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Die Microsauria Des Mitteleuropäischen Rotliegend
Die Microsauria des mitteleuropäischen Rotliegend Dissertation zur Erlangung des Grades „Doktor der Naturwissenschaften" im Promotionsfach Geologie/Paläontologie am Fachbereich Chemie, Pharmazie und Geowissenschaften der Johannes Gutenberg-Universität in Mainz Sabine Glienke geb. in Worms Mainz, 2011 http://d-nb.info/1058187503 Inhalt Inhalt 1. Einleitung 6 1.1. Allgemeine Merkmale und Bearbeitungsgeschichte 6 1.2. Fundorte und Erhaltung 9 2. Methoden, Abkürzungen und Material 16 2.1. Methoden 16 2.1.1. Bearbeitung der Skelette 16 2.1.2. Gewinnung und Bearbeitung der Einzelknochen 16 2.1.3. Kladogramme 17 2.2. Abkürzungen 17 2.2.1. Sammlungen 17 2.2.2. In den Zeichnungen verwendete Abkürzungen 17 2.3. Übersicht über die untersuchten Skelette 19 3. Beschreibungen 21 3.1. Die Familie Brachystelechidae CARROLL& GASKILL, 1978 21 3.1.1. Systematische Stellung 21 3.1.2. Diagnose 21 3.2. Die Gattung Batropetes CARROLL & GASKILL, 1971 21 3.2.1. Systematische Stellung 21 3.2.2. Diagnose 22 3.2.3. Die vier Spezies der Gattung Batropetes 22 3.3. Batropetes niederkirchensis n. sp 26 3.3.1. Diagnose 26 3.3.2. Beschreibung 28 3.3.2.1. Schädel 28 3.3.2.1.1. Schädeldach 28 3.3.2.1.2. Gaumen 38 3.3.2.1.3. Hirnkapsel 43 3.3.2.1.4. Unterkiefer 46 3.3.2.2. Postcraniales Skelett 48 Inhalt 3.4. Batropetes palatinus n. sp 62 3.4.1. Diagnose 62 3.4.2. Beschreibung 63 3.4.2.1. Schädel 74 3.4.2.1.1. Schädeldach 74 3.4.2.1.2. -
Reptile Family Tree
Reptile Family Tree - Peters 2015 Distribution of Scales, Scutes, Hair and Feathers Fish scales 100 Ichthyostega Eldeceeon 1990.7.1 Pederpes 91 Eldeceeon holotype Gephyrostegus watsoni Eryops 67 Solenodonsaurus 87 Proterogyrinus 85 100 Chroniosaurus Eoherpeton 94 72 Chroniosaurus PIN3585/124 98 Seymouria Chroniosuchus Kotlassia 58 94 Westlothiana Casineria Utegenia 84 Brouffia 95 78 Amphibamus 71 93 77 Coelostegus Cacops Paleothyris Adelospondylus 91 78 82 99 Hylonomus 100 Brachydectes Protorothyris MCZ1532 Eocaecilia 95 91 Protorothyris CM 8617 77 95 Doleserpeton 98 Gerobatrachus Protorothyris MCZ 2149 Rana 86 52 Microbrachis 92 Elliotsmithia Pantylus 93 Apsisaurus 83 92 Anthracodromeus 84 85 Aerosaurus 95 85 Utaherpeton 82 Varanodon 95 Tuditanus 91 98 61 90 Eoserpeton Varanops Diplocaulus Varanosaurus FMNH PR 1760 88 100 Sauropleura Varanosaurus BSPHM 1901 XV20 78 Ptyonius 98 89 Archaeothyris Scincosaurus 77 84 Ophiacodon 95 Micraroter 79 98 Batropetes Rhynchonkos Cutleria 59 Nikkasaurus 95 54 Biarmosuchus Silvanerpeton 72 Titanophoneus Gephyrostegeus bohemicus 96 Procynosuchus 68 100 Megazostrodon Mammal 88 Homo sapiens 100 66 Stenocybus hair 91 94 IVPP V18117 69 Galechirus 69 97 62 Suminia Niaftasuchus 65 Microurania 98 Urumqia 91 Bruktererpeton 65 IVPP V 18120 85 Venjukovia 98 100 Thuringothyris MNG 7729 Thuringothyris MNG 10183 100 Eodicynodon Dicynodon 91 Cephalerpeton 54 Reiszorhinus Haptodus 62 Concordia KUVP 8702a 95 59 Ianthasaurus 87 87 Concordia KUVP 96/95 85 Edaphosaurus Romeria primus 87 Glaucosaurus Romeria texana Secodontosaurus -
Morphology, Phylogeny, and Evolution of Diadectidae (Cotylosauria: Diadectomorpha)
Morphology, Phylogeny, and Evolution of Diadectidae (Cotylosauria: Diadectomorpha) by Richard Kissel A thesis submitted in conformity with the requirements for the degree of doctor of philosophy Graduate Department of Ecology & Evolutionary Biology University of Toronto © Copyright by Richard Kissel 2010 Morphology, Phylogeny, and Evolution of Diadectidae (Cotylosauria: Diadectomorpha) Richard Kissel Doctor of Philosophy Graduate Department of Ecology & Evolutionary Biology University of Toronto 2010 Abstract Based on dental, cranial, and postcranial anatomy, members of the Permo-Carboniferous clade Diadectidae are generally regarded as the earliest tetrapods capable of processing high-fiber plant material; presented