DOCSLIB.ORG

Non-Anguimorph Lizards of the Late Oligocene and Early Miocene Of

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Non-Anguimorph Lizards of the Late Oligocene and Early Miocene Of East Tennessee State University Digital Commons @ East Tennessee State University Electronic Theses and Dissertations Student Works 5-2014 Non-anguimorph Lizards of the Late Oligocene and Early Miocene of Florida and Implications for the Reorganization of the North American Herpetofauna Kevin Chovanec East Tennessee State University Follow this and additional works at: https://dc.etsu.edu/etd Part of the Biodiversity Commons, Ecology and Evolutionary Biology Commons, and the Paleontology Commons Recommended Citation Chovanec, Kevin, "Non-anguimorph Lizards of the Late Oligocene and Early Miocene of Florida and Implications for the Reorganization of the North American Herpetofauna" (2014). Electronic Theses and Dissertations. Paper 2384. https://dc.etsu.edu/ etd/2384 This Thesis - Open Access is brought to you for free and open access by the Student Works at Digital Commons @ East Tennessee State University. It has been accepted for inclusion in Electronic Theses and Dissertations by an authorized administrator of Digital Commons @ East Tennessee State University. For more information, please contact [email protected]. Non-anguimorph Lizards of the Late Oligocene and Early Miocene of Florida and Implications for the Reorganization of the North American Herpetofauna ________________________ A thesis presented to the faculty of the Department of Geosciences East Tennessee State University In partial fulfillment of the requirements for the degree Master of Science in Geosciences ________________________ by Kevin Robert Chovanec May 2014 ________________________ Blaine W. Schubert, Chair Jim I. Mead Steven C. Wallace Keywords: biogeography, Squamata, provincialism, Neotropical, niche conservatism 1 ABSTRACT Non-anguimorph Lizards of the Late Oligocene and Early Miocene of Florida and Implications for the Reorganization of the North American Herpetofauna by Kevin R. Chovanec Paleokarst deposits from the Oligo-Miocene of northern Florida preserve undescribed herpetofaunal remains that fill important temporal and geographic gaps in our understanding of Cenozoic lizard evolution. Here I describe and discuss the non- anguimorph lizard diversity of the Brooksville 2 (Ar2) and Miller (He1) local faunas to test for patterns of regional and latitudinal provincialism in the contemporary North American record. Collectively, the sites are significant for documenting 1) extralimital occurrences of the tropical clades Anolis and Corytophaninae, 2) a substantial temporal range extension of the modern southeastern endemic Rhineuridae, 3) the earliest record of eublepharid gekkotans from North America, and 4) the early Miocene arrival of “cnemidophorine” teiids from South America. This work complements recent studies of older, Eocene lizards by others and lends paleontological support to aspects of the tropical conservatism hypothesis: lineages now confined to the tropics were present at higher latitudes when megathermal climates were more extensive. 2 ACKNOWLEDGEMENTS Thank you to Drs. Blaine Schubert, Jim Mead, and Wally “Steven” Wallace for their guidance, patience, and support. Fellow graduate students helped shape this endeavor into the experience that it has been. Jeff Martin, in particular, has been a dependable source of useful and not-so-useful discussion. He also fixed my car and helped make significant improvements to my oral presentation; I feel compelled to thank him on behalf of my audience for the latter. Mark Hutchinson provided some encouraging words at the start of my program that were (and still are) much appreciated. Richard Hulbert and Kenneth Krysko of the Florida Museum of Natural History facilitated access to fossil and modern specimens under their care, and Andreas Kerner kindly donated material from the Miller locality. Not a single aspect of this project would have been possible without the help of Sandy Swift. Sandy showed me the art of picking and sorting, taught me much of what I know of small vertebrate anatomy, and tried her best to get me to produce the same kinds of high-quality figures that she does. My questions were always met with enthusiasm, and her curiosity has been infectious. She filled equally important roles of mentor and total distraction, so thanks to her for being a source of chocolate and of motivation. 3 TABLE OF CONTENTS Page ABSTRACT .........................................................................................................................2 ACKNOWLEDGEMENTS .................................................................................................3 TABLE OF CONTENTS .....................................................................................................4 TABLE OF FIGURES .........................................................................................................9 Chapter 1. INTRