2008 – Fourth Symposium on Hemlock
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Studies of the Laboulbeniomycetes: Diversity, Evolution, and Patterns of Speciation
Studies of the Laboulbeniomycetes: Diversity, Evolution, and Patterns of Speciation The Harvard community has made this article openly available. Please share how this access benefits you. Your story matters Citable link http://nrs.harvard.edu/urn-3:HUL.InstRepos:40049989 Terms of Use This article was downloaded from Harvard University’s DASH repository, and is made available under the terms and conditions applicable to Other Posted Material, as set forth at http:// nrs.harvard.edu/urn-3:HUL.InstRepos:dash.current.terms-of- use#LAA ! STUDIES OF THE LABOULBENIOMYCETES: DIVERSITY, EVOLUTION, AND PATTERNS OF SPECIATION A dissertation presented by DANNY HAELEWATERS to THE DEPARTMENT OF ORGANISMIC AND EVOLUTIONARY BIOLOGY in partial fulfillment of the requirements for the degree of Doctor of Philosophy in the subject of Biology HARVARD UNIVERSITY Cambridge, Massachusetts April 2018 ! ! © 2018 – Danny Haelewaters All rights reserved. ! ! Dissertation Advisor: Professor Donald H. Pfister Danny Haelewaters STUDIES OF THE LABOULBENIOMYCETES: DIVERSITY, EVOLUTION, AND PATTERNS OF SPECIATION ABSTRACT CHAPTER 1: Laboulbeniales is one of the most morphologically and ecologically distinct orders of Ascomycota. These microscopic fungi are characterized by an ectoparasitic lifestyle on arthropods, determinate growth, lack of asexual state, high species richness and intractability to culture. DNA extraction and PCR amplification have proven difficult for multiple reasons. DNA isolation techniques and commercially available kits are tested enabling efficient and rapid genetic analysis of Laboulbeniales fungi. Success rates for the different techniques on different taxa are presented and discussed in the light of difficulties with micromanipulation, preservation techniques and negative results. CHAPTER 2: The class Laboulbeniomycetes comprises biotrophic parasites associated with arthropods and fungi. -
COLEOPTERA COCCINELLIDAE) INTRODUCTIONS and ESTABLISHMENTS in HAWAII: 1885 to 2015
AN ANNOTATED CHECKLIST OF THE COCCINELLID (COLEOPTERA COCCINELLIDAE) INTRODUCTIONS AND ESTABLISHMENTS IN HAWAII: 1885 to 2015 JOHN R. LEEPER PO Box 13086 Las Cruces, NM USA, 88013 [email protected] [1] Abstract. Blackburn & Sharp (1885: 146 & 147) described the first coccinellids found in Hawaii. The first documented introduction and successful establishment was of Rodolia cardinalis from Australia in 1890 (Swezey, 1923b: 300). This paper documents 167 coccinellid species as having been introduced to the Hawaiian Islands with forty-six (46) species considered established based on unpublished Hawaii State Department of Agriculture records and literature published in Hawaii. The paper also provides nomenclatural and taxonomic changes that have occurred in the Hawaiian records through time. INTRODUCTION The Coccinellidae comprise a large family in the Coleoptera with about 490 genera and 4200 species (Sasaji, 1971). The majority of coccinellid species introduced into Hawaii are predacious on insects and/or mites. Exceptions to this are two mycophagous coccinellids, Calvia decimguttata (Linnaeus) and Psyllobora vigintimaculata (Say). Of these, only P. vigintimaculata (Say) appears to be established, see discussion associated with that species’ listing. The members of the phytophagous subfamily Epilachninae are pests themselves and, to date, are not known to be established in Hawaii. None of the Coccinellidae in Hawaii are thought to be either endemic or indigenous. All have been either accidentally or purposely introduced. Three species, Scymnus discendens (= Diomus debilis LeConte), Scymnus ocellatus (=Scymnobius galapagoensis (Waterhouse)) and Scymnus vividus (= Scymnus (Pullus) loewii Mulsant) were described by Sharp (Blackburn & Sharp, 1885: 146 & 147) from specimens collected in the islands. There are, however, no records of introduction for these species prior to Sharp’s descriptions. -
Yorkhill Green Spaces Wildlife Species List
Yorkhill Green Spaces Wildlife Species List April 2021 update Yorkhill Green Spaces Species list Draft list of animals, plants, fungi, mosses and lichens recorded from Yorkhill, Glasgow. Main sites: Yorkhill Park, Overnewton Park and Kelvinhaugh Park (AKA Cherry Park). Other recorded sites: bank of River Kelvin at Bunhouse Rd/ Old Dumbarton Rd, Clyde Expressway path, casual records from streets and gardens in Yorkhill. Species total: 711 Vertebrates: Amhibians:1, Birds: 57, Fish: 7, Mammals (wild): 15 Invertebrates: Amphipods: 1, Ants: 3, Bees: 26, Beetles: 21, Butterflies: 11, Caddisflies: 2, Centipedes: 3, Earthworms: 2, Earwig: 1, Flatworms: 1, Flies: 61, Grasshoppers: 1, Harvestmen: 2, Lacewings: 2, Mayflies: 2, Mites: 4, Millipedes: 3, Moths: 149, True bugs: 13, Slugs & snails: 21, Spiders: 14, Springtails: 2, Wasps: 13, Woodlice: 5 Plants: Flowering plants: 174, Ferns: 5, Grasses: 13, Horsetail: 1, Liverworts: 7, Mosses:17, Trees: 19 Fungi and lichens: Fungi: 24, Lichens: 10 Conservation Status: NameSBL - Scottish Biodiversity List Priority Species Birds of Conservation Concern - Red List, Amber List Last Common name Species Taxon Record Common toad Bufo bufo amphiban 2012 Australian landhopper Arcitalitrus dorrieni amphipod 2021 Black garden ant Lasius niger ant 2020 Red ant Myrmica rubra ant 2021 Red ant Myrmica ruginodis ant 2014 Buff-tailed bumblebee Bombus terrestris bee 2021 Garden bumblebee Bombus hortorum bee 2020 Tree bumblebee Bombus hypnorum bee 2021 Heath bumblebee Bombus jonellus bee 2020 Red-tailed bumblebee Bombus -
Coccinellidae)
ECOLOGY AND BEHAVIOUR OF THE LADYBIRD BEETLES (COCCINELLIDAE) Edited by I. Hodek, H.E van Emden and A. Honek ©WILEY-BLACKWELL A John Wiley & Sons, Ltd., Publication CONTENTS Detailed contents, ix 8. NATURAL ENEMIES OF LADYBIRD BEETLES, 375 Contributors, xvii Piotr Ccryngier. Helen E. Roy and Remy L. Poland Preface, xviii 9. COCCINELLIDS AND [ntroduction, xix SEMIOCHEMICALS, 444 ]an Pettcrsson Taxonomic glossary, xx 10. QUANTIFYING THE IMPACT OF 1. PHYLOGENY AND CLASSIFICATION, 1 COCCINELLIDS ON THEIR PREY, 465 Oldrich Nedved and Ivo Kovdf /. P. Mid'laud and James D. Harwood 2. GENETIC STUDIES, 13 11. COCCINELLIDS IN BIOLOGICAL John J. Sloggett and Alois Honek CONTROL, 488 /. P. Midland 3. LIFE HISTORY AND DEVELOPMENT, 54 12. RECENT PROGRESS AND POSSIBLE Oldrkli Nedved and Alois Honek FUTURE TRENDS IN THE STUDY OF COCCINELLIDAE, 520 4. DISTRIBUTION AND HABITATS, 110 Helmut /; van Emden and Ivo Hodek Alois Honek Appendix: List of Genera in Tribes and Subfamilies, 526 5. FOOD RELATIONSHIPS, 141 Ivo Hodek and Edward W. Evans Oldrich Nedved and Ivo Kovdf Subject index. 532 6. DIAPAUSE/DORMANCY, 275 Ivo Hodek Colour plate pages fall between pp. 250 and pp. 251 7. INTRAGUILD INTERACTIONS, 343 Eric Lucas VII DETAILED CONTENTS Contributors, xvii 1.4.9 Coccidulinae. 8 1.4.10 Scymninae. 9 Preface, xviii 1.5 Future Perspectives, 10 References. 10 Introduction, xix Taxonomic glossary, xx 2. GENETIC STUDIES, 13 John J. Sloggett and Alois Honek 1. PHYLOGENY AND CLASSIFICATION, 1 2.1 Introduction, 14 Oldrich Nedved and Ivo Kovdf 2.2 Genome Size. 14 1.1 Position of the Family. 2 2.3 Chromosomes and Cytology. -
Faunističko-Ekološke Značajke Bubamara (Coleoptera,Coccinellidae) Grada Nove Gradiške I Okolnih Naselja
Faunističko-ekološke značajke bubamara (Coleoptera,Coccinellidae) grada Nove Gradiške i okolnih naselja Ribarić, Ana Master's thesis / Diplomski rad 2020 Degree Grantor / Ustanova koja je dodijelila akademski / stručni stupanj: Josip Juraj Strossmayer University of Osijek, Department of biology / Sveučilište Josipa Jurja Strossmayera u Osijeku, Odjel za biologiju Permanent link / Trajna poveznica: https://urn.nsk.hr/urn:nbn:hr:181:754773 Rights / Prava: In copyright Download date / Datum preuzimanja: 2021-10-06 Repository / Repozitorij: Repository of Department of biology, Josip Juraj Strossmayer University of Osijek Sveučilište Josipa Jurja Strossmayera u Osijeku Odjel za biologiju Diplomski sveučilišni studij Biologija i kemija; smjer: nastavnički Ana Ribarić Faunističko – ekološke značajke bubamara (Coleoptera, Coccinellidae) grada Nove Gradiške i okolnih naselja Diplomski rad Osijek, 2020. TEMELJNA DOKUMENTACIJSKA KARTICA Diplomski rad Sveučilište Josipa Jurja Strossmayera u Osijeku Odjel za biologiju Diplomski sveučilišni studij Biologija i kemija; smjer: nastavnički Znanstveno područje: Prirodne znanosti Znanstveno polje: Biologija FAUNISTIČKO – EKOLOŠKE ZNAČAJKE BUBAMARA (COLEOPTERA, COCCINELLIDAE) GRADA NOVE GRADIŠKE I OKOLNIH NASELJA Ana Ribarić Rad je izrađen na: Odjel za biologiju, Zavod za zoologiju Mentor: Dr. sc. Nataša Turić, docent Komentor: Dr. sc. Goran Vignjević, docent Kratak sažetak diplomskog rada: Bubamare ili božje ovčice su kukci (Insecta) iz reda kornjaša (Coleoptera) i porodice Coccinellidae. Cilj ovog rada bio je utvrditi prisutnost i faunistički sastav bubamara na području grada Nove Gradiške i okolnih mjesta unutar Brodsko – posavske županije. Tijekom dvomjesečnog istraživanja 2020. godine, na 6 lokaliteta, uzorkovane su ukupno 622 jedinke iz porodice Coccinellidae. Jedinke su svrstane u 10 vrsta, 10 rodova i 3 potporodice. Kao najbrojnija vrsta utvrđena je Harmonia axyridis s uzorkovanih 373 jedinke odnosno 59,97% ukupnog broja jedinki, a slijedi vrsta Coccinella septempunctata sa 153 uzorkovane jedinke, tj. -
Heteroptera: Anthocoridae, Lasiochilidae)
2018 ACTA ENTOMOLOGICA 58(1): 207–226 MUSEI NATIONALIS PRAGAE doi: 10.2478/aemnp-2018-0018 ISSN 1804-6487 (online) – 0374-1036 (print) www.aemnp.eu RESEARCH PAPER Annotated catalogue of the fl ower bugs from India (Heteroptera: Anthocoridae, Lasiochilidae) Chandish R. BALLAL1), Shahid Ali AKBAR2,*), Kazutaka YAMADA3), Aijaz Ahmad WACHKOO4) & Richa VARSHNEY1) 1) National Bureau of Agricultural Insect Resources, Bengaluru, India; e-mail: [email protected] 2) Central Institute of Temperate Horticulture, Srinagar, 190007 India; e-mail: [email protected] 3) Tokushima Prefectural Museum, Bunka-no-Mori Park, Mukoterayama, Hachiman-cho, Tokushima, 770–8070 Japan; e-mail: [email protected] 4) Department of Zoology, Government Degree College, Shopian, Jammu and Kashmir, 192303 India; e-mail: [email protected] *) Corresponding author Accepted: Abstract. The present paper provides a checklist of the fl ower bug families Anthocoridae th 6 June 2018 and Lasiochilidae (Hemiptera: Heteroptera) of India based on literature and newly collected Published online: specimens including eleven new records. The Indian fauna of fl ower bugs is represented by 73 5th July 2018 species belonging to 26 genera under eight tribes of two families. Generic transfers of Blap- tostethus pluto (Distant, 1910) comb. nov. (from Triphleps pluto Distant, 1910) and Dilasia indica (Muraleedharan, 1978) comb. nov. (from Lasiochilus indica Muraleedharan, 1978) are provided. A lectotype is designated for Blaptostethus pluto. Previous, as well as new, distribu- -
Taxon Vernacular Date Stage Comment Adalia Decempunctata 10-Spot Ladybird 09-Apr-06 Adult on Ilex Adalia Decempunctata 10-Spot L
