Notes on some Scandinavian (Diptera: )

JAROSLAW TRZYWTNSTT

Krzywiriski, J. Notes on some Scandinavian Palpomyiini (Diptera: Ceratopogonidae). [Data om nigra skandinavistiska svidknott av tribus Palpomyiini (Diptera: Ceratopogoni' dae).] - Ent. Tidskr. I l7 (3): I 1 3- I 19. Uppsala, Sweden 1996. ISSN 0013-886x.

Records of ten of Palpomyiini (Diptera: Ceratopogonidae) new to the faunas of Swe- den lPalpomyia brachialis (Hal.), P luteifemorata Edw., P nigripes (Meig'), P. semifumosa Gtgh., P spinipes (Panzer), rubiginosa (Winn.), albicornis (Meig.), B. nobilis (Winn.), B. taeniata (Hal. in Walker)l and Norway fP. pubescens Kieff.l are given along with the diagnoses allowing for their recognition. Data on general distribution of each species are presented. The genus Phaenobezzia is reported for the first time in Scandinavia.

J. Krzywiiski, Department of Invertebrate Zoology, University of Gdansk, Al. Pilsudskiego 46, 81-378 Gdynia, Poland.

Introduction The Ceratopogonidae or biting midges are small lection of the Zoological Museum, University of (length l-4 mm) nematocerous . Except for Lund) l9 species have been determined, among hematophagous forms attacking man and live- them 9 species of Bezzia, and Phaeno- stock they have received little attention. Even bezzia not recorded in this country as yet. In the relatively big in size predaceous species, such as collection of biting midges in the Natural History those belonging to the tribe Palpomyiini, that in Museum (London) one species of Palpomyiafrom immature stages are common inhabitants of va- Norway new to the fauna of this country has also rious aquatic and semi-aquatic situations, are still been found. Specimens belonging to each species poorly known. were carefully compared with existing types Scandinavian biting midges have been studied (listed in the paper) and other specimens from out- by several authors. Data concerning species of side Scandinavia, data of which are not included Palpomyiini were given by Staeger (1839), here. Zetterstedt (1838, 1850, 1855), Lundstrtim (1910, 1916, 1917), Kieffer (1915, l9l6), Edwards (1931), Stori (1939) and more recently by Results Clastrier (1962a, b). Hackman ( I 980) summarized All species new to the Scandinavian fauna are re- Finnish species in a check list. A total number of ported below. Diagnoses and other notes allowing 27 species of Palpomyiini have been encountered for their recognition as well as data on their gene- in Scandinavia (Tab. I .). The present paper reports ral distribution are given along with the records. P additional species new to the faunas of two pubescens, P. semifumosa and P. spinipe.t are Pre- countries of the region. sented in the new interpretation, justification of which is given in a revisionary paper ot Pal- pomyia of Central Europe (in preparation). The Material studied abbreviations for parts of Russia follow "Cata- the explanation The present study is based upon the examination logue of Palaearctic Diptera". For of Swedish and Norwegian specimens of of the special terms used in the paper, see (in print). Palpomyiini. In Swedish material (from the col- Szadziewski et al. r13 Jaroslaw Krzywiriski Ent. Tidskr. ll7 (1996)