here is a review of diadectid morphology, phylogeny, taxonomy, and paleozoogeography. Phylogenetic analyses support the monophyly of Diadectidae within Diadectomorpha, the sister-group to Amniota, with Limnoscelis as the sister-taxon to Tseajaia + Diadectidae. Analysis of diadectid interrelationships of all known taxa for which adequate specimens and information are known—the first of its kind conducted—positions Ambedus pusillus as the sister-taxon to all other forms, with Diadectes sanmiguelensis, Orobates pabsti, Desmatodon hesperis, Diadectes absitus, and (Diadectes sideropelicus + Diadectes tenuitectes + Diasparactus zenos) representing progressively more derived taxa in a series of nested clades. In light of these results, it is recommended herein that the species Diadectes sanmiguelensis be referred to the new genus -
71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas. -
Early Tetrapod Relationships Revisited
Biol. Rev. (2003), 78, pp. 251–345. f Cambridge Philosophical Society 251 DOI: 10.1017/S1464793102006103 Printed in the United Kingdom Early tetrapod relationships revisited MARCELLO RUTA1*, MICHAEL I. COATES1 and DONALD L. J. QUICKE2 1 The Department of Organismal Biology and Anatomy, The University of Chicago, 1027 East 57th Street, Chicago, IL 60637-1508, USA ([email protected]; [email protected]) 2 Department of Biology, Imperial College at Silwood Park, Ascot, Berkshire SL57PY, UK and Department of Entomology, The Natural History Museum, Cromwell Road, London SW75BD, UK ([email protected]) (Received 29 November 2001; revised 28 August 2002; accepted 2 September 2002) ABSTRACT In an attempt to investigate differences between the most widely discussed hypotheses of early tetrapod relation- ships, we assembled a new data matrix including 90 taxa coded for 319 cranial and postcranial characters. We have incorporated, where possible, original observations of numerous taxa spread throughout the major tetrapod clades. A stem-based (total-group) definition of Tetrapoda is preferred over apomorphy- and node-based (crown-group) definitions. This definition is operational, since it is based on a formal character analysis. A PAUP* search using a recently implemented version of the parsimony ratchet method yields 64 shortest trees. Differ- ences between these trees concern: (1) the internal relationships of aı¨stopods, the three selected species of which form a trichotomy; (2) the internal relationships of embolomeres, with Archeria -
Physical and Environmental Drivers of Paleozoic Tetrapod Dispersal Across Pangaea
ARTICLE https://doi.org/10.1038/s41467-018-07623-x OPEN Physical and environmental drivers of Paleozoic tetrapod dispersal across Pangaea Neil Brocklehurst1,2, Emma M. Dunne3, Daniel D. Cashmore3 &Jӧrg Frӧbisch2,4 The Carboniferous and Permian were crucial intervals in the establishment of terrestrial ecosystems, which occurred alongside substantial environmental and climate changes throughout the globe, as well as the final assembly of the supercontinent of Pangaea. The fl 1234567890():,; in uence of these changes on tetrapod biogeography is highly contentious, with some authors suggesting a cosmopolitan fauna resulting from a lack of barriers, and some iden- tifying provincialism. Here we carry out a detailed historical biogeographic analysis of late Paleozoic tetrapods to study the patterns of dispersal and vicariance. A likelihood-based approach to infer ancestral areas is combined with stochastic mapping to assess rates of vicariance and dispersal. Both the late Carboniferous and the end-Guadalupian are char- acterised by a decrease in dispersal and a vicariance peak in amniotes and amphibians. The first of these shifts is attributed to orogenic activity, the second to increasing climate heterogeneity. 1 Department of Earth Sciences, University of Oxford, South Parks Road, Oxford OX1 3AN, UK. 2 Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstraße 43, 10115 Berlin, Germany. 3 School of Geography, Earth and Environmental Sciences, University of Birmingham, Birmingham B15 2TT, UK. 4 Institut -
