Homalothecium trichoides (Lindb.) Boros; Camptothecium nitens (Hedw.) Schimp.; Homalothecium nitens (Hedw.) H. Rob.; Camptothecium trichoides Broth.) Tortella bambergeri (Schimp.) Broth. – not reported by BOROS (1968), ORBÁN and VAJDA (1983), but in MARSTALLER (1993a); three specimens from JE could be confirmed (Erzberger, unpubl.); another specimen was found during revision: Herb. Erzberger, Berlin, 4702: Ungarn, Bükk-Geb., Hór-völgy, Ódor-vár, 06.08.1998, leg. P. Erzberger sub T. tortuosa (untypisch) (Erzberger, unpubl.). Tortella inclinata (R. Hedw.) Limpr. Tortella tortuosa (Hedw.) Limpr. Tortula atrovirens (Sm.) Lindb. (Desmatodon convolutus (Brid.) Grout). – ORBÁN and VAJDA (1983) includes var. gasilieni (Venturi) Limpr., which is treated under Pottia lanceolata in BOROS (1968), but placed in T. atrovirens e.g.inZANDER (1993); taxonomic value and position require further studies. Tortula brevissima Schiffn. – For an account of this species in Hungary, its interesting bryogeography and bryosociology see ERZBERGER (1998), KÜRSCHNER and PAROLLY (1998), KÜRSCHNER and PÓCS (2002) and KÜRSCHNER (2002). Tortula calcicolens W. A. Kramer (Tortula ruralis var. calcicola (J. J. Amann) Barkman; Syntrichia calcicola J. J. Amann) Tortula caninervis (Mitt.) Broth. subsp. spuria (J. J. Amann) W. A. Kramer (Tortula caninervis subsp. spuria var. gypsophila (Roth) W. A. Kramer; Tortula desertorum Broth.; Syntrichia caninervis Mitt. var. spuria (J. J. Amann) R. H. Zander ). – var. spuria; discovered as new for the Hungarian bryoflora during herbarium studies by TÓTH (1987); for nomenclature see WERNER and HÉBRARD (1986); vulnerable according to RAJCZY (1990) (as Tortula desertorum Broth.). Tortula crinita (De Not.) De Not. (Tortula intermedia (Brid.) Berk., nom. illeg., hom. post.; Syntrichia montana Nees; Syntrichia intermedia Brid.). – Two varieties occur in Hungary (TÓTH 1986): var. crinita and var. calva (Durieu et Sagot) Nebel et Heinrichs (= T. intermedia var. calva (Durieu et Sagot) Wijk et Margad).

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Tortula inermis (Brid.) Mont. (Syntrichia inermis (Brid.) Bruch). – Specimen: BP 4452. Comit. Pest. In valle “Apátkúti völgy” ad saxa trachyt. pr. pag. Visegrád. 07.06.1932. leg. J. Szepesfalvi, teste J. Kučera 1998. Tortula laevipila (Brid.) Schwägr. (Syntrichia laevipila Brid.) Tortula latifolia Bruch ex Hartm. (Syntrichia latifolia (Hartm.) Huebener) Tortula mucronifolia Schwägr. (Syntrichia mucronifolia (Schwägr.) Brid.) Tortula muralis L. ex Hedw. – var. muralis and var. aestiva Brid. ex Hedw. Tortula norvegica (F. Weber) Wahlenb. ex Lindb. (Syntrichia norvegica F. Weber). – first published for Hungary by TÓTH (1987), discovered by L. Meinunger. Tortula obtusifolia (Schwägr.) Mathieu (Tortula muralis var. obtusifolia (Schwägr.) Boros). – BOROS (1968) treats this taxon as a variety of T. muralis. Tortula papillosa Wilson (Syntrichia papillosa (Wilson) Jur.) Tortula papillosissima (Copp.) Broth. (Syntrichia ruralis var. hirsuta (Venturi) Podp., [syn. of var. papillosissima]; Syntrichia ruralis var. submamillosa (W. A. Kramer) R. H. Zander (syn. of var. submamillosa)). – Two varieties occur in Hungary (TÓTH 1986): var. papillosissima (= T. ruralis subsp. hirsuta (Venturi) W. A. Kramer var. hirsuta) and var. submamillosa (W. A. Kramer) Hein- richs et Caspari (= T. ruralis subsp. hirsuta (Venturi) W. A. Kramer var. submamillosa W. A. Kramer). Tortula ruraliformis (Besch.) Ingham (Tortula ruralis var. arenicola Braithw., nom. illeg., nom. superfl.; Syntrichia ruralis var. ruraliformis (Besch.) Durand, nom. illeg.; Syntrichia ruralis var. arenicola (Braithw.) J. J. Amann) Tortula ruralis (Hedw.) P. Gaertn., B. Mey. et Scherb. (Syntrichia ruralis (Hedw.) F. Weber et D. Mohr) Tortula subulata Hedw. (Syntrichia subulata (Hedw.) F. Weber et D. Mohr). – var. subulata and var. angustata (Schimp.) Limpr. Tortula virescens (De Not.) De Not. (Syntrichia virescens (De Not.) Ochyra) Trichostomum brachydontium Bruch (Trichostomum mutabile Bruch). – It is not clear, which of the two varieties (var. brachydontium and var. cuspidatum (Braithw.) L. I. Savicz) is present; the report of var. cuspidatum in DÜLL (1984) is based on PODPERA (1954), but there no details are given. Trichostomum crispulum Bruch. – BOROS (1968) mentions T. viridulum Bruch (as doubtful), a taxon that is now placed as var. angustifolium Bruch et Schimp. in T. crispulum (KOPERSKI et al. 2000). Ulota bruchii Hornsch. ex Brid. (Ulota crispa var. norvegica (Grönvall) A. J. E. Sm. et M. O. Hill) Ulota coarctata (P. Beauv.) Hammar (Ulota ludwigii (Brid.) Brid.) Ulota crispa (Hedw.) Brid. (Ulota ulophylla Broth.; Ulota crispula Brid.). – Although treated in BOROS (1968), ORBÁN and VAJDA (1983) as a separate taxon, U. crispula is now generally included in U. crispa (SMITH and PROCTOR 1993, ERZBERGER 2003). Ulota hutchinsiae (Sm.) Hammar (Ulota americana (P. Beauv.) Limpr., nom. illeg., hom. post.). – extinct according to RAJCZY (1990); no specimen could be located in BP and EGR. Warnstorfia exannulata (Schimp.) Loeske (Drepanocladus exannulatus (Schimp.) Warnst.) Weissia brachycarpa (Nees et Hornsch.) Jur. (Hymenostomum microstomum (Hedw.) R. Br. ex Nees et Hornsch.; Weissia microstoma (Hedw.) Müll. Hal., nom. illeg., hom. post.) Weissia condensa (Voit) Lindb. (Hymenostomum tortile (Schwägr.) Bruch et Schimp.; Weissia tortilis (Schwägr.) Müll. Hal., nom. illeg., hom. post.) Weissia controversa Hedw. (Weissia viridula Brid., nom. illeg., nom. superfl.). – var. controversa; synonym: W. viridula var. stenocarpa (Nees et Hornsch.) Bruch et Schimp.; this name is used

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in BOROS (1968) and ORBÁN and VAJDA (1983) in addition to the “type”, the meaning of which is not clear. Weissia fallax Sehlm. (Weissia tortilis var. fallax (Sehlm.) Mönk.; Weissia controversa var. crispata (Nees et Hornsch.) Nyholm) Weissia longifolia Mitt. (Astomum crispum (Hedw.) Hampe) Weissia rostellata (Brid.) Lindb. (Hymenostomum rostellatum (Brid.) Schimp.). – in BOROS (1968) missing, in ORBÁN and VAJDA (1983) reported from a single locality (for which a single, sterile and therefore doubtful specimen could be located: BP 75551 Comit. Abauj-Torna. In rupibus humidis vallis Senyővölgy pr. pag. Telkibánya; montes Sátorhegység); however, a recent collection (specimen: BP 167684 Budapest County. Budai-mountains, mount Hárs-hegy at Budapest on acidic soil on the northeastern part of the hill, 30.06.2000, leg. et det. B. Papp) proves the presence of this species in Hungary. Weissia rutilans (Hedw.) Lindb. Zygodon rupestris Schimp. ex Lorentz (Zygodon viridissimus var. vulgaris Malta, nom. illeg., nom. superfl.). – BOROS (1968), ORBÁN and VAJDA (1983) report only Z. viridissimus, but according to revision, specimens in BP and JE are Z. rupestris (B. Papp, P. Erzberger, unpubl.).

APPENDIX 1. EXCLUDED SPECIES (Taxa reported in ORBÁN and VAJDA (1983) are marked with an asterisk)

HEPATICAE

Cephaloziella elachista (Gottsche et Rabenh.) Schiffn. – A doubtful record from Mecsek Mts is rejected by BOROS (1968); not in Hungary according to SÖDERSTRÖM et al. (2002). Gymnomitrion corallioides Nees. – doubtful according to SÖDERSTRÖM et al. (2002); source: MÜLLER (1954–1957): “Ungarn, Liptauer Gebirge” – present-day Slovakia, no records from Hun- gary known. *Nardia insecta Lindb. – although reported in BOROS (1968), ORBÁN and VAJDA (1983) and SÖDERSTÖM et al. (2002), all specimens in BP were revised to N. geoscyphus by Váňa; since all former reports refer to a single locality (representative specimen: BP 39074/H Comit. Abauj-Torna. In arenoso-argillosis silvat. montis Farkas-hegy supra Telkibánya, 19.10.1959, leg. Á. Boros), there is no evidence for the occurrence of this species in Hungary. *Porella baueri (Schiffn.) C. E. O. Jensen. – see annotation to Porella platyphylla. *Porella platyphylloidea (Schwein.) Lindb. (Madotheca platyphylloidea (Schwein.) Dumort.). – according to THERRIEN et al. (1998) not specifically distinct from P. platyphylla. Ptilidium ciliare (L.) Hampe. – not in Hungary, according to BOROS (1968), SÖDERSTRÖM et al. 2002. Sphaerocarpos michelii Bellardi (Sphaerocarpos terrestris Sm.). – Erroneously reported in DÜLL (1983), RAJCZY (1990) and accepted in SÖDERSTRÖM et al. (2002); Düll in litt.: “pers. com- munication by T. Pócs: “Only once collected, specimen in BP””. According to T. Pócs (pers. comm.) this statement of Düll is erroneous and refers to S. texanus.

Studia bot. hung. 35, 2004 130 ERZBERGER, P. and PAPP, B.

MUSCI

Aloina bifrons (De Not.) Delgad. – Missing in BOROS (1968), ORBÁN and VAJDA (1983), reports in DÜLL (1984, 1992) are based on an erroneous information from Orbán. Aloina obliquifolia (Müll. Hal.) Broth. – see note under A. rigida. *Anoectangium aestivum (Hedw.) Mitt. – The report in ORBÁN and VAJDA (1983) (Appen- dix) is based on a single specimen, which was revised to Hymenostylium recurvirostrum (B. Papp, unpubl.). Specimen: BP 160192. Comit. Borsod. Lillafüred. Ad margines sylvarum. 10.03.1978. leg. L. Vajda. Brachythecium starkei (Brid.) Schimp. – excluded in BOROS (1968), omitted in ORBÁN and VAJDA (1983). *Bryum veronense De Not. – in BOROS (1968) as subspecies of B. argenteum; see also ORBÁN and VAJDA (1983), VAJDA (1966b), VAJDA and ORBÁN (1975); the specimens (BP 118573 and BP 57956: Comit. Nógrád. In rupibus irrigatis rivi Rózsapatak prope Királyháza, montes Börzsöny. 03.02.1957, leg. L. Vajda) were revised to B. argenteum (B. Papp, unpubl.). Calliergon sarmentosum (Wahlenb.) Kindb. (Warnstorfia sarmentosa (Wahlenb.) Hedenäs). – erroneously reported in HEDENÄS (2003) for Hungary; the specimen on which this report is based was collected in present-day Slovakia (Hedenäs, pers. comm.), and no other report is known. *Coscinodon cribrosus (Hedw.) Spruce. – first published by VAJDA (1966a), this record is listed in BOROS (1968), ORBÁN and VAJDA (1983); however, the specimens (BP 71290 and BP 111088 Comit. Veszprém. In rupibus siccis pr. pag. Salföld, 12.05.1956, leg. L. Vajda) were revised to Grimmia trichophylla (B. Papp, unpubl.). *Dichodontium flavescens (Dicks.) Lindb. (Dichodontium pellucidum var. flavescens (Dicks.) Moore). – Taxonomy according to WERNER (2002); no D. flavescens could be found during revision of all specimens of D. pellucidum in BP (Erzberger, unpubl.). Until specimens are located, D. flavescens must be considered absent from the Hungarian bryoflora. Dicranella palustris (Dicks.) Crundw. ex E. F. Warb. – erroneously reported by DÜLL (1984). Didymodon fallax var. adriatica (Baumgartner) Düll. – The record of Latzel (as Barbula adriatica Baumgartner) is rejected by BOROS (1968); but Latzel’s record is repeated in PODPERA (1954), DÜLL (1984, 1992). Düll (pers. comm.) considers the variety to be presumably without taxonomic value. Didymodon icmadophilus (Müll. Hal.) K. Saito. – erroneously reported by DÜLL (1984). *Drepanocladus revolvens (anon.) Warnst. – see note under D. cossonii; not reported for Hun- gary according to HEDENÄS (2003). *Entodon concinnus (De Not.) Paris (Entodon orthocarpus (Brid.) Lindb.; Cylindrothecium concinnatum (De Not.) Schimp.). – All specimens that could be located in BP (1 specimen), EGR, SZO (4 specimens, Piers-herb.) belong to different species e.g. Pleurozium schreberi or Calliergonella cuspidata. These specimens refer to records from Kőszeg Mts; according to BOROS (1968), ORBÁN and VAJDA (1983) all other records are doubtful. Fissidens kosaninii Latzel. – reported in BOROS (1968), VAJDA (1958), missing in ORBÁN and VAJDA (1983), because the material has been revised to F. exiguus (VAJDA 1975). *Fissidens osmundoides Hedw. – The only report in ORBÁN and VAJDA (1983) (Appendix) is based on a single specimen, which has been revised to Fissidens taxifolius (B. Papp, unpubl.). Speci- men: BP 160761. Comit. Somogy, in salicetis ad margines fluvii Dráva prope pag. Drávatamási. 21.04.1979. Leg. I. Galambos, det. L. Vajda sub Fissidens osmundoides Hedw. *Grimmia donniana Sm.–MUÑOZ and PANDO (2000) quote a Hungarian specimen from BP (L. Vajda s.n.). In MUÑOZ (1998b), p. 381, probably the same specimen is quoted in more detail:

Studia bot. hung. 35, 2004 ANNOTATED CHECKLIST OF HUNGARIAN BRYOPHYTES 131

“HUNGARY, Montes Szeben, prope Paltanis, Vajda s.n. (BP–73974)”. This collection originates from Romania, outside the present territory of Hungary. No specimen of Grimmia donniana from present-day Hungary annotated by Muñoz could be located in BP. Thus, there is no evidence for the presence of G. donniana in Hungary. In ORBÁN and VAJDA (1983), this name appears erroneously (see note under G. montana). Heterocladium heteropterum Schimp. – BOROS (1968): the specimen of Szepesfalvi was revised by Boros to Pterigynandrum filiforme. Hylocomium pyrenaicum (Spruce) Lindb. – specimens have been revised to H. splendens (BOROS (1968)). *Hypnum fertile Sendtn. – A single occurrence of this species was reported by I. Galambos (ORBÁN and VAJDA (1983), Appendix). However, the specimen does not belong to H. fertile (B. Papp and P. Erzberger, unpubl.). Specimen: BP 159411. Comit. Somogy. In ripa Nagyberek, ad truncos putr., prope pago Darány. 14.11.1976. leg. I. Galambos, det. L. Vajda sub Hypnum fertile Sendtn. *Isopterygiopsis muelleriana (Schimp.) Z. Iwats. (Isopterygium muellerianum (Schimp.) A. Jaeger; Plagiothecium muellerianum Schimp.). – Two specimens under this name were located in BP, both represent different species. BP 143883: Comit. Hont. In rupibus andesit. sept. Fagetorum vallis rivi Nagyvasfazék-patak pr. Királyrét, 13.04.1958, leg. Á. Boros, rev. by B. Papp as Plagiothecium succulentum (Wilson) Lindb. (unpubl.); this locality was omitted in BOROS (1968), ORBÁN and VAJDA (1983). BP 143878 Com. Heves. In rupibus andesiticis umbrosis vallis Hórakó prope Mátraháza, 25.10.1952, leg. L. Vajda, rev. by B. Papp as Isoperygiopsis pulchella (Hedw.) Iwats. (unpubl.). Since this is the only locality mentioned in BOROS (1968), ORBÁN and VAJDA (1983), we consider I. muelleriana to be absent from the Hungarian bryoflora. Myurella tenerrima (Brid.) Lindb. – erroneously reported by DÜLL (1985). Palustriella decipiens (De Not.) Ochyra (Cratoneuron decipiens (De Not.) Loeske). – Re- jected in BOROS (1968); HEDENÄS (2003) erroneously reports this species from Hungary; the specimen on which this record is based was collected by Győrffy in present-day Slovakia (Hedenäs, pers. comm.). *Pohlia bulbifera (Warnst.) Warnst. – All specimens in BP labelled P. bulbifera proved to be different species (Erzberger, in press). *Pohlia drummondii (Müll. Hal.) A. L. Andrews (Pohlia commutata Lindb.). – Synonymy according to KOPERSKI et al. (2000); synonymies in ORBÁN and VAJDA (1983) erroneous. No specimen of P. drummondii could be found during revision in BP (Erzberger, in press). Pohlia filum (Schimp.) Mårtensson (Pohlia gracilis (Bruch et Schimp.) Lindb.). – No specimen of P. filum could be found during revision in BP (Erzberger, in press). *Pseudoleskea incurvata (Hedw.) Loeske (Pseudoleskea atrovirens (Brid.) Schimp.; Lescuraea atrovirens (Brid.) Kindb.). – The reports in BOROS (1968), ORBÁN and VAJDA (1983) are based on a single collection, which has been revised to Pseudoleskeella nervosa (Brid.) Nyholm (B. Papp, unpubl.). Specimens: BP 75667 and EGR. Agriense, in silva Corno-Quercetum mtis Rókalyuktető, pr. praed. Markaz. 22.06.1966. leg. M. Kovács, det. L. Vajda sub Lescuraea incurvata (Hedw.) Lawton var. incurvata. Pseudoleskea patens (Lindb.) Kindb. (Lescuraea atrovirens var. patens (Lindb.) Mönk.). – The reports in BOROS (1968), ORBÁN and VAJDA (1983) are based on a single collection, which has been revised to Thuidium abietinum (B. Papp, unpubl.). Specimens: BP 127617 and EGR. Comit. Heves. In rupestribus andesit. “Kecskebérc” prope Mátraháza. 28.10.1952. leg. Á. Boros. Racomitrium aciculare (Hedw.) Brid. – reports in DÜLL (1985, 1992) are purely conjectural (Düll in litt.). Racomitrium lanuginosum (Hedw.) Brid. – rejected in BOROS (1968).

Studia bot. hung. 35, 2004 132 ERZBERGER, P. and PAPP, B.

Rhytidiadelphus loreus (Hedw.) Warnst. – BOROS (1968) doubts the only report by Győrffy. Scorpiurium circinatum (Brid.) M. Fleisch. et Loeske (Eurhynchium circinatum (Brid.) Schimp.). – BOROS (1968) rejects the only report by Latzel. Sphagnum molle Sull. – BOROS (1968) rejects the only report by Szepesfalvi. Specimens in BP (BP 6353, BP 3017/S, BP 329/S and BP 6351: Hungaria occid. In sphagneto piscinarum pr. pag. Lesenceistvánd ad lac. Balaton, 01.09.1933, leg. J. Szepesfalvi) represent S. capillifolium (rev. J. Klawitter, Berlin). Splachnum ampullaceum L. ex Hedw. – Erroneously reported by DÜLL (1985). Stegonia latifolia (Schwägr.) Venturi ex Broth. – Erroneously reported by DÜLL (1985) (source: Limpricht: “Liptauer Alpen“). *Tortella flavovirens (Bruch) Broth. – var. viridiflava (De Not.) Casares-Gil. (= T. esterelensis (Roth) Giacom.); in BOROS (1968) as T. viridiflava (De Not.) Broth. (= subsp. viridiflava (De Not.) Giacom.); all specimens located in BP were revised to other species of Tortella (B. Papp, unpubl.). Tortella humilis (Hedw.) Jenn. (T. caespitosa (Schwägr.) Limpr.). – According to BOROS (1968) reports of this species are based on misidentified T. inclinata. Tortella nitida (Lindb.) Broth. – According to BOROS (1968) reports of this species are based on misidentified T. tortuosa (revised by Reimers). Tortula cuneifolia (Dicks.) Turner. – The report in DÜLL (1984) is based on PODPERA (1954), but there reference is given to Loeske, who rejected a report by Győrffy (“Hajdúszoboszló”, 1932) as erroneous, as communicated by Boros (in litt.). Warnstorfia fluitans (Hedw.) Loeske (Drepanocladus fluitans (Hedw.) Warnst.). – Estimated doubtful in BOROS (1968), omitted in ORBÁN and VAJDA (1983); erroneously reported in HEDENÄS (2003) for Hungary; the specimen on which the report is based, was collected in present-day Slovakia (Hedenäs, pers. comm.).