Tab. 1. A list of species of PaLpomyiini recorded from Scandinavian specimens. SWEDEN: Sk: Scandinavia (D - Denmark, S - Sweden, F - Finland, N - Forsakar, 2.08. 198 I , 1 Q; T. Hiillestad, Grytiingen, Norway). 21.07.1975,1 Q; Kullaberg, 6.08.1975, 2 9; Kul- laberg,22.O8.1983,2 Q. All specimens H. Anders- Arter av svidknott ur gruppen Palpomyiini som rappor- son leg. (ZML). terats frdn Norden (D Danmark, S Sverige, F Fin- - - - Distribution. P brachialis is an arboreal Euro- land, N - Norge). pean species known from ?Spain, Great Britain, Belgium, Germany, ?Austria, Hungary, Slovakia, D S F N Lithuania, Byelorussia, Ukraine and Russia Palpomyia armipe s (Meigen) + + + + (CET). For the first time recorded from Sweden. P. ate rrima Goetghebuer + + Note. In colour of legs and somewhat swollen P. bispinosa Kieffer + fore femur P. brachialis resembles P. distincta P concoloripes Clastrier + (Haliday) the most. However, the latter species P distincta (Haliday) + + differs in broadly dark apices of mid and hind P. (Meigen) + + + + flavipes femora, furthermore female yellow P. lineata (Meigen) + + + in scutellum, P. lundstroemiRemm + + claws with small basal inner tooth and presence of P pubescens Kieffer + + two lobes arising from the eighth sternum, and P. rufipes (Meigen) + male in setose aedeagus and the presence of P. serripes (Meigen) + + mesoventral lobe on gonocoxite. P spinipes (Panzer) + P. tibialis (Meigen) + + + Be zzia affinis (Staeger) + B. annulipes (Meigen) + + + Palp o my ia luteifu morata Edwards B. bicolor (Meigen) + + Palpom1-ia Iuteifemorata Edwards, 1926 419. B. coracina (Zetterstedt) + + Diagnosis. Body covered numerous small B. flav ico rnis (Staeger) + with B. le uco gaste r (Zetterstedt) + + + spear-shaped setae. Femora yellow; fore and mid B. nigrita Clastrier + tibiae brownish yellow or brown (mid tibia lighter B. ni gritula (Zenersted0 + + + than fore one), hind tibia blackish brown. B. ornata (Meigen) + Types. Female holotype of P. luteifemorataEd- B. p il ipennis Lundstr6m + wards, paratypes - 3 females and 2 males in NHM, B. signata (Meigen) + London. B. solstitialis (Winnertz) + Scandinavian specimens. SWEDEN: Dlr: B. transfuga (Staeger) + + + Leksand, 18.07.1979, B. winne rtziano Kieffer + Singin, T. Tjeder leg., lQ (zML). Distribution. This is an arboreal European spe- cies known from Great Britain, Belgium, Ger- Palp o my (Haliday) ia brac hialis many, Estonia and Ukraine (Crimea). For the first (Fie. l.) time recorded from Sweden. Ceratopogon brachialis Haliday, 1833: 152. Note. Among European species of Palpomyia Palpomyia brachialis (Haliday); Kieffer, I 906. only P luteftmorata and P. floralis (Meigen) Palpomyia longipennis Kieffer, I 919: 104. have spear-shaped setae. However, P. floralis can Palpomyia fusciclava Kieffer, I 9 l9: I 09. be easily distinguished by darker legs (at least Palpomyia nemorivaga Goetghebuer, 1920: 85. hind femur is dark in distal halfl. Diagnosis. Legs almost totally yellow, only hind femur black at apex, knees brownish black, distal end of hind tibia brown; fore femur slightly Palpomyia n igrip e s (Meigen) thickened, with 7-11 spines. Body covered with Ceratopogon nigripes Meigen, 1830: 265. simple, not modified setae. Halteres brown. Palpoml-ia nigripes (Meigen); Kieffer, 1906: 63. Gonostyle weakly bent, with straight end. Ceratopogon pratensis Meigen, 1830: 264. Tlpes. Three female syntypes of P. nemorivaga Palpomyia prarensls (Meigen); Kieffer, 1906: 63. Goetghebuer in IRScNB, Brussels. Polpomyia brr^ocrypta Kieffer, 1925a: 425. fi4 Ent. Tidskr. 