From the Late Permian of Eastern Europe V
Paleontological Journal, Vol. 32, No. 3, 1998, pp. 278–287. Translated from Paleontologicheskii Zhurnal, No. 3, 1998, pp. 64–73. Original Russian Text Copyright © 1998 by Golubev. English Translation Copyright © 1998 by åÄàä ç‡Û͇ /Interperiodica Publishing (Russia). Narrow-armored Chroniosuchians (Amphibia, Anthracosauromorpha) from the Late Permian of Eastern Europe V. K. Golubev Paleontological Institute, Russian Academy of Sciences, ul. Profsoyuznaya 123, Moscow, 117647 Russia Received January 21, 1997 Abstract—The Permian and Triassic chroniosuchians are revised and the morphology of the dorsal armor scutes is discussed in detail. The first narrow-armored chroniosuchid Uralerpeton tverdochlebovae from the Late Permian Vyazniki faunistic assemblage of Eastern Europe is described and the age of the Vyazniki fauna discussed. INTRODUCTION armor occurred late in the process of decomposition of the animal: the head, the brachial and pelvic girdles and even The chroniosuchians (Chroniosuchia), an unusual the vertebral column became disarticulated sooner. group of Late Permian and Triassic reptiliomorph amphibians, dominated the aquatic tetrapod assem- In horizontal plane the chroniosuchian scutes are blages of Eastern Europe during the Late Tatarian age. rectangular (Figs. 1a and 1b; 2b and 2c; 4a–4d), except Conventionally this group, consisting of two families: for the anteriormost scute, which is shaped like a semi- the Chroniosuchidae and the Bystrowianidae (Tatar- circle or semiellipsis (Fig. 1d). The scute consists of a inov, 1972; Ivakhnenko and Tverdokhlebova, 1980; massive axial part, or the scute body (corpus scutulumi) Shishkin and Novikov, 1992) is included within the and two lateral horizontal plates, or the scute wings order Anthracosauromorpha as suborder. According to (alae scutulumi) (Fig. -
The Devonian Tetrapod Acanthostega Gunnari Jarvik: Postcranial Anatomy, Basal Tetrapod Interrelationships and Patterns of Skeletal Evolution M
Transactions of the Royal Society of Edinburgh: Earth Sciences, 87, 363-421, 1996 The Devonian tetrapod Acanthostega gunnari Jarvik: postcranial anatomy, basal tetrapod interrelationships and patterns of skeletal evolution M. I. Coates ABSTRACT: The postcranial skeleton of Acanthostega gunnari from the Famennian of East Greenland displays a unique, transitional, mixture of features conventionally associated with fish- and tetrapod-like morphologies. The rhachitomous vertebral column has a primitive, barely differentiated atlas-axis complex, encloses an unconstricted notochordal canal, and the weakly ossified neural arches have poorly developed zygapophyses. More derived axial skeletal features include caudal vertebral proliferation and, transiently, neural radials supporting unbranched and unsegmented lepidotrichia. Sacral and post-sacral ribs reiterate uncinate cervical and anterior thoracic rib morphologies: a simple distal flange supplies a broad surface for iliac attachment. The octodactylous forelimb and hindlimb each articulate with an unsutured, foraminate endoskeletal girdle. A broad-bladed femoral shaft with extreme anterior torsion and associated flattened epipodials indicates a paddle-like hindlimb function. Phylogenetic analysis places Acanthostega as the sister- group of Ichthyostega plus all more advanced tetrapods. Tulerpeton appears to be a basal stem- amniote plesion, tying the amphibian-amniote split to the uppermost Devonian. Caerorhachis may represent a more derived stem-amniote plesion. Postcranial evolutionary trends spanning the taxa traditionally associated with the fish-tetrapod transition are discussed in detail. Comparison between axial skeletons of primitive tetrapods suggests that plesiomorphic fish-like morphologies were re-patterned in a cranio-caudal direction with the emergence of tetrapod vertebral regionalisation. The evolution of digited limbs lags behind the initial enlargement of endoskeletal girdles, whereas digit evolution precedes the elaboration of complex carpal and tarsal articulations. -