APPENDIX 2. TAXA OF DOUBTFUL OCCURRENCE IN HUNGARY (not listed in main list)

HEPATICAE

Moerckia hibernica (Hook.) Gottsche (Moerckia flotoviana (Nees) Schiffn.). – a single record by Szepesfalvi, but there exists no specimen; omitted in ORBÁN and VAJDA (1983) but accepted by BOROS (1968), RAJCZY (1990), SÖDERSTRÖM et al. (2002); sometimes spelled Moerchia, e.g. DAMSHOLT (2002).

MUSCI

Atrichum tenellum (Röhl.) Bruch et Schimp. – doubtful according to BOROS (1968), missing in ORBÁN and VAJDA (1983). Barbula crocea (Brid.) F. Weber et D. Mohr – doubtful according to BOROS (1968), missing in ORBÁN and VAJDA (1983), but reported by MARSTALLER (1993a); no specimen collected by Marstaller in Hungary was sent upon request from JE.

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Barbula enderesii Garov. – doubtful according to BOROS (1968), missing in ORBÁN and VAJDA (1983). Bryum barnesii J. B. Wood. – reported by DÜLL (1985), but since B. barnesii was considered by Düll as synonym of B. bicolor, this record is doubtful. No specimen seen. Bryum cyclophyllum (Schwägr.) Bruch et Schimp. – doubtful according to BOROS (1968), missing in ORBÁN and VAJDA (1983). Cratoneuron curvicaule (Jur.) G. Roth. – not reported for Hungary according to HEDENÄS (2003), but reported by DÜLL (1985); however, this report is unlikely because in Central Europe C. curvicaule is a montane-alpine taxon (HEDENÄS 2003, GRIMS 1999); no specimen seen. Cynodontium gracilescens (F. Weber et D. Mohr) Schimp. – the only specimen determined by Latzel is rejected in BOROS (1968) (Boros revised it to Dichodontium pellucidum); however, Boros’s determination is probably wrong (Erzberger, unpubl.). The identity of the specimen still awaits reso- lution. Dicranum bergeri Blandow ex Hoppe. – reported by DÜLL (1984), but unlikely, since it is a boreo-montane taxon, no specimen seen. Dicranum fuscescens Sm. – the only reports of the taxon as var. falcifolium Braithw. and var. congestum Brid. by Latzel are estimated doubtful in BOROS (1968); not reported in ORBÁN and VAJDA (1983); in DÜLL (1984) D. fuscescens is not reported, whereas D. congestum is, but in DÜLL (1992) this report is obviously thought unreliable, because it “has to be verified for Hu.”. Encalypta alpina Sm. – doubtful according to BOROS (1968), not reported in ORBÁN and VAJDA (1983), nor in DÜLL (1984). Encalypta rhaptocarpa Schwägr. – doubtful according to BOROS (1968), not reported in ORBÁN and VAJDA (1983), nor in DÜLL (1984); according to a remark in BOROS (1968) the specimens from the Bükk Mts are intermediate between E. rhaptocarpa and E. vulgaris, perhaps aff. E. trachymitria Ripart (cf. NYHOLM 1998). The true identity of these specimens remains to be estab- lished. *Entosthodon obtusus (Hedw.) Lindb. (Funaria obtusa (Hedw.) Lindb.). – The report in ORBÁN and VAJDA (1983) is based on a single record by Degen, which is questioned in BOROS (1968). A doubtful specimen from a different locality was revised to E. cf. fascicularis (B. Papp, unpubl.). Specimen: BP 114587. Comit. Győr. In loco inculto paludoso prope pag. Nagyszentpál- puszta, prope Győrszemere. 08.07.1941, leg. S. Polgár, det. Á. Boros. No other specimen of E. obtusus from Hungary could be located in BP, EGR or SZO. Grimmia decipiens (Schultz) Lindb. – in BOROS (1968) one report by Szepesfalvi and one report by Latzel are estimated doubtful, reports from Mátra Mts are rejected; omitted in ORBÁN and VAJDA (1983) and DÜLL (1984); not reported for Hungary in MUÑOZ and PANDO (2000); no specimen seen. Grimmia incurva Schwägr. – MUÑOZ and PANDO (2000) quote a specimen (A. Boros s.n., BP) that could not be located. MAIER and GEISSLER (1995) report the altitudinal range of G. incurva between 1,900 and 3,080 m a.s.l. The occurrence in present-day Hungary thus seems highly unlikely. Until appropriate material is located, the occurrence of G. incurva in Hungary cannot be accepted. *Grimmia sessitana De Not. (Grimmia reflexidens Müll. Hal.; Grimmia alpestris (F. Weber et D. Mohr) Schleich. var. subsulcata (Limpr.) Broth.). – Nomenclature according to Maier (2002b); MUÑOZ (1998b) quotes G. reflexidens Müll. Hal. as the correct name for G. sessitana De Not., but this is rejected by MAIER (2002b); in MUÑOZ (1998b) as well as in MUÑOZ and PANDO (2000) Hun- gary is missing from the world distribution, but the species is reported in ORBÁN and VAJDA (1983) (as G. sessitana f. subsulcata) and in BOROS (1968) (as Grimmia alpestris var. subsulcata). How- ever, the specimens in BP belong to G. funalis, revision by J. Muñoz and E. Maier. Until appropriate material is located, G. sessitana must be considered to be absent from the Hungarian bryoflora.

Studia bot. hung. 35, 2004 134 ERZBERGER, P. and PAPP, B.

Hypnum andoi A. J. E. Sm. – reported as doubtful in DÜLL (1985), no specimen seen; but could perhaps be expected. Hypnum pratense W. D. J. Koch ex Spruce. – reported as doubtful in BOROS (1968), omitted in ORBÁN and VAJDA (1983), DÜLL (1985). Meesia longiseta Hedw. – reported as doubtful in BOROS (1968), omitted in ORBÁN and VAJDA (1983), DÜLL (1985). Mnium spinosum (Voit) Schwägr. – reported as doubtful in BOROS (1968), omitted in ORBÁN and VAJDA (1983), DÜLL (1985), no Hungarian records reported in ORBÁN (1975). Pohlia sphagnicola (Bruch et Schimp.) Broth. – reported as doubtful in BOROS (1968), omitted in ORBÁN and VAJDA (1983), DÜLL (1985). Pseudoleskeella tectorum (Brid.) Kindb. ex Broth. (Leskea tectorum (Brid.) Lindb.). – reported as doubtful in BOROS (1968), omitted in ORBÁN and VAJDA (1983), but reported in DÜLL (1985). Schistidium confertum (Funck) Bruch et Schimp. – in BOROS (1968) transient forms to S. brunnescens are reported; specimens should be examined. Schistidium rivulare (Brid.) Podp. – missing from BOROS (1968), ORBÁN and VAJDA (1983), but listed as vulnerable by RAJCZY (1990); specimens should be checked. Sphagnum rubellum Wilson. – neither in BOROS (1968) nor ORBÁN and VAJDA (1983), but in RAJCZY (1990) “endangered”; no specimen could be located in BP and EGR; a recent collection by E. Szurdoki was revised to S. capillifolium (rev. J. Klawitter, Berlin and P. Erzberger, unpubl.); the taxonomic status of S. rubellum and its separation from S. capillifolium are controversial (MCQUEEN 1989, HILL 1978, KOPERSKI et al. 2000). Tortula canescens Mont. – not reported by BOROS (1968), ORBÁN and VAJDA (1983), but in MARSTALLER (1993a); from JE a single specimen (Südungarn Villanier Gebirge Nagyharsany-h., 27.05.88, leg. R. Marstaller) consisting of a few sterile plants only was obtained, that does not allow unequivocal determination. Tortula princeps De Not. (Syntrichia princeps (De Not.) Mitt.). – reported in BOROS (1968), ORBÁN and VAJDA (1983) and MARSTALLER (1993a), but all specimens were revised by J. Kučera, České Budejovice, to T. crinita, among them a specimen from JE (Ungarn Villanier Gebirge Nagyharsány-hegy, 06.06.88, leg. R. Marstaller). Weissia squarrosa (Nees et Hornsch.) Müll. Hal. – reported as doubtful in BOROS (1968), omitted in ORBÁN and VAJDA (1983), DÜLL (1984). Weissia triumphans (De Not.) M. O. Hill. – not reported in BOROS (1968), ORBÁN and VAJDA (1983), but reported by MARSTALLER (1993a), no specimen collected by Marstaller in Hungary was sent upon request from JE. Zygodon viridissimus (Dicks.) Brid. – var. viridissimus may be included in the reports in BOROS (1968), ORBÁN and VAJDA (1983); reported by MARSTALLER (1993a, b); a single specimen collected by Marstaller in Hungary was sent upon request from JE, which proved to be Z. rupestris (Erzberger, unpubl.): JE Ungarn Baranya Wälder um Dravafok, 07.06.89, leg. R. Marstaller.

Studia bot. hung. 35, 2004 ANNOTATED CHECKLIST OF HUNGARIAN BRYOPHYTES 135

APPENDIX 3. TAXA OF DOUBTFUL OCCURRENCE IN HUNGARY (listed in main list, see notes under individual taxa)

HEPATICAE

Cephalozia lacinulata J. B. Jack ex Spruce Marsupella sprucei (Limpr.) Bernet Metzgeria conjugata Lindb. subsp. simplex (Müll. Frib.) R. M. Schust. (Metzgeria simplex Lorb. ex Müll. Frib.) Metzgeria fruticulosa (Dicks.) A. Evans Radula lindenbergiana Gottsche ex C. Hartm. Riccia duplex Lorb. ex Müll. Frib.

MUSCI

Brachythecium campestre (Müll. Hal.) Schimp. Ceratodon conicus (Hampe) Lindb. Fissidens gymnandrus Büse Pterygoneurum crossidioides W. Frey, Herrnst. et Kürschner Racomitrium affine (F. Weber et D. Mohr) Lindb. Racomitrium obtusum (Brid.) Brid.