117 (1996) Some S cand inav ian P alpomyi ini Diagnosis. Legs brownish black; fore femur yellow or light brown in a proximal half, mid and hind femora yellowish at bases. Fore femur with 4-5 slightly elevated spines. Female claws without basal inner tooth; 9th sternum of female abdomen very narrow, not divided, heavily sclerotized. Parameres fused, forming a slender median pro- cess basally and widened, spoon-shaped distal portion; aedeagus greatly reduced; gonocoxite recurved 90" basally. Types. Four female syntypes of C. nigripes Meigen in MNHN. Paris. Scandinavian specimens. SWEDEN: Ha: Breared, Klovberget, 19.06. 1984, H. Andersson leg., I I @ML). Distribution. This is arboreal European species known from Great Britain, France, Belgium, the Netherlands, Germany, ?Czech, ?Slovakia, Pol- and and Russia (CET). Recorded for the first time from Sweden. Note. P. nigripes belongs to a complex of four species, the females of which are characterized by the shape of ninth sternum and males by highly characteristic structure of genitalia described above. Besides P. nigripes two other species, P remmi Havelka and P globulifera Remm inhabit the Palaearctic Region; the fourth, P. canadensis Grogan & Wirth is known from the Nearctic Re- gion. P nigripes differs from other members of the complex in dark colour of legs, mainly in entirely dark tibiae. Fig. 1. Palpomyia brachi.alis, female habitus (Length bar below = I mm). Drawing: J. Krzywiriski.

p ube sce ns Kieffer Palpomyia Hona av svidknottet Palpomyia brachialis (skalstreck = Palpomyio pubescens Kieffer, 1919: 105. 1 mm). De 14 mm Ldnga honorna av gruppen Palpo' Palpomyia crassipes Goetghebuer, 1920: 112. myiini fdngar ,sina byten i flykten. De attackerar ncistan Palpomyia spinipes sensu Goetghebuer nec Panzer, enbart hannar av andra myqqor eller mindre dag' 1922:53 sliindor. Honan iiter vanligen ocksd upp hannen under Palpomyia turfacea Kieffer, 1925b: 155. parningen, varvid hans kropp reduceras till en tom kuti- livncir sig honorna av blomnektar Diagnosis. Small , wing length (measured kula. Dcirutdver (frr)mst av vdxter), liksom han- from the basal arculus to the wing tip) 1.55-2.18 familjenflockblomstriga narna, som inte dr predatorer mm in female and 1.23-1.32 mm in male. Body black. Halter yellow. Legs yellow; fore femur with black tip, distal 1/3 of mid and 1/2 of hind femora brown or black; fore and mid tibiae with brown distal end, hind tibia totally brown or black. Fore four flagellomeres divided by the sum of the femur greatly swollen, with l1-15 spines. Male lengths of the preceding nine) lower than 1.10. anepisternum with 1-7 setae, katepisternum bare; Types. Female syntype of P. crassipes parameres gradually narrowing in the distal part, Goetghebuer in IRScNB, Brussels. with undivided, more or less sharp, setose tip. Scandinavian specimens. NORWAY: Laerdal, Male antennal ratio (sum of the lengths of distal 24-27.07,1923,F. W. Edwards leg., I 9 (NHM). l15 Jaroslaw Krzywifiski Ent. Tidskr. 117 (1996)