Species list for Deep Dene, Bitterne Park (SU443137) by Philip Budd Taxon Vernacular Date Stage Comment Adalia decempunctata 10-spot Ladybird 09-Apr-06 Adult On Ilex Adalia decempunctata 10-spot Ladybird 11-Sep-06 Adult Off Betula Adalia bipunctata 2-spot Ladybird 11-Sep-06 Adult Coccinella septempunctata 7-spot Ladybird 09-Apr-06 Adult In dead Quercus leaves Episyrphus balteatus a hoverfly 26-Mar-12 Adult Poa annua Annual Meadow-grass 26-Mar-12 Not recorded Fraxinus excelsior Ash 26-Mar-12 Not recorded Laurus nobilis Bay 26-Mar-12 Not recorded Birdsown Laurus nobilis Bay 09-Apr-06 Not recorded 1 seen Laurus nobilis Bay 19-Nov-04 Not recorded 2 seen Fagus sylvatica Beech 26-Mar-12 Not recorded Pinus wallichiana Bhutan Pine 26-Mar-12 Not recorded 2 planted Elasmostethus interstinctus Birch Shieldbug 11-Sep-06 Adult Off Ilex and Betula Hyacinthoides non-scripta Bluebell 26-Mar-12 Not recorded Psilocybe cyanescens Blueleg Brownie 24-Nov-04 Not recorded Gonocerus acuteangulatus Box Bug 11-Sep-06 Adult 1 off Ligustrum ovalifolium Pteridium aquilinum Bracken 26-Mar-12 Not recorded Rubus fruticosus agg. Bramble 26-Mar-12 Not recorded Rumex obtusifolius Broad-leaved Dock 26-Mar-12 Not recorded Troilus luridus Bronze Shieldbug 09-Apr-06 Adult In dead Fagus leaves Buddleja davidii Butterfly-bush 26-Mar-12 Not recorded Buteo buteo Buzzard 07-Dec-11 Adult 1 seen Xylaria hypoxylon Candle Snuff 24-Nov-04 Not recorded Prunus laurocerasus Cherry Laurel 26-Mar-12 Not recorded Planted Prunus cerasifera Cherry Plum 26-Mar-12 Not recorded Probably birdsown -
An Inventory of Nepal's Insects
An Inventory of Nepal's Insects Volume III (Hemiptera, Hymenoptera, Coleoptera & Diptera) V. K. Thapa An Inventory of Nepal's Insects Volume III (Hemiptera, Hymenoptera, Coleoptera& Diptera) V.K. Thapa IUCN-The World Conservation Union 2000 Published by: IUCN Nepal Copyright: 2000. IUCN Nepal The role of the Swiss Agency for Development and Cooperation (SDC) in supporting the IUCN Nepal is gratefully acknowledged. The material in this publication may be reproduced in whole or in part and in any form for education or non-profit uses, without special permission from the copyright holder, provided acknowledgement of the source is made. IUCN Nepal would appreciate receiving a copy of any publication, which uses this publication as a source. No use of this publication may be made for resale or other commercial purposes without prior written permission of IUCN Nepal. Citation: Thapa, V.K., 2000. An Inventory of Nepal's Insects, Vol. III. IUCN Nepal, Kathmandu, xi + 475 pp. Data Processing and Design: Rabin Shrestha and Kanhaiya L. Shrestha Cover Art: From left to right: Shield bug ( Poecilocoris nepalensis), June beetle (Popilla nasuta) and Ichneumon wasp (Ichneumonidae) respectively. Source: Ms. Astrid Bjornsen, Insects of Nepal's Mid Hills poster, IUCN Nepal. ISBN: 92-9144-049 -3 Available from: IUCN Nepal P.O. Box 3923 Kathmandu, Nepal IUCN Nepal Biodiversity Publication Series aims to publish scientific information on biodiversity wealth of Nepal. Publication will appear as and when information are available and ready to publish. List of publications thus far: Series 1: An Inventory of Nepal's Insects, Vol. I. Series 2: The Rattans of Nepal. -
Implementation and Status of Biological Control of the Hemlock Woolly Adelgid