APPENDIX 4. TAXA RECORDED AFTER 1983

HEPATICAE

Anastrophyllum hellerianum (Lindenb.) R. M. Schust. – ÓDOR (2000). Riccia crinita Taylor (Riccia intumescens (Bisch.) Underw.; Riccia canescens Steph.; Riccia trichocarpa M. Howe). – MARSTALLER (1993a); confirmed specimen see main list.

MUSCI

Aloina aloides (Schultz) Kindb. – MARSTALLER (1993a); confirmed specimens see main list. Barbula indica (Hook.) Spreng. – GALAMBOS (1992). Bryum gemmiferum R. Wilczek et Demaret. – ZANTEN (1999a). Bryum rubens Mitt. – first collected by I. Galambos (unpubl., see note in main list); JAKAB (1997), ZANTEN (1999a), PAPP and RAJCZY (1999, 2000). Bryum violaceum Crundw. et Nyholm. – ZANTEN (1999a). Ceratodon conicus (Hampe) Lindb. – ZANTEN (1999b).

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Crossidium crassinerve (De Not.) Jur. – reported by ORBÁN (1991: found by Galambos), DÜLL (1984). Crossidium laxefilamentosum W. Frey et Kürschner. – PÓCS et al. (2004). Ditrichum crispatissimum (Müll. Hal.) Paris – ERZBERGER (2001). Drepanocladus sordidus (Müll. Hal.) Hedenäs – HEDENÄS (2003) Ephemerum minutissimum Lindb. (E. serratum var. minutissimum (Lindb.) Grout) – B. Papp (unpubl.) Ephemerum recurvifolium (Dicks.) Boulay. – NOVOTNY (1988). Ephemerum sessile (Bruch) Müll. Hal. – confirmed specimen see note in main list. Grimmia dissimulata E. Maier. – confirmed specimens see note in main list. Hedwigia stellata Hedenäs – ERZBERGER (1996). Orthotrichum sprucei Mont. – ERZBERGER and PAPP (2000). Phascum leptophyllum Müll. Hal. – ZANTEN (2000). Pohlia andalusica (Höhn.) Broth. – (ERZBERGER, in press). Pterygoneurum compactum M. J. Cano, J. Guerra et Ros. – PÓCS (1999). Pterygoneurum crossidioides W. Frey, Herrnst. et Kürschner. – PÓCS (1999). Pterygoneurum squamosum Segarra et Kürschner. – PÓCS (1999). Seligeria campylopoda Kindb. – confirmed specimens see note in main list. Seligeria trifaria (Brid.) Lindb. var. longifolia (Broth.) Ochyra et Gos. – confirmed specimen see note in main list. Tortella bambergeri (Schimp.) Broth. – MARSTALLER (1993a) from Villany Mts, Erzberger (unpubl.) from Bükk Mts. Tortula brevissima Schiffn. – ERZBERGER (1998). Tortula caninervis (Mitt.) Broth. subsp. spuria var. spuria (J. J. Amann) W. A. Kramer. – TÓTH (1987). Tortula norvegica (F. Weber) Lindb. – TÓTH (1987), leg. L. Meinunger. Tortula papillosissima (Copp.) Broth. var. papillosissima and var. submamillosa (W. A. Kramer) Heinrichs et Caspari. – TÓTH (1987)

APPENDIX 5. TABLE OF USEFUL SYNONYMS

SYNONYM ACCEPTED NAME Hepaticae Alicularia scalaris (Gray) Corda Nardia scalaris Gray Aneura latifrons Lindb. Riccardia latifrons (Lindb.) Lindb. Aneura multifida Dumort. Riccardia multifida (L.) Gray Aneura palmata Dumort. Riccardia palmata (Hedw.) Carruth. Aneura sinuata Dumort. Riccardia chamedryfolia (With.) Grolle Anthoceros crispulus auct. non (Mont.) Douin Anthoceros agrestis Paton Anthoceros laevis auct. Phaeoceros carolinianus (Michx.) Proskauer Anthoceros punctatus auct. non L. Anthoceros agrestis Paton Barbilophozia obtusa (Lindb.) Meyl. Lophozia obtusa (Lindb.) A. Evans Calypogeia meylanii H. Buch Calypogeia integristipula Steph. Calypogeia trichomanis auct. Calypogeia azurea Stotler et Crotz Cephalozia lammersiana (Huebener) F. Lees Cephalozia bicuspidata (L.) Dumort.

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SYNONYM ACCEPTED NAME Cephalozia media Lindb. Cephalozia lunulifolia (Dumort.) Dumort. Cephaloziella starkei (Funck) Schiffn. Cephaloziella divaricata (Sm.) Schiffn. Cephaloziella subdentata Warnst. Cephaloziella spinigera (Lindb.) Warnst. Chiloscyphus coadunatus (Sw.) J. J. Engel et Lophocolea bidentata var. bidentata (L.) R. M. Schust. Dumort. Chiloscyphus latifolius (Nees) J. J. Engel et R. Lophocolea bidentata var. rivularis (Raddi) M. Schust. Schiffn. Chiloscyphus minor (Nees) J. J. Engel et R. M. Lophocolea minor Nees Schust. Chiloscyphus profundus (Nees) J. J. Engel et Lophocolea heterophylla (Schrad.) Dumort. R. M. Schust. Clevea hyalina (Sommerf.) Lindb. Athalamia hyalina (Sommerf.) S. Hatt. Eucalyx hyalinus (Lyell) Breidl. Jungermannia hyalina Lyell Fimbriaria fragrans Nees Asterella saccata (Wahlenb.) A. Evans Fimbriaria saccata (Wahlenb.) Nees Asterella saccata (Wahlenb.) A. Evans Fossombronia dumortieri (Huebener et Genth) Fossombronia foveolata Lindb. Lindb. Frullania cleistostoma Schiffn. et Wollny Frullania inflata Gottsche Frullania saxicola Austin Frullania inflata Gottsche Grimaldia barbifrons Bischof Mannia fragrans (Balbis) Frye et L. Clark Grimaldia fragrans (Balbis) Corda Mannia fragrans (Balbis) Frye et L. Clark Grimaldia rupestris (Nees) Lindenb. Mannia triandra (Scop.) Grolle Haplozia caespiticia (Lindenb.) Dumort. Jungermannia caespiticia Lindenb. Haplozia crenulata (Sm.) Dumort. Jungermannia gracillima Sm. Haplozia lanceolata (L.) Dumort. Jungermannia leiantha Grolle Haplozia pumila (With.) Dumort. Jungermannia pumila With. Haplozia sphaerocarpa (Hook.) Dumort. Jungermannia sphaerocarpa Hook. Isopaches bicrenatus (Hoffm.) H. Buch Lophozia bicrenata (Hoffm.) Dumort. Jungermannia amakawana Grolle Jungermannia subulata A. Evans Jungermannia lanceolata L. Jungermannia leiantha Grolle Leiocolea alpestris (F. Weber) Isov. Leiocolea collaris (Nees) Schljakov Leiocolea muelleri (Lindenb.) Jörg. Leiocolea collaris (Nees) Schljakov Lejeunea serpyllifolia Lib. Lejeunea cavifolia (Ehrh.) Lindb. Liochlaena lancoelata (L.) Nees Jungermannia leiantha Grolle Lophocolea alata Larter Lophocolea bidentata var. bidentata (L.) Dumort. Lophocolea bidentata auct. non (L.) Dumort., Lophocolea bidentata var. rivularis (Raddi) sensu Limpricht, BOROS (1968), ORBÁN Schiffn. and VAJDA (1983) Lophocolea cuspidata auct. non (Nees) Lophocolea bidentata var. bidentata (L.) Limpr., sensu Limpricht, BOROS (1968), Dumort. ORBÁN and VAJDA (1983) Lophozia alpestris auct. Lophozia sudetica (Huebener) Grolle Lophozia badensis (Gottsche) Schiffn. Leiocolea badensis (Gottsche) Jörg. Lophozia collaris (Nees) Dumort. Leiocolea collaris (Nees) Schljakov Lophozia confertifolia Schiffn. Lophozia wenzelii (Nees) Steph. Lophozia gracillima H. Buch Lophozia ascendens (Warnst.) R. M. Schust.

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SYNONYM ACCEPTED NAME Lophozia guttulata (Lindb. et H. Arnell) Evans Lophozia longiflora (Nees) Schiffn. Lophozia heterocolpos (Hartm.) M. Howe Leiocolea heterocolpos (Hartm.) H. Buch Lophozia muelleri (Lindenb.) Dumort. Leiocolea collaris (Nees) Schljakov Lophozia porphyroleuca auct. Lophozia longiflora (Nees) Schiffn. Lophozia quinquedentata (Huds.) Cogn. Tritomaria quinquedentata (Huds.) H. Buch Madotheca cordaeana (Huebener) Dumort. Porella cordaeana (Huebener) Moore Madotheca laevigata (Schrad.) Dumort. Porella arboris-vitae (With.) Grolle Madotheca platyphylla (L.) Dumort. Porella platyphylla (L.) Pfeiff. Madotheca platyphylloidea (Schwein.) Porella platyphylla (L.) Pfeiff. Dumort. Mannia rupestris (Nees) Frye et L. Clark Mannia triandra (Scop.) Grolle Marchantia aquatica (Nees) Burgeff Marchantia polymorpha L. subsp. polymorpha Marchantia polymorpha auct. Marchantia polymorpha L. subsp. ruderalis Bischl. et Boisselier Marsupella hungarica Boros et Vajda Marsupella funckii (F. Weber et D. Mohr) Dumort. Metzgeria pubescens (Schrank) Raddi Apometzgeria pubescens (Schrank) Kuwah. Metzgeria simplex Müll. Frib. Metzgeria conjugata Lindb. subsp. simplex (Müll. Frib.) R. M. Schust. Oxymitra paleacea Lindenb. Oxymitra incrassata (Brot.) Sergio et Sim-Sim Pellia calycina Taylor Pellia endiviifolia (Dicks.) Dumort. Pellia fabbroniana auct. Pellia endiviifolia (Dicks.) Dumort. Plagiochila major (Nees) S. W. Arnell Plagiochila asplenioides (L.) Dumort. Plectocolea hyalina (Lyell) Mitt. Jungermannia hyalina Lyell Porella platyphylloidea (Schwein.) Lindb. Porella platyphylla (L.) Pfeiff. Riccardia pinguis (L.) Gray Aneura pinguis (L.) Dumort. Riccardia sinuata (Hook.) Trevis Riccardia chamedryfolia (With.) Grolle Riccia baumgartneri Schiffn. Riccia subbifurca Crozals Riccia bischoffii Huebener Riccia ciliifera Lindenb. Riccia crystallina auct. non L. Riccia cavernosa Hoffm. Riccia pseudo-frostii Schiffn. Riccia huebeneriana Lindenb. Riccia pseudopapillosa Steph. Riccia papillosa Moris Riccia subtumida Milde Riccia ciliata Hoffm. Scapania microphylla Warnst. Scapania lingulata H. Buch Scapania nemorosa Dumort. Scapania nemorea (L.) Grolle Scapania parvifolia Warnst. Scapania scandica (Arnell et H. Buch) Macvicar Solenostoma caespiticium (Lindenb.) Steph. Jungermannia caespiticia Lindenb. Solenostoma crenulatum Mitt. Jungermannia gracillima Sm. Solenostoma lanceolata Steph. Jungermannia leiantha Grolle Solenostoma pumilum (With.) Müll. Frib. Jungermannia pumila With. Solenostoma riparia (Taylor) Dumort. Jungermannia atrovirens Dumort. Solenostoma sphaerocarpum (Hook.) Steph. Jungermannia sphaerocarpa Hook. Solenostoma triste (Nees) Müll. Frib. Jungermannia atrovirens Dumort. Sphaerocarpos californicus Austin Sphaerocarpos texanus Austin Sphenolobus exsectiformis (Breidl.) Steph. Tritomaria exsectiformis (Breidl.) Loeske Sphenolobus exsectus (Schmidel) Steph. Tritomaria exsecta (Schmidel) Loeske Sphenolobus michauxii (F. Weber) Steph. Anastrophyllum michauxii (F. Weber) H. Buch