Distribution. According to Remm (1976) P. Palp omy ia s pinip e s (Panzer) pubescens inhabits cold situations - north taiga Ceratopogon spinipes Panzer, I 806: 14. and the mountains in the south. Known from Great P a lpomy ia sp inip e s (Panzer); Kieffer, 1 90 1 : I 57. Britain, Belgium, the Netherlands, Germany, Ceratopogon fulvus Macquart, I 826: 18 1. Austria, ?Hungary, Czech, Slovakia, Poland, Swe- P alpomy ia fulva (Macquart); Kieffer, 1906: 63. den, Finland, Estonia, Latvia, Ukraine (Crimea) Ceratopogon ferrugineus Meigen, 1830; 265. i (Meigen) 1 and Russia (Kaliningrad Distr., NET, ES, FE). P alpo my ia fe rrug nea ; Kieffer, 90 I : I 57. Palpomyia parvtforceps Kieffer, 1925a: 98. This is the first record from Norway. Note. From other species of distincta group of Diagnosis. Female: head reddish, thorax yel- Palpomyia with pale halteres females of P lowish red or red, abdomen whitish. Legs yellow; pubescens differ in totally dark hind tibia, male in distal l/5 of mid femur and l/4 of hind femur small size, lack of setae on katepisternum, and black, fore and mid tibiae brown apically, distal 1/ parameres undivided in distal part. Male of P 5 of hind tibia black; femora slender; fore femur distincta, which is the most similar to P bearing 8-13, mid 0-3 and hind 2-5 spines. Male: pubescens, differs from this species in higher body brown; legs similar to female in colour, value of antennal ratio (over 1. l0) and bigger size however distal l/4 of mid and l/3 of hind femora (wing length 1.27-1.68 mm). brown; parameres fused at base and deeply divided in a distal half. Ilpes. Female holotype of C. ferrugineus Meigen in MNHN, Paris. Palpomy ia s e mifumo sa Goetghebuer Scandinavian specimens. SWEDEN: Sk: Norr Palpomyia semifumosa Goetghebuer, 1922: 53 (= Krankesjdn, 3.07.1975,4 9; Norr Krankesjrin, hortulana sensu Goetghebuer, 1920). Fukt. Lrivskog,3.08.1974, I 9; Norr Krankesjrin, Palpomyia hortulana sensu Goetghebuer nec Meigen, Sjristrand, sand, 3.07.1975, 3 9; Norr Krankesjtin, 1920:82. Torrhed, 23.07 .1980, 2 9. Kullaber g, I 1.07 .197 6, Diagnosis. Body covered with small simple setae. | 9;22.07.1978, 1 9;5.08.1982, I Q. All speci- Legs yellow; fore femur with black extreme tip; mens H. Andersson leg. (ZML). distal 1/3 of mid and hind femora dark brown; mid Distribution. This is an arboreal Palaearctic tibia yellowish brown, in proximal half and distal species, known from Great Britain, France, Bel- end darkened, hind tibia totally blackish brown; gium, the Netherlands, Germany, Austria, Italy, hind tarsus brown. Fore femur with 4-7 slightly Hungary, Poland, Denmark and Georgia. For the elevated spines. Wing narroq darkened in a distal first time recorded from Sweden. half. Female with small, spherical spermathecae. Note. P fulvescens Kieffer described from Male: gonostyle weakly bent, with straight end; Hungary is similar to P. spinipes, but differs in aedeagus narrow above basal arms; distal portion dark brown halteres and slightly thickened fore of parameres slender, with slightly bulbous tip. femora. Ilpes. Two female syntypes of P. semifumosa Goetghebuer in IRScNB, Brussels. Scandinavian specimens. SWEDEN: Sk: Phaenob ezzia rub igino s a (Winnertz) Torna Hiillestad, I1.06.1982, C. Hansson leg., I d Ceratopogon rubiginosus Winnertz, 1852 72. (zML). Bellia rubiginosc (Winnertz); Kieffer, 1901: 153. Distribution. Species known from Belgium, Probezzia rubiginosa (Winnertz); Kieffer, 1925a: 121. the Netherlands, Great Britain, Armenia and Be77ia glyceriae Kieffer, 1913: 9. P robezzia nitidivent ris Goetghebuer, Georgia. This is the first record from Sweden. 