Lopingian, Permian) of North China
Foss. Rec., 23, 205–213, 2020 https://doi.org/10.5194/fr-23-205-2020 © Author(s) 2020. This work is distributed under the Creative Commons Attribution 4.0 License. The youngest occurrence of embolomeres (Tetrapoda: Anthracosauria) from the Sunjiagou Formation (Lopingian, Permian) of North China Jianye Chen1 and Jun Liu1,2,3 1Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China 2Chinese Academy of Sciences Center for Excellence in Life and Paleoenvironment, Beijing 100044, China 3College of Earth and Planetary Sciences, University of Chinese Academy of Sciences, Beijing 100049, China Correspondence: Jianye Chen ([email protected]) Received: 7 August 2020 – Revised: 2 November 2020 – Accepted: 16 November 2020 – Published: 1 December 2020 Abstract. Embolomeri were semiaquatic predators preva- 1 Introduction lent in the Carboniferous, with only two species from the early Permian (Cisuralian). A new embolomere, Seroher- Embolomeri are a monophyletic group of large crocodile- peton yangquanensis gen. et sp. nov. (Zoobank Registration like, semiaquatic predators, prevalent in the Carboniferous number: urn:lsid:zoobank.org:act:790BEB94-C2CC-4EA4- and early Permian (Cisuralian) (Panchen, 1970; Smithson, BE96-2A1BC4AED748, registration: 23 November 2020), is 2000; Carroll, 2009; Clack, 2012). The clade is generally named based on a partial right upper jaw and palate from the considered to be a stem member of the Reptiliomorpha, taxa Sunjiagou Formation of Yangquan, Shanxi, China, and is late that are more closely related to amniotes than to lissamphib- Wuchiapingian (late Permian) in age. It is the youngest em- ians (Ruta et al., 2003; Vallin and Laurin, 2004; Ruta and bolomere known to date and the only embolomere reported Coates, 2007; Clack and Klembara, 2009; Klembara et al., from North China Block. -
Karpinskiosaurus Ultimus (Seymouriamorpha, Parareptilia) from the Upper Permian of European Russia V
Paleontological Journal, Vol. 36, No. 1, 2002, pp. 72–79. Translated from Paleontologicheskii Zhurnal, No. 1, 2002, pp. 77–84. Original Russian Text Copyright © 2002 by Bulanov. English Translation Copyright © 2002 by åÄIä “Nauka /Interperiodica” (Russia). Karpinskiosaurus ultimus (Seymouriamorpha, Parareptilia) from the Upper Permian of European Russia V. V. Bulanov Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya ul. 123, Moscow, 117997 Russia e-mail: [email protected] Received April 14, 1999 Abstract—The genus Raphanodon Ivachnenko, 1987 (Leptorophidae, Seymouriamorpha) is not a valid taxon, since its type species Nycteroleter ultimus Tchudinov et Vjuschkov, 1956 is assigned to the genus Karpinskio- saurus. The cranial anatomy of K. ultimus is described in the present paper with newly collected material from the Babintsevo locality. The genus Karpinskiosaurus is assigned to the subfamily Karpinskiosaurinae, which is affiliated to the Lower Permian Discosauriscinae (Discosauriscus and Ariekanerpeton) to form the family Karpinskiosauridae. INTRODUCTION the skull structure of this form will be published in the future. In 1956, Tchudinov and Vjuschkov described a new species of the genus Nycteroleter, N. ultimus based on The genus Karpinskiosaurus was established by jaw material from the Pron’kino locality (Orenburg Sushkin (1925) based on two specimens (an incomplete Region) (Tchudinov and Vjuschkov, 1956). Later, Iva- skeleton and a skull) described by Amalitsky (1921) as khnenko separated this form from the nycteroleters and Kotlassia secunda from the excavations at the Malaya placed it in the family Leptorophidae (Seymouriamor- Severnaya Dvina River (Sokolki locality). Sushkin pha, Parareptilia) under a new name, Raphanodon. This ranked this genus as a new family, Karpinskiosauridae, paper included reconstructions of the skull roof and based on its distinctions from Kotlassia proper (Kotlas- palatal complex of R. -