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SYNONYM ACCEPTED NAME Sphenolobus minutus (Schreb.) Steph. Anastrophyllum minutum (Schreb.) R. M. Schust. Tesselina pyramidata Dumort. Oxymitra incrassata (Brot.) Sergio et Sim-Sim Musci Abietinella abietina (Hedw.) M. Fleisch. Thuidium abietinum (Hedw.) Schimp. Acrocladium cuspidatum Lindb. Calliergonella cuspidata (Hedw.) Loeske Amblystegiella confervoides (Brid.) Loeske Amblystegium confervoides (Brid.) Schimp. Amblystegiella subtilis (Hedw.) Loeske Amblystegium subtile (Hedw.) Schimp. Amblystegium compactum (Müll. Hal.) Austin Conardia compacta (Müll. Hal.) H. Rob. Amblystegium densum Milde Conardia compacta (Müll. Hal.) H. Rob. Amblystegium filicinum (Hedw.) De Not. Cratoneuron filicinum (Hedw.) Spruce Amblystegium juratzkanum Schimp. Amblystegium serpens (Hedw.) Schimp. Amblystegium kochii Schimp. Amblystegium humile (P. Beauv.) Crundw. Amblystegium riparium (Hedw.) Schimp. Leptodictyum riparium (Hedw.) Warnst. Amblystegium saxatile Schimp. Amblystegium radicale (P. Beauv.) Schimp. Amblystegium trichopodium (Schultz) C. Amblystegium humile (P. Beauv.) Crundw. Hartm. Andreaea petrophila Fürnr. Andreaea rupestris Hedw. Anisothecium crispum (Hedw.) C. E. O. Jensen Dicranella crispa (Hedw.) Schimp. Anisothecium rigidulum (Hedw.) C. E. O. Dicranella humilis R. Ruthe Jensen Anisothecium rubrum Lindb. Dicranella varia (Hedw.) Schimp. Anisothecium rufescens (Dicks.) Lindb. Dicranella rufescens (Dicks.) Schimp. Anisothecium schreberianum (Hedw.) Dixon Dicranella schreberiana (Hedw.) Hilf. ex H. A. Crum et L. E. Anderson Anisothecium staphylinum (H. Whitehouse) Dicranella staphylina H. Whitehouse Sipman, Rubers et Rieman Anisothecium vaginale (With.) Loeske Dicranella crispa (Hedw.) Schimp. Anisothecium varium (Hedw.) Mitt. Dicranella varia (Hedw.) Schimp. Anomodon apiculatus Schimp. Anomodon rugelii (Müll. Hal.) Keissl. Arbuscula alopecura (Hedw.) H. A. Crum, Thamnobryum alopecurum (Hedw.) Gangulee Steere et L. E. Anderson Archidium phascoides Brid. Archidium alternifolium (Hedw.) Schimp. Astomum crispum (Hedw.) Hampe Weissia longifolia Mitt. Atrichum haussknechtii Jur. et Milde Atrichum undulatum var. gracilisetum Besch. Barbula acuta (Brid.) Brid. Didymodon acutus (Brid.) K. Saito Barbula cordata (Jur.) Braithw. Didymodon cordatus Jur. Barbula cylindrica (Taylor) Schimp. Didymodon insulanus (De Not.) M. O. Hill Barbula fallax Hedw. Didymodon fallax (Hedw.) R. H. Zander Barbula gracilis Schwägr. Didymodon acutus (Brid.) K. Saito Barbula hornschuchiana Schultz Pseudocrossidium hornschuchianum (Schultz) R. H. Zander Barbula lurida (Spreng.) Lindb. Didymodon luridus Hornsch. ex Spreng. Barbula recurvirostra (Hedw.) Dixon Bryoerythrophyllum recurvirostrum (Hedw.) P. C. Chen Barbula reflexa (Brid.) Brid. Didymodon ferrugineus (Besch.) M. O. Hill Barbula revoluta Brid. Pseudocrossidium revolutum (Brid.) R. H. Zander Barbula rigidula (Hedw.) Mitt. Didymodon rigidulus Hedw.

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SYNONYM ACCEPTED NAME Barbula sinuosa (Mitten) Garov. Didymodon sinuosus (Mitt.) Delogne Barbula spadicea (Mitten) Braithw. Didymodon spadiceus (Mitt.) Limpr. Barbula tophacea (Brid.) Mitt. Didymodon tophaceus (Brid.) Lisa Barbula trifaria auct. non (Hedw.) Mitt. Didymodon luridus Hornsch. ex Spreng. Barbula vinealis Brid. Didymodon vinealis (Brid.) R. H. Zander Barbula vinealis var. cylindrica (Taylor) Boulay Didymodon insulanus (De Not.) M. O. Hill Bartramia norvegica Lindb. Bartramia halleriana Hedw. Bartramia oederi Brid. Plagiopus oederianus (Sw.) H. A. Crum et L. E. Anderson Brachythecium laetum (Brid.) Schimp. Brachythecium oxycladum (Brid.) A. Jaeger Brachythecium lanceolatum Warnst. Brachythecium salebrosum (F. Weber et D. Mohr) Schimp. Brachythecium rotaeanum De Not. Brachythecium capillaceum (F. Weber et D. Mohr) Giacom. Brachythecium salebrosum (F. Weber et D. Brachythecium capillaceum (F. Weber et D. Mohr) Schimp. var. capillaceum F. Mohr) Giacom. Weber et D. Mohr Bryum affine Lindb. et Arnell Bryum creberrimum Taylor Bryum alpinum var. mildeanum (Jur.) Podp. Bryum mildeanum Jur. Bryum angustirete Macoun Bryum algovicum Müll. Hal. Bryum atropurpureum Bruch et Schimp. Bryum bicolor Dicks. Bryum badium (Brid.) Schimp. Bryum caespiticium Hedw. Bryum capillare subsp. elegans (Brid.) Podp. Bryum elegans Brid. Bryum cernuum Lindb. Bryum uliginosum (Brid.) Bruch et Schimp. Bryum cirrhatum Hoppe et Hornsch. Bryum pallescens Schleich. ex Schwägr. Bryum duvalii Voit Bryum weigelii Spreng. Bryum flaccidum Brid. Bryum laevifilum Syed Bryum inclinatum (Brid.) Blandow Bryum imbricatum (Schwägr.) Bruch et Schimp. Bryum murale Wilson Bryum radiculosum Brid. Bryum pendulum (Hornsch.) Schimp. Bryum algovicum Müll. Hal. Bryum ventricosum Relhan Bryum pseudotriquetrum (Hedw.) P. Gaertn., B. Mey. et Scherb. Buxbaumia indusiata Brid. Buxbaumia viridis (Lam. et DC.) Brid. ex Moug. et Nestl. Calliergon cuspidatum Kindb. Calliergonella cuspidata (Hedw.) Loeske Calliergonella lindbergii (Mitt.) Hedenäs Hypnum lindbergii Mitt. Camptothecium lutescens (Hedw.) Schimp. Homalothecium lutescens (Hedw.) H. Rob. Camptothecium nitens (Hedw.) Schimp. Tomentypnum nitens (Hedw.) Loeske Camptothecium philippeanum (Spruce) Kindb. Homalothecium philippeanum (Spruce) Schimp. Camptothecium sericeum (Hedw.) Kindb. Homalothecium sericeum (Hedw.) Schimp. Camptothecium trichoides Broth. Tomentypnum nitens (Hedw.) Loeske Campyliadelphus chrysophyllus (Brid.) Kanda Campylium chrysophyllum (Brid.) Lange Campyliadelphus elodes (Lindb.) Kanda Campylium elodes (Lindb.) Kindb. Campylium helodes (Spruce) Broth. Campylium elodes (Lindb.) Kindb. Campylium radicale (P. Beauv.) Grout Amblystegium radicale (P. Beauv.) Schimp. Campylopus torfaceus Bruch et Schimp. Campylopus pyriformis (Schultz) Brid. Catharinea angustata (Brid.) Brid. Atrichum angustatum (Brid.) Bruch et Schimp.