1923: 104 Note. P semifumosa is very similar to P Diagnosis. Head, thorax, and abdomen as well as flavipes Meigen aod P. Jloralis (Meigen) in leg halteres brown. Legs brownish yellow or light coloration, but is easily distinguished by rhe brown. Femora unarmed. presence of small simple unmodified setae. P Scandinavian specimens. SWEDEN: Sk: Norr citrinipes Kieffer and P. puberula Remm, that are Krankesjdn, Fukt. Lrivskog, 30.07.1974, H. An- probably closely related to P. semifumosa ate derssonleg., 19 @}l{L). much smaller in size. Distribution. P. rubiginosa is an arboreal Pa- ll6 Ent. Tidskr. ll7 (1996) S ome S c and inav ian P a lp omy i ini laearctic species known from Great Britain, morphology of their genitalia and females have Belgium, the Netherlands, Germany, Austria, Pol- strongly sclerotized ninth sternum with very and, Estonia, Latvia, Lithuania and Russia (CEl narrow, nearly straight and pointed arms. At pre- ES, FE). For the first time reported from Sweden. sent only males can be determined to species in This is also the first record of the genus this group. P haeno b e zzia in Scandinavia. Note. This is the only species of Phaenobezzia known in Europe. On the basis of male Tokunaga (Winnertz) (1939) described from Japan closely related P Ceratopogon nobilis Winnertz, 1852: '19. minutistyla which is distinguished ftom P. ru- Bezzia nobilis (Winnertz); Kieffer, I 90 I : I 53. biginosa by much wider and shorter gonocoxites. Ceratopogon setulosus Loew, 1861: 312. The Nearctic Region inhabits very similar P Bezzia setulosa (Loew); Johannsen, 1905: 102. Ceratopogon barbe ri Coquillett, : 601. opaca (Loew), which differs from P rubiginosain l90l Bezzia barberi (Coquillett); Malloch, 19l4: 282. having dark brown hind tibia and higher value of Bezzia leucosticta Kieffer, l9l9: 124. (Wirth antennal ratio and Grogan 1982). Bezzia chrysocoma Kieffer, 1922l 355. Bezzia acanthodes Macfie, l94O: 192. Bezzia atlantica Wirth & Williams, 1957: 13. Bezzia albicornis (Meigen) B e zzia no b il iformis Clastrier, 1962a: 7 4. Ceratopogon albicornis Meigen, l8l8: 74. Diagnosis. Female: thorax brown, scutellum yel- Ceratolophus albicornis (Meigen); Kieffer, 1906: 60. lowish, abdomen whitish; legs yellow, fore femur Bezzia albicornis (Meigen); Goetghebuel 1922: 58. with a narrow praeapical brown ring, distal l/3 of Ceratopo Bon pallidetarsatus Strobl, I 900: I 7 l. mid femur dark brown, basal and B e zzia p a I I ide tarsata (Strobl) ; Kieffer, 1 924 : 264. distal l/4 of hind Bezzia brevinenis Kieffer, l9l9: 122. femur dark brown or black; fore tibia with a Bezzia strobli Kieffer, 1919: 122. narrow, subbasal ring, mid and hind tibiae broadly Homobezzia atrata Macfie, 1944: 126. black on both ends; fore femur with 2-4 spines, Bezzia atrata (Macfie); Clastrier, 1962a: 112. often mid and hind femora armed with l-2 spines; Diagnosis. Legs black; tarsi whitish. Fore leg with proximal flagellomeres yellow basally; postscu- 2-4 spines. Parameres robust distally, expanded in tellum bare. Male: abdomen brown; legs similar in the middle, with lateral surfaces coarselly corru- colour to female; proximal flagellomeres almost gated in the widened area; end of parameres totally yellow, plume reddish. hyaline, formed by four small lobes, inner two of Tlpes. Two male syntypes of B. nobilifurmis them longer and divided by a deep notch. Clastrier in MNHN, Paris. Ilpes: Male holotype of C. albicornis Meigen Scandinavian specimens. SWEDEN: Sk: Norr in MNHN, Paris. One male and three female Krankesjrin, 3.01 .1975, H. Andersson leg.,2 d; syntypes of H. atrata Macfie in MNH, London. Norr Krankesjcin, Sjcistrand, sand, 3.07.1975, H. Scandinavian specimens. SWEDEN: Sk: Andersson leg. I d, 29 @ML). Klagshamnsudde, 1.06.1983, H. Andersson leg., I Distribution. B. nobiLis is the most widespread d (zML). species occurring in Holarctic and Neotropical Distribution. This is one of the most widely Regions; in Palaearctic known from Great Britain, distributed species of Palpomyiini, known from France, Belgium, the Netherlands, Germany, Central and western Africa, in Palaearctic from Austria, Hungary, Poland, Estonia, Lithuania, Algeria, Egypt, Spain, Great Britain, Belgium, Ukraine (Crimea), Russia (NET, CET, SET, WS, Germany, Serbia, Croatia, Austria, Hungary, Pol- ES, FE), Kazakhstan, Mongolia, China and Japan. and, Lithuania, Ukraine, Russia (SET, SMA, WS, For the first time reported from Sweden. ES, FE), Armenia, Azerbaijan, Turkmenistan, Note. B. nobilis is closelv related to B. taeniata Kazakhstan, Afghanistan, Mongolia and north (Haliday in Walker). China. Now recorded for the first time from Swe- den. (Haliday Note. B. albicornis belongs to a small group of Bezzia taeniala in Walker) species, males of which are very distinctive in the Ceratopogon taeniatus Haliday in Walker, 1856: 238.

tt'7 Jaroslaw Krzywiiski Ent. Tidskr. ll7 (1996)

Bezzia taeniata(Haliday in Walker); Kieffer, l9l9: I 14. Meig. et Johansennomyla Mall. - Arch. Institut Pas- teur d'Alg6rie. 4O: 225-288. Diagnosis. Female: thorax matt, brownish black; Coquiffett, D. W. 1901. New Diptera in the U. S. Natio- scutellum yellowish brown; abdomen yellowish, nal Museum. - Proc. U. S. Nat. Mus. 23: 593-618. dorsally with a narrow brown median stripe Edwards, F. W. 1926. On the British biting midges interrupted on joints of segments and with dark (Diptera, Ceratopogoniclae). - Trans. Entomol. Soc. band along each side. Legs yellow; basal 1/4 of Lond. ''14: 389-426. femora brown, distal l/4 of femora brownish Edwards, F. W. 1931. Insektfaunan inom Abisko Natio- under ledning av Yngve Sjtistedt. black; fore tibia with a wanting brown subbasal nalpark. III. Studier 7. Myggor Nematocera. K. Svensk. Vet. Skrift. brown tip, mid and hind tibiae broadly - - ring and Natursk. l8:32-36. brown or black on both ends; fore femur with 2-5 Goetghebuer, M. 1920. Ceratopogoninae de Belgique. - spines, other femora unarmed; bases of proximal Mem. Mus. R. Hist. Nat. Belg. 8: l-116. flagellomeres light brown; postscutellum covered Goetghebuer, M. 1922. Etude critique des Ceratopogon with setae. Male: legs darker than in female, hind de la collection Meigen conserv6e au Museum femur totally brownish black; flagellum black, d'Histoire Naturelle de Paris. - Bull. Soc. Entomol. plume brownish black. Belg. 4: 50-59. pour Scandinavian specimens. SWEDEN: Nb: N. Goetghebuer, M. 1923. Nouveaux mat6riaux l'6tude de la faune des Chironomides de Belgique. 