Mapping, Stratigraphy, and Tectonic Implications of Lowerrermian
...MAPPING, STRATIGRAPHY, AND TECTONIC IMPLICATIONS OF LOWERRERMIAN STRATA, EASTERN ~ICHITA MOUNTAINS, OKLAHOMA By STEVE DWAYNE BRIDGES \\ Bachelor of Science Arkansas Tech University Russellville, Arkansas 1982 Submitted to the Faculty of the Graduate College of the Oklahoma State University in partial fulfillment of the requirements for the Degree of MASTER OF SCIENCE December, 1985 . ~ T h.ts\.s \tl r 5' ~g S)m car.?- ; . t: .. ' MAPPING, STRATIGRAPHY, AND TECTONIC IMPLICATIONS OF LOWER PERMI-AN STRATA, EASTERN WICHITA MOUNTAINS, OKLAHOMA Thesis Approved: Thesis Adviser ,/JrdL'VV\ Ce~ ean of the Graduate College ii 1236335 PREFACE The purpose of this thesis is to map, propose a revised stratigraphic section for, and comment on the tectonic implications of the Lower Permian deposits surrounding the eastern Wichita Mountains of southwestern Oklahoma. It is hoped that this thesis will raise questions and encourage investigators to scrutinize the geologic literature so that our geologic interpretations can continue to evolve. I am deeply indebted to the many individuals that provided useful comments, ideas, and suggestions which helped in the completion of this study. Deepest appreciations are conveyed to my major advisor Dr. R. Nowell Donovan for his counsel, guidance and friendship and to my committee members Dr. Gary F. Stewart and Dr. Ibrahim Cemen for their suggestions and review of the text. Sincere thanks are expressed to Deborah Ragland who ordered photos and provided interpretations which were vital in the preparation of this thesis. Gratitude is expressed to Mr. and Mrs. Charlie Oliver for their hospitality, friendship and for allowing me the "run" of the Kimbell Ranch. -
Bones, Molecules, and Crown- Tetrapod Origins
TTEC11 05/06/2003 11:47 AM Page 224 Chapter 11 Bones, molecules, and crown- tetrapod origins Marcello Ruta and Michael I. Coates ABSTRACT The timing of major events in the evolutionary history of early tetrapods is discussed in the light of a new cladistic analysis. The phylogenetic implications of this are com- pared with those of the most widely discussed, recent hypotheses of basal tetrapod interrelationships. Regardless of the sequence of cladogenetic events and positions of various Early Carboniferous taxa, these fossil-based analyses imply that the tetrapod crown-group had originated by the mid- to late Viséan. However, such estimates of the lissamphibian–amniote divergence fall short of the date implied by molecular studies. Uneven rates of molecular substitutions might be held responsible for the mismatch between molecular and morphological approaches, but the patchy quality of the fossil record also plays an important role. Morphology-based estimates of evolutionary chronology are highly sensitive to new fossil discoveries, the interpreta- tion and dating of such material, and the impact on tree topologies. Furthermore, the earliest and most primitive taxa are almost always known from very few fossil localities, with the result that these are likely to exert a disproportionate influence. Fossils and molecules should be treated as complementary approaches, rather than as conflicting and irreconcilable methods. Introduction Modern tetrapods have a long evolutionary history dating back to the Late Devonian. Their origins are rooted into a diverse, paraphyletic assemblage of lobe-finned bony fishes known as the ‘osteolepiforms’ (Cloutier and Ahlberg 1996; Janvier 1996; Ahlberg and Johanson 1998; Jeffery 2001; Johanson and Ahlberg 2001; Zhu and Schultze 2001).