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SYNONYM ACCEPTED NAME Catharinea haussknechtii Broth. Atrichum undulatum var. gracilisetum Besch. Catharinea undulata (Hedw.) Weber et Mohr Atrichum undulatum (Hedw.) P. Beauv. Chenia leptophylla (Müll. Hal.) R. H. Zander Phascum leptophyllum Müll. Hal. Chenia rhizophylla (Sakurai) R. H. Zander Phascum leptophyllum Müll. Hal. Chrysohypnum calcareum (Crundw. et Campylium calcareum Crundw. et Nyholm Nyholm) Boros Chrysohypnum chrysophyllum (Brid.) Loeske Campylium chrysophyllum (Brid.) Lange Chrysohypnum helodes (Spruce) Loeske Campylium elodes (Lindb.) Kindb. Chrysohypnum hygrophilum (Jur.) Loeske Amblystegium radicale (P. Beauv.) Schimp. Chrysohypnum polygamum (Schimp.) Loeske Campylium polygamum (Schimp.) C. E. O. Jensen Chrysohypnum radicale (P. Beauv.) Boros Amblystegium radicale (P. Beauv.) Schimp. Chrysohypnum sommerfeltii auct. Campylium calcareum Crundw. et Nyholm Chrysohypnum stellatum (Hedw.) Loeske Campylium stellatum (Hedw.) C. E. O. Jensen Cinclidotus minor Lindb. Cinclidotus fontinaloides (Hedw.) P. Beauv. Cinclidotus mucronatus (Brid.) Guim. Dialytrichia mucronata (Brid.) Broth. Cinclidotus nigricans (Brid.) Wijk et Margad. Cinclidotus riparius (Brid.) H. Arnell Cirriphyllum crassinervium (Wilson) Loeske Eurhynchium crassinervium (Wilson) Schimp. et M. Fleisch. Cirriphyllum filiforme Broth. Eurhynchium flotowianum (Sendtn.) Kartt. Cirriphyllum germanicum (Grebe) Loeske et Rhynchostegiella tenuicaulis (Spruce) Kartt. M. Fleisch. Cirriphyllum reichenbachianum (Huebener) Eurhynchium flotowianum (Sendtn.) Kartt. Wijk et Margad. Cirriphyllum tenuinerve Wijk et Margad. Cirriphyllum tommasinii (Boulay) Grout Cirriphyllum vaucheri Loeske et M. Fleisch. Cirriphyllum tommasinii (Boulay) Grout Cirriphyllum velutinoides (Schimp.) Loeske et Eurhynchium flotowianum (Sendtn.) Kartt. M. Fleisch. Cratoneuron commutatum (Hedw.) G. Roth Palustriella commutata (Hedw.) Ochyra Cynodontium capillaceum Hedw. Distichium capillaceum (Hedw.) Bruch et Schimp. Cynodontium schisti (F. Weber et D. Mohr) Cnestrum schisti (F. Weber et D. Mohr) I. Hagen Lindb. Desmatodon convolutus (Brid.) Grout Tortula atrovirens (Sm.) Lindb. Dicranella curvata (Hedw.) Schimp. Dicranella subulata (Hedw.) Schimp. Dicranella rubra Lindb. Dicranella varia (Hedw.) Schimp. Dicranella schreberi Schimp. Dicranella schreberiana (Hedw.) Hilf. ex H. A. Crum et L. E. Anderson Dicranella secunda Lindb. Dicranella subulata (Hedw.) Schimp. Dicranella varia var. tenella Bruch et Schimp. Dicranella humilis R. Ruthe Dicranodontium longirostre (F. Weber et D. Dicranodontium denudatum (Brid.) E. Britton Mohr) Bruch et Schimp. Dicranum longifolium Ehrh. ex Hedw. Paraleucobryum longifolium (Hedw.) Loeske Dicranum palustre Bruch et Schimp. Dicranum bonjeanii De Not. Dicranum rugosum (Funck) Hoffm. ex Brid. Dicranum polysetum Sw. Dicranum strictum D. Mohr Dicranum tauricum Sapjegin Dicranum undulatum Ehrh. ex F. Weber et D. Dicranum polysetum Sw. Mohr

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SYNONYM ACCEPTED NAME Didymodon austriacus Schiffn. et Baumgartner Didymodon cordatus Jur. Didymodon rigidulus var. glaucus (Ryan) Didymodon glaucus Ryan Wijk. et Margad. Didymodon rubellus Bruch et Schimp. Bryoerythrophyllum recurvirostrum (Hedw.) P. C. Chen Didymodon trifarius auct. non (Hedw.) Röhl. Didymodon luridus Hornsch. ex Spreng. Diphyscium sessile Lindb. Diphyscium foliosum (Hedw.) D. Mohr Distichium montanum I. Hagen Distichium capillaceum (Hedw.) Bruch et Schimp. Ditrichum flexicaule var. longifolium (J. E. Ditrichum crispatissimum (Müll. Hal.) Paris Zetterst.) I. Hagen Ditrichum flexicaule var. sterile (De Not.) Ditrichum crispatissimum (Müll. Hal.) Paris Limpr. Ditrichum giganteum R. S. Williams Ditrichum crispatissimum (Müll. Hal.) Paris Ditrichum gracile (Mitt.) O. Kuntze Ditrichum crispatissimum (Müll. Hal.) Paris Ditrichum homomallum (Hedw.) Hampe Ditrichum heteromallum (Hedw.) E. Britton Ditrichum tenuifolium Lindb. Ditrichum cylindricum (Hedw.) Grout Ditrichum tortile (Schrad.) Brockm. Ditrichum pusillum (Hedw.) Hampe Dolichotheca seligeri (Brid.) Loeske Herzogiella seligeri (Brid.) Z. Iwats. Dolichotheca silesiaca (F. Weber et D. Mohr) Herzogiella seligeri (Brid.) Z. Iwats. M. Fleisch. Drepanocladus exannulatus (Schimp.) Warnst. Warnstorfia exannulata (Schimp.) Loeske Drepanocladus intermedius (Lindb.) Warnst. Drepanocladus cossonii (Schimp.) Loeske Drepanocladus polygamus (Schimp.) Hedenäs Campylium polygamum (Schimp.) C. E. O. Jensen Drepanocladus uncinatus (Hedw.) Warnst. Sanionia uncinata (Hedw.) Loeske Drepanocladus vernicosus (Mitt.) Warnst. Hamatocaulis vernicosus (Mitt.) Hedenäs Dryptodon hartmanii (Schimp.) Limpr. Grimmia hartmanii Schimp. Encalypta contorta Lindb. Encalypta streptocarpa Hedw. Encalypta extinctoria Lindb. Encalypta vulgaris Hedw. Entodon schreberi (Brid.) Mönk. Pleurozium schreberi (Brid.) Mitt. Erythrophyllum rubellum Hilp. Bryoerythrophyllum recurvirostrum (Hedw.) P. C. Chen Eurhynchium praelongum auct. non (Hedw.) Eurhynchium hians (Hedw.) Sande Lac. Schimp. Eurhynchium praelongum var. abbreviatum Eurhynchium schleicheri (R. Hedw.) Jur. Turner Eurhynchium rusciforme Milde Platyhypnidium riparioides (Hedw.) Dixon Eurhynchium stokesii (Turner) Schimp. Eurhynchium praelongum (Hedw.) Schimp. Eurhynchium striatum subsp. zetterstedtii Eurhynchium angustirete (Broth.) T. J. Kop. (Størmer) Podp. Eurhynchium swartzii (Turner) Curn. Eurhynchium hians (Hedw.) Sande Lac. Eurhynchium tommasinii (Sendtn.) R. Ruthe Cirriphyllum tommasinii (Boulay) Grout Eurhynchium velutinoides Schimp. Eurhynchium flotowianum (Sendtn.) Kartt. Eurhynchium zetterstedtii Størmer Eurhynchium angustirete (Broth.) T. J. Kop. Fabronia octoblepharis (Schleich.) Schwägr. Fabronia ciliaris (Brid.) Brid. Fissidens crassipes var. mildeanus (Schimp.) Fissidens crassipes Bruch et Schimp. subsp. Mönk. crassipes

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SYNONYM ACCEPTED NAME Fissidens crassipes var. philibertii Besch. Fissidens crassipes Bruch et Schimp. subsp. warnstorfii (M. Fleisch.) Brugg.-Nann. Fissidens crassipes var. rufipes Schimp. Fissidens crassipes Bruch et Schimp. subsp. crassipes Fissidens crassipes var. submarginatus M. Fissidens crassipes Bruch et Schimp. subsp. Fleisch. et Warnst. warnstorfii (M. Fleisch.) Brugg.-Nann. Fissidens cristatus Wilson ex Mitt. Fissidens dubius P. Beauv. Fissidens decipiens De Not. Fissidens dubius P. Beauv. Fissidens minutulus Sull. Fissidens pusillus (Wilson) Milde Fissidens minutulus var. tenuifolius (Boulay) Fissidens gracilifolius Brugg.-Nann. et Nyholm Norkett Fissidens obtusifolius auct. non Wilson Fissidens arnoldii R. Ruthe Fissidens pusillus auct. pp. Fissidens pusillus (Wilson) Milde Fissidens pusillus auct. pp. Fissidens gracilifolius Brugg.-Nann. et Nyholm Fissidens pusillus var. minutulus (Sull.) Husn. Fissidens gracilifolius Brugg.-Nann. et Nyholm Fissidens tamarindifolius (Turner) Brid. Fissidens incurvus Starke ex Röhl. Funaria calcarea Wahlenb. Funaria muhlenbergii Turner Funaria dentata Crome Funaria muhlenbergii Turner Funaria fascicularis (Hedw.) Lindb. Entosthodon fascicularis (Hedw.) Müll. Hal. Funaria hibernica Hook. Funaria muhlenbergii Turner Funaria hungarica Boros Entosthodon hungaricus (Boros) Loeske Funaria mediterranea Lindb. Funaria muhlenbergii Turner Georgia pellucida (Hedw.) Rabenh. Tetraphis pellucida Hedw. Grimmia affinis Hornsch. Grimmia longirostris Hook. Grimmia apocarpa Hedw. Schistidium apocarpum (Hedw.) Bruch et Schimp. Grimmia campestris Hook. Grimmia laevigata (Brid.) Brid. Grimmia commutata Huebener Grimmia ovalis (Hedw.) Lindb. Grimmia flaccida (De Not.) Lindb. Schistidium flaccidum (De Not.) Ochyra Grimmia leucophaea Grev. Grimmia laevigata (Brid.) Brid. Grimmia trichophylla var. muehlenbeckii Grimmia muehlenbeckii Schimp. (Schimp.) Husn. Grimmia trichophylla var. tenuis (Wahlenb.) Grimmia muehlenbeckii Schimp. Wijk et Margad. Gymnostomum rupestre Schwägr. Gymnostomum aeruginosum Sm. Hedwigia albicans Lindb. p.p. Hedwigia ciliata (Hedw.) Ehrh. ex P. Beauv. Hedwigia albicans Lindb. p.p. Hedwigia stellata Hedenäs Hennediella heimii (Hedw.) R. H. Zander Desmatodon heimii (Hedw.) Mitt. Heterocladium squarrosulum Voit Heterocladium dimorphum (Brid.) Schimp. Heterocladium squarrulosum Lindb. Heterocladium dimorphum (Brid.) Schimp. Homalothecium geheebii (Milde) Wigh Brachythecium geheebii Milde Homalothecium nitens (Hedw.) H. Rob. Tomentypnum nitens (Hedw.) Loeske Homalothecium trichoides (Lindb.) Boros Tomentypnum nitens (Hedw.) Loeske Hygramblystegium filicinum (Hedw.) Loeske Cratoneuron filicinum (Hedw.) Spruce Hygramblystegium fluviatile (Hedw.) Loeske Amblystegium fluviatile (Hedw.) Schimp. Hygramblystegium irriguum Loeske Amblystegium tenax (Hedw.) C. E. O. Jensen Hygramblystegium tenax (Hedw.) Jenn. Amblystegium tenax (Hedw.) C. E. O. Jensen Hygrohypnum palustre Loeske Hygrohypnum luridum (Hedw.) Jenn.