2e Lulefl, S. Sunderbyn, 4.O1 .1972, I d, 29; 5.07 . note (l). Biol. lacustr. 12:.103-120. 1972, 3 2 7.01.t972, 4 Nb, Rine6, Rrir- -Annls. d, 9; 9. Goetghebuer, M. 1934b. Heleidae (Ceratopogonidae), All specimens H. An- biick,7.07.1912,2d,3 9. A. Die Imagines. - /n: Goetghebuer, M., Lenz, F. Die dersson leg. (ZML). Fliegen der Paliiearktischen Region, l3a, 133 pp. Distribution. B. taeniata is known from Ire- Stuttgart. land only. Now recorded for the first time in Swe- Hackman, W. 1980. A check list of the Finnish Diptera I. den. Nematocera and Brachycera (s. str.). - Notulae Note. B. taeniata was described by Haliday in Entomol. 60: l7-48. 1833. Catalogue of Diptera occuring Walker's ( 1856) "Insecta Britannica". The species Haliday, A. H. about Holywood in Downshire. Entomol. Mag. l: has not been found by any later author though its - I 47- | 80. name appeared in several papers of a compilatory Johannsen, O. A. 1905. Aquatic nematocerous Diptera (Kieffer, 1919 - in key; Edwards,1926; character II. - Buf l. N. Y. State Mus. 86: 76-352 + 2l pl. Goetghebuer, 1934b; Remm, 1988). Kieffer, J. J. 1901. Synopse des repr6sentants europ6ens du groupe Ceratopogon avec description de quelques espEces nouvelles. - Bull. Soc. Hist. Nat. Metz. 9: Acknowledgements 143-165. Kieffer, J. J. 1906. Diptera. Fam. Chironomidae. - ln: Danielsson, Zoolo- I am very much indebted to Dr. R. Wytsman, P Genera Insectorum. 42:78 pp. gical Museum, University of Lund who kindly arranged Kieffer, J. J. 1913. Nouveaux Chironomides (Ten- the loan of undetermined specimens of Swedish Pal- dip6dides) d'Allemagne. - Bull. Soc. Hist. Nat. Metz. very grateful Dr. P. Grootaert of pomyiini. I am also to 28:'1-35. the Institut Royal des Sciences Naturelles de Belgique. Kieffer. J. J. 1915. Uber diinische Chironomiden. - Brussels (lRScNB), Dr. B. Pitkin of the Natural History Entomol. Meddel. lO: 28O-29'7 . Museum, London (MHN) and Dr. L. Matile of the Mu- Kieffer, J. J. 1916. Schwedische Chironomiden. - Arch. Paris (MNHN) for seum National d'Histoire Naturelle, Hydrobiol. Suppl. 2: 483-554. their help given to me during my study of Palpomyiini. Kieffer, J. J. I 9 I 9. Chironom ides d' Europe conserves au My special thanks are due to S.-A. Bergtind and L. Hed- Mus6e National Hongrois de Budapest. Ann. Mus. translations. str6m for the Swedish nat. Hung. l7: l-160. Kieffer, l. J. 1924. Chrionomides Isic!] d'Egypte (Dipt.). - Bull. Soc. R. Ent. Egypte. 8:244-313. References Kieffer, l. J. 1925a. Diptdres (ndmatoceres piqueurs): Clastrier, J.1962a. Notes sur les Cdratopogonid6s. XVI. Chironomidae Ceratopogoninae. Faune de France. Espbces nouvelles de la R6gion pal6arctique, ou 11, 139 pp. Paris. apparent6es, du genre Bezzia Kieffer. - Arch. Institut Kieffer J. J., 1925b, Nouveaux genres et nouvelles Pasteur d'Alg6rie. 40: 53- I 25. espdces de Chironomides piqueurs, Arch. Inst. Pas- Clastrier, J. 1962b. Notes sur Cdratopogonid6s de la teur A196r., 3: 405-430. R6gion pal6arctique. XVII. Nouveaux Palpomyia Loew, H. 1861. Diptera Americae septentrionalis

I t8 Ent. Tidskr. 117 (1996) S o me S c andinav ian P alpo my i ini indigena. Centuria prima. - Berl. Entomol. Z. 5: Strobl, G. 1900. Spanische Dipteren. XI Theil. - Wien. 307 -359. Ent. Zeit. 19: 169-l'74. Lundstr

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