Forest Health Technology Enterprise Team TECHNOLOGY TRANSFER Biological Control IMPLEMENTATION AND STAtuS OF BIOLOGICAL CONTRol OF THE HEMLOCK WOOLLY AdelgId United States Forest Forest Health FHTET-2011-04 Department Service Technology December 2011 of Agriculture Enterprise Team Forest Health Technology Enterprise Team TECHNOLOGY TRANSFER Biological Control IMPLEMENTATION AND STAtuS OF BIOLOGICAL CONTRol OF THE HEMLOCK WOOLLY AdelgId United States Forest Forest Health FHTET-2011-04 Department Service Technology December 2011 of Agriculture Enterprise Team IMPLEMENTATION AND STATUS OF BIOLOGICAL CONTROL OF THE HEMLOCK WOOLLY ADELGID The Forest Health Technology Enterprise Team (FHTET) was created in 1995 by the Deputy Chief for State and Private Forestry, Forest Service, U.S. Department of Agriculture, to develop and deliver technologies to protect and improve the health of American forests. This book was published by FHTET as part of the technology transfer series. http://www.fs.fed.us/foresthealth/technology/ On the cover: Center photo: Hemlock woolly adelgid white woolly masses on a hemlock branch (USDA Forest Service, Karen Felton) Top right: Sasajiscymnus tsugae predatory beetle (USDA Forest Service, Lynn Jones); Middle right: Collecting and checking hemlock branch samples for Laricobius nigrinus predatory beetle larvae (USDA Forest Service, Brad Onken); Bottom center: Collecting Laricobius nigrinus predatory beetles in Idaho (USDA Forest Service, Brad Onken); Middle left: Releasing Laricobius nigrinus predatory beetles (USDA Forest Service, Brad Onken); Top left: Laricobius nigrinus predatory beetle (USDA Forest Service, Lynn Jones) For additional copies of this publication, contact: Brad Onken Richard Reardon U.S. Forest Service U.S. Forest Service 180 Canfi eld Street 180 Canfi eld Street Morgantown, WV 26505 Morgantown, WV 26505 (304) 285-1546 (304) 285-1566 [email protected] [email protected] The entire publication is available online at http://www.fs.fed.us/na/morgantown/fhp/hwa The U.S. -
Phylogeny of Ladybirds (Coleoptera: Coccinellidae): Are the Subfamilies Monophyletic?
Molecular Phylogenetics and Evolution 54 (2010) 833–848 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev Phylogeny of ladybirds (Coleoptera: Coccinellidae): Are the subfamilies monophyletic? A. Magro a,b,1, E. Lecompte b,c,*,1, F. Magné b,c, J.-L. Hemptinne a,b, B. Crouau-Roy b,c a Université de Toulouse, ENFA, EDB (Laboratoire Evolution et Diversité Biologique), 2 route de Narbonne, F-31320 Castanet Tolosan, France b CNRS, EDB (Laboratoire Evolution et Diversité Biologique), F-31062 Toulouse, France c Université de Toulouse, UPS, EDB (Laboratoire Evolution et Diversité Biologique), 118 route de Narbonne, F-31062 Toulouse, France article info abstract Article history: The Coccinellidae (ladybirds) is a highly speciose family of the Coleoptera. Ladybirds are well known Received 20 April 2009 because of their use as biocontrol agents, and are the subject of many ecological studies. However, little Revised 15 October 2009 is known about phylogenetic relationships of the Coccinellidae, and a precise evolutionary framework is Accepted 16 October 2009 needed for the family. This paper provides the first phylogenetic reconstruction of the relationships Available online 10 November 2009 within the Coccinellidae based on analysis of five genes: the 18S and 28S rRNA nuclear genes and the mitochondrial 12S, 16S rRNA and cytochrome oxidase subunit I (COI) genes. The phylogenetic relation- Keywords: ships of 67 terminal taxa, representative of all the subfamilies of the Coccinellidae (61 species, 37 genera), Phylogeny and relevant outgroups, were reconstructed using multiple approaches, including Bayesian inference Coccinellidae Partitioned analyses with partitioning strategies. The recovered phylogenies are congruent and show that the Coccinellinae Evolution is monophyletic but the Coccidulinae, Epilachninae, Scymninae and Chilocorinae are paraphyletic. -
Glasgow Necropolis (November 2010) Group Formal Name