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SYNONYM ACCEPTED NAME Hylocomium proliferum (Brid.) Lindb. Hylocomium splendens (Hedw.) Schimp. Hylocomium squarrosum (Hedw.) Schimp. Rhytidiadelphus squarrosus (Hedw.) Warnst. Hylocomium triquetrum (Hedw.) Schimp. Rhytidiadelphus triquetrus (Hedw.) Warnst. Hymenostomum microstomum (Hedw.) R. Weissia brachycarpa (Nees et Hornsch.) Jur. Braun ex Nees et Hornsch. Hymenostomum rostellatum (Brid.) Schimp. Weissia rostellata (Brid.) Lindb. Hymenostomum tortile (Schwägr.) Bruch et Weissia condensa (Voit) Lindb. Schimp. Hymenostylium curvirostre Mitt. Hymenostylium recurvirostrum (Hedw.) Dixon Hypnum arcuatum Lindb. Hypnum lindbergii Mitt. Hypnum incurvatum Brid. Homomallium incurvatum (Brid.) Loeske Hypnum reptile Michx. Hypnum pallescens (Hedw.) P. Beauv. Isopterygium elegans (Brid.) Lindb. Pseudotaxiphyllum elegans (Brid.) Z. Iwats. Isopterygium pulchellum (Hedw.) A. Jaeger et Isopterygiopsis pulchella (Hedw.) Z. Iwats. Sauerb. Isopterygium seligeri (Brid.) Dixon Herzogiella seligeri (Brid.) Z. Iwats. Isothecium filescens (Brid.) Mönk. Eurhynchium striatulum (Spruce) Schimp. Isothecium myurum Brid. Isothecium alopecuroides (Dubois) Isov. Isothecium viviparum Lindb. Isothecium alopecuroides (Dubois) Isov. Leptobryum lutescens (Limpr.) Mönk. Pohlia lutescens (Limpr.) Lindb. Leptodictyum kochii (Schimp.) Warnst. Amblystegium humile (P. Beauv.) Crundw. Leptodictyum trichopodium (Schultz) Warnst. Amblystegium humile (P. Beauv.) Crundw. Lescuraea saviana (De Not.) E. Lawton Pseudoleskea saviana (De Not.) Latzel Leskea catenulata (Schrad.) Mitt. Pseudoleskeella catenulata (Schrad.) Kindb. Leskea nervosa (Brid.) Myrin Pseudoleskeella nervosa (Brid.) Nyholm Leskeella nervosa (Brid.) Loeske Pseudoleskeella nervosa (Brid.) Nyholm Microbryum curvicolle (Hedw.) R. H. Zander Phascum curvicolle Hedw. Microbryum davallianum (Sm.) R. H. Zander Pottia davalliana (Sm.) C. E. O. Jensen Microbryum floerkeanum (F. Weber et D. Phascum floerkeanum F. Weber et D. Mohr Mohr) Schimp. Microbryum starckeanum R. H. Zander Pottia starckeana (Hedw.) Müll. Hal. Microbryum starckeanum var. brachyodus Pottia mutica Venturi (Bruch et Schimp.) R. H. Zander Mniobryum albicans (Wahlenb.) Limpr. Pohlia wahlenbergii (F. Weber et D. Mohr) A. L. Andrews Mniobryum carneum (F. Weber et D. Mohr) Pohlia melanodon (Brid.) A. J. Shaw Limpr. Mniobryum delicatulum (Hedw.) Dixon Pohlia melanodon (Brid.) A. J. Shaw Mniobryum wahlenbergii (F. Weber et D. Pohlia wahlenbergii (F. Weber et D. Mohr) A. L. Mohr) Jenn. Andrews Mnium affine Blandow Plagiomnium affine (Blandow) T. J. Kop. Mnium affine var. elatum Bruch et Schimp. Plagiomnium elatum (Bruch et Schimp.) T. J. Kop. Mnium ambiguum H. Müll. Mnium lycopodioides Schwägr. Mnium cuspidatum auct. p.p. Plagiomnium affine (Blandow) T. J. Kop. Mnium cuspidatum Hedw. Plagiomnium cuspidatum (Hedw.) T. J. Kop. Mnium longirostre Brid. Plagiomnium rostratum (anon.) T. J. Kop.

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SYNONYM ACCEPTED NAME Mnium medium Bruch et Schimp. Plagiomnium medium (Bruch et Schimp.) T. J. Kop. Mnium orthorrhynchum auct. non Brid. Mnium thomsonii Schimp. Mnium orthorrhynchum var. lycopodioides Mnium lycopodioides Schwägr. (Schwägr.) Husn. Mnium punctatum Hedw. Rhizomnium punctatum (Hedw.) T. J. Kop. Mnium rostratum anon. Plagiomnium rostratum (anon.) T. J. Kop. Mnium rugicum Laurer Plagiomnium ellipticum (Brid.) T. J. Kop. Mnium seligeri auct. Plagiomnium elatum (Bruch et Schimp.) T. J. Kop. Mnium undulatum Hedw. Plagiomnium undulatum (Hedw.) T. J. Kop. Neckera besseri (Lobarz.) Jur. Homalia besseri Lobarz. Neckera sendtneriana Schimp. Homalia besseri Lobarz. Orthodicranum flagellare (Hedw.) Loeske Dicranum flagellare Hedw. Orthodicranum montanum (Hedw.) Loeske Dicranum montanum Hedw. Orthodicranum strictum (D. Mohr) Culm. Dicranum tauricum Sapjegin Orthodicranum tauricum (Sapjegin) Z. Dicranum tauricum Sapjegin Smirnova Orthotrichum braunii Bruch et Schimp. Orthotrichum stellatum Brid. Orthotrichum fallax Bruch ex Brid. Orthotrichum pumilum Sw. Orthotrichum fastigiatum Brid. Orthotrichum affine Brid. Orthotrichum leiocarpum Bruch et Schimp. Orthotrichum striatum Hedw. Orthotrichum leucomitrium Bruch et Schimp. Orthotrichum scanicum Grönv. Orthotrichum octoblepharum Brid. Orthotrichum affine Brid. Orthotrichum sardagnanum Venturi Orthotrichum cupulatum Brid. Orthotrichum saxatile Brid. Orthotrichum anomalum Hedw. Orthotrichum schimperi Hammar Orthotrichum pumilum Sw. Orthotrichum sturmii Hoppe et Hornsch. Orthotrichum rupestre Schleich. ex Schwägr. Oxyrhynchium hians (Hedw.) Loeske Eurhynchium hians (Hedw.) Sande Lac. Oxyrhynchium praelongum var. stokesii Eurhynchium praelongum (Hedw.) Schimp. (Turner) Podp. Oxyrhynchium pumilum (Wilson) Loeske Eurhynchium pumilum (Wilson) Schimp. Oxyrhynchium schleicheri (R. Hedw.) Roell Eurhynchium schleicheri (R. Hedw.) Jur. Oxyrhynchium speciosum (Brid.) Warnst. Eurhynchium speciosum (Brid.) Jur. Oxyrhynchium swartzii (Turner) Warnst. Eurhynchium hians (Hedw.) Sande Lac. Oxystegus cylindricus (Brid.) Hilp. Oxystegus tenuirostris (Hook. et Taylor) A. J. E. Sm. Oxystegus sinuosus Hilp. Didymodon sinuosus (Mitt.) Delogne Phascum acaulon L. ex With. Phascum cuspidatum Hedw. Phascum alternifolium Hedw. Archidium alternifolium (Hedw.) Schimp. Phascum halophilum Šmarda Phascum cuspidatum Hedw. Philonotis capillaris auct. Philonotis arnelli Husn. Physcomitrella patens (Hedw.) Schimp. Aphanorrhegma patens (Hedw.) Lindb. Physcomitrium acuminatum Bruch et Schimp. Physcomitrium eurystomum Sendtn. Plagiopus oederi (Brid.) Limpr. Plagiopus oederianus (Sw.) H. A. Crum et L. E. Anderson Plagiothecium curvifolium Limpr. Plagiothecium laetum Schimp. Plagiothecium densifolium (Broth.) Limpr. Taxiphyllum densifolium (Broth.) Reimers Plagiothecium elegans (Brid.) Sull. Pseudotaxiphyllum elegans (Brid.) Z. Iwats.

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SYNONYM ACCEPTED NAME Plagiothecium neglectum Mönk. Plagiothecium nemorale (Mitt.) A. Jaeger Plagiothecium pulchellum (Hedw.) Schimp. Isopterygiopsis pulchella (Hedw.) Z. Iwats. Plagiothecium roeseanum Schimp. Plagiothecium cavifolium (Brid.) Z. Iwats. Plagiothecium silesiacum (F. Weber et D. Herzogiella seligeri (Brid.) Z. Iwats. Mohr) Schimp. Plagiothecium sylvaticum auct. Plagiothecium nemorale (Mitt.) A. Jaeger Plasteurhynchium striatulum (Spruce) M. Eurhynchium striatulum (Spruce) Schimp. Fleisch. Platydictya confervoides (Brid.) H. A. Crum Amblystegium confervoides (Brid.) Schimp. Platydictya subtilis (Hedw.) H. A. Crum Amblystegium subtile (Hedw.) Schimp. Pleuridium alternifolium auct. non (Hedw.) Pleuridium subulatum (Hedw.) Rabenh. Brid. Pleuridium subulatum auct. non (Hedw.) Pleuridium acuminatum Lindb. Rabenh. Pohlia carnea Lindb. Pohlia melanodon (Brid.) A. J. Shaw Pohlia grandiflora H. Lindb. Pohlia annotina (Hedw.) Lindb. Polytrichastrum alpinum (Hedw.) G. L. Sm. Polytrichum alpinum Hedw. Polytrichastrum formosum (Hedw.) G. L. Sm. Polytrichum formosum Hedw. Polytrichastrum longisetum (Brid.) G. L. Sm. Polytrichum longisetum Sw. ex Brid. Polytrichum alpestre Hoppe Polytrichum strictum Brid. Polytrichum angustatum Brid. Atrichum angustatum (Brid.) Bruch et Schimp. Polytrichum attenuatum Menzies ex Brid. Polytrichum formosum Hedw. Polytrichum gracile Dicks. Polytrichum longisetum Sw. ex Brid. Pottia heimii (Hedw.) Hampe Desmatodon heimii (Hedw.) Mitten Pottia truncatula (With.) Buse Pottia truncata (Hedw.) Bruch et Schimp. Pseudephemerum axillare (Lindb.) I. Hagen Pseudephemerum nitidum (Hedw.) Reimers Pseudocalliergon lycopodioides (Brid.) Drepanocladus lycopodioides (Brid.) Warnst. Hedenäs Pseudoisothecium myosuroides (Brid.) Grout Isothecium myosuroides Brid. Pseudoleskea illyrica Glowacki Pseudoleskea saviana (De Not.) Latzel Pseudoscleropodium purum (Hedw.) M. Scleropodium purum (Hedw.) Limpr. Fleisch. ex Broth. Pterogonium ornithopodioides (F. Weber et D. Pterogonium gracile (Hedw.) Sm. Mohr) Lindb. Pterygoneurum cavifolium Jur. Pterygoneurum ovatum (Hedw.) Dixon Pterygoneurum pusillum (Lindb.) C. E. O. Pterygoneurum ovatum (Hedw.) Dixon Jensen Racomitrium protensum (A. Braun) Huebener Racomitrium aquaticum (Schrad.) Brid. Rhabdoweisia schisti (F. Weber et D. Mohr) Cnestrum schisti (F. Weber et D. Mohr) I. Hagen Bruch et Schimp. Rhabdoweisia striata (Schrad.) Lindb. Rhabdoweisia fugax (Hedw.) Bruch et Schimp. Rhodobryum spathulatum auct. Rhodobryum ontariense (Kindb.) Kindb. Rhynchostegiella algiriana (P. Beauv. ) Rhynchostegiella tenella (Dicks.) Limpr. Warnst. Rhynchostegiella compacta (Müll. Hal.) Conardia compacta (Müll. Hal.) H. Rob. Loeske

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SYNONYM ACCEPTED NAME Rhynchostegiella jacquinii (Garov.) Limpr. Rhynchostegiella teneriffae (Mont.) Dirkse et Bouman Rhynchostegiella pallidirostra (Brid.) Loeske Eurhynchium pumilum (Wilson) Schimp. Rhynchostegium riparioides (Hedw.) Cardot Platyhypnidium riparioides (Hedw.) Dixon Saelania caesia (P. Beauv.) Lindb. Saelania glaucescens (Hedw.) Broth. Schistidium apocarpum var. brunnescens Schistidium brunnescens Limpr. (Limpr.) Loeske Schistidium pulvinatum auct. non (Hedw.) Schistidium flaccidum (De Not.) Ochyra Brid. Schistidium pulvinatum var. flaccidum (De Schistidium flaccidum (De Not.) Ochyra Not.) De Not. Scorpidium cossonii (Schimp.) Hedenäs Drepanocladus cossonii (Schimp.) Loeske Seligeria setacea Lindb. Seligeria recurvata (Hedw.) Bruch et Schimp. Sharpiella seligeri (Brid.) Z. Iwats. Herzogiella seligeri (Brid.) Z. Iwats. Sphagnum acutifolium Ehrh. ex Schrad. Sphagnum capillifolium (Ehrh.) Hedw. Sphagnum nemoreum auct. Sphagnum capillifolium (Ehrh.) Hedw. Sphagnum parvifolium (Warnst.) Warnst. Sphagnum angustifolium (Warnst.) C. E. O. Jensen Sphagnum plumulosum Roell Sphagnum subnitens Russow et Warnst. Sphagnum recurvum P. Beauv. var. Sphagnum flexuosum Dozy et Molk. amblyphyllum (Russow) Warnst. Sphagnum recurvum P. Beauv. var. Sphagnum fallax (H. Klinggräff) H. Klinggräff mucronatum (Russow) Warnst. Sphagnum robustum (Warnst.) Roell Sphagnum russowii Warnst. Sphagnum rufescens (Nees et Hornsch.) Sphagnum denticulatum var. denticulatum Brid. Warnst.) Sphagnum subbicolor auct. non Hampe Sphagnum centrale C. E. O. Jensen Splachnobryum wrightii Müll. Hal. Splachnobryum obtusum (Brid.) Müll. Hal. Steppomitra hungarica (Boros) Vondraček et Entosthodon hungaricus (Boros) Loeske Hadač Straminergon stramineum (Brid.) Hedenäs Calliergon stramineum (Brid.) Kindb. Streblotrichum convolutum (Hedw.) P. Beauv. Barbula convoluta Hedw. Swartzia montana Lindb. Distichium capillaceum (Hedw.) Bruch et Schimp. Syntrichia calcicola J. J. Amann Tortula calcicolens W. A. Kramer Syntrichia inermis (Brid.) Bruch Tortula inermis (Brid.) Mont. Syntrichia intermedia Brid. Tortula crinita (De Not.) De Not. Syntrichia laevipila Brid. Tortula laevipila (Brid.) Schwägr. Syntrichia latifolia (Hartm.) Huebener Tortula latifolia Bruch ex Hartm. Syntrichia montana Nees Tortula crinita (De Not.) De Not. Syntrichia mucronifolia (Schwägr.) Brid. Tortula mucronifolia Schwägr. Syntrichia norvegica F. Weber Tortula norvegica (F. Weber) Lindb. Syntrichia papillosa (Wilson) Jur. Tortula papillosa Wilson Syntrichia princeps (De Not.) Mitt. Tortula princeps De Not. Syntrichia ruralis (Hedw.) F. Weber et D. Tortula ruralis (Hedw.) P. Gaertn., B. Mey. et Mohr Scherb. Syntrichia ruralis var. arenicola (Braithw.) Tortula ruraliformis (Besch.) Ingham Amann Syntrichia ruralis var. hirsuta (Venturi) Podp. Tortula papillosissima (Copp.) Broth.

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SYNONYM ACCEPTED NAME Syntrichia ruralis var. ruraliformis (Besch.) Tortula ruraliformis (Besch.) Ingham Durand Syntrichia ruralis var. submamillosa (W. A. Tortula papillosissima (Copp.) Broth. Kramer) R. H. Zander Syntrichia subulata (Hedw.) F. Weber et D. Tortula subulata Hedw. Mohr Syntrichia virescens (De Not.) Ochyra Tortula virescens (De Not.) De Not. Taxiphyllum depressum (Brid.) Reimers Taxiphyllum wissgrillii (Garov.) Wijk et Margad. Thamnium alopecurum (Hedw.) Schimp. Thamnobryum alopecurum (Hedw.) Gangulee Thuidium erectum Duby Thuidium delicatulum (Hedw.) Schimp. Thuidium lanatum (Brid.) I. Hagen Helodium blandowii (F. Weber et D. Mohr) Warnst. Thuidium tamariscifolium Lindb. Thuidium tamariscinum (Hedw.) Schimp. Tortella cylindrica (Brid.) Loeske Oxystegus tenuirostris (Hook. et Taylor) A. J. E. Sm. Tortula acaulon (With.) R. H. Zander Phascum cuspidatum Hedw. Tortula caninervis subsp. spuria var. Tortula caninervis (Mitt.) Broth. subs. spuria (J. gypsophila (Roth) W. A. Kramer J. Amann) W. A. Kramer Tortula cernua (Huebener) Lindb. Desmatodon cernuus (Huebener) Bruch et Schimp. Tortula desertorum Broth. Tortula caninervis (Mitt.) Broth. subs. spuria (J. J. Amann) W. A. Kramer Tortula intermedia (Brid.) Berk. Tortula crinita (De Not.) De Not. Tortula lanceola R. H. Zander Pottia lanceolata (Hedw.) Müll. Hal. Tortula modica R. H. Zander Pottia intermedia (Turner) Fürnr. Tortula muralis var. obtusifolia (Schwaegr.) Tortula obtusifolia (Schwägr.) Mathieu Boros Tortula protobryoides R. H. Zander Pottia bryoides (Dicks.) Mitt. Tortula rhizophylla (Sakurai) Z. Iwats. et K. Phascum leptophyllum Müll. Hal. Saito Tortula ruralis var. arenicola Braithw. Tortula ruraliformis (Besch.) Ingham Tortula ruralis var. calcicola (J. J. Amann) Tortula calcicolens W. A. Kramer Barkman Tortula truncata (Hedw.) Mitt. Pottia truncata (Hedw.) Bruch et Schimp. Tortula velenovskyi Schiffn. Hilpertia velenovskyi (Schiffn.) R. H. Zander Trichodon cylindricum (Hedw.) Schimp. Ditrichum cylindricum (Hedw.) Grout Trichostomum cylindricum (Brid.) Müll. Hal. Oxystegus tenuirostris (Hook. et Taylor) A. J. E. Sm. Trichostomum tenuirostre (Hook. et Taylor) Oxystegus tenuirostris (Hook. et Taylor) A. J. E. Lindb. Sm. Ulota americana (P. Beauv.) Limpr. Ulota hutchinsiae (Sm.) Hammar Ulota crispa var. norvegica (Grönv.) A. J. E. Ulota bruchii Brid. Sm. et M. O. Hill Ulota crispula Brid. Ulota crispa (Hedw.) Brid. Ulota ludwigii (Brid.) Brid. Ulota coarctata (P. Beauv.) Hammar Ulota ulophylla Broth. Ulota crispa (Hedw.) Brid.

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SYNONYM ACCEPTED NAME Weissia controversa var. crispata (Nees et Weissia fallax Sehlm. Hornsch.) Nyholm Weissia microstoma (Hedw.) Müll. Hal. Weissia brachycarpa (Nees et Hornsch.) Jur. Weissia tortilis (Schwägr.) Müll. Hal. Weissia condensa (Voit) Lindb. Weissia tortilis var. fallax (Sehlm.) Mönk. Weissia fallax Sehlm. Weissia viridula Brid. Weissia controversa Hedw. Zygodon viridissimus var. vulgaris Malta Zygodon rupestris Lorentz

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