Glasgow Necropolis (November 2010) Group Formal Name Common Name Earliest RecordLatest Record annelid Lumbricus terrestris Linnaeus, 1758 Common Earthworm 10/05/2007 10/05/2007 bird Accipiter nisus (Linnaeus, 1758) Sparrowhawk 29/07/2010 29/07/2010 bird Aegithalos caudatus (Linnaeus, 1758) Long-tailed Tit 18/03/2009 18/03/2009 bird Carduelis carduelis (Linnaeus, 1758) Goldfinch 05/04/2008 26/05/2010 bird Carduelis chloris (Linnaeus, 1758) Greenfinch 05/04/2008 09/06/2010 bird Erithacus rubecula (Linnaeus, 1758) Robin 05/04/2008 05/04/2008 bird Falco tinnunculus Linnaeus, 1758 Kestrel 09/06/2010 09/06/2010 bird Fringilla coelebs Linnaeus, 1758 Chaffinch 05/04/2008 26/05/2010 bird Parus major Linnaeus, 1758 Great Tit 05/04/2008 05/04/2008 bird Pica pica (Linnaeus, 1758) Magpie 11/05/2005 11/05/2005 bird Prunella modularis (Linnaeus, 1758) Dunnock 26/05/2010 26/05/2010 bird Sturnus vulgaris Linnaeus, 1758 Starling 11/05/2005 11/05/2005 bird Troglodytes troglodytes (Linnaeus, 1758) Wren 05/04/2008 09/06/2010 bird Turdus merula Linnaeus, 1758 Blackbird 11/05/2005 26/05/2010 bird Turdus viscivorus Linnaeus, 1758 Mistle Thrush 09/05/2006 29/07/2010 centipede Geophilus flavus (De Geer, 1778) 10/05/2007 10/05/2007 centipede Lithobius (Lithobius) forficatus (Linnaeus, 1758) the common centipede 12/05/2008 12/05/2008 centipede Stigmatogaster subterranea (Shaw, 1789) 10/05/2007 10/05/2007 clubmoss Lycopodium clavatum L. Stag's-horn Clubmoss 25/08/1985 1988 conifer Chamaecyparis pisifera (Siebold & Zucc.) Siebold & Zucc. Sawara Cypress July 2006 July 2006 fern Asplenium ruta-muraria L. -
Integrative Taxonomy Reveals Hidden Species Within a Common Fungal Parasite of Ladybirds Received: 19 April 2018 Danny Haelewaters 1,2, André De Kesel3 & Donald H
www.nature.com/scientificreports OPEN Integrative taxonomy reveals hidden species within a common fungal parasite of ladybirds Received: 19 April 2018 Danny Haelewaters 1,2, André De Kesel3 & Donald H. Pfster1 Accepted: 8 October 2018 Our understanding of fungal diversity is far from complete. Species descriptions generally focus on Published: xx xx xxxx morphological features, but this approach may underestimate true diversity. Using the morphological species concept, Hesperomyces virescens (Ascomycota, Laboulbeniales) is a single species with global distribution and wide host range. Since its description 120 years ago, this fungal parasite has been reported from 30 species of ladybird hosts on all continents except Antarctica. These host usage patterns suggest that H. virescens could be made up of many diferent species, each adapted to individual host species. Using sequence data from three gene regions, we found evidence for distinct clades within Hesperomyces virescens, each clade corresponding to isolates from a single host species. We propose that these lineages represent separate species, driven by adaptation to diferent ladybird hosts. Our combined morphometric, molecular phylogenetic and ecological data provide support for a unifed species concept and an integrative taxonomy approach. What is a species? Tis is a perennial question in evolutionary biology. Te answer is complex and has been intensely argued for decades. Diferent species concepts corresponding to multiple biological properties pro- vide a means to recognize, delineate and describe species. Tese properties include diferences in morphological traits, nucleotide divergence and monophyly, reproductive isolation, ecological niches or adaptive zones, mate recognition or mating systems, geographic range, exclusive coalescence of alleles, etc. However, biologists from various research felds have advocated diferent and sometimes incompatible species concepts, leading to varying conclusions regarding delimitation of species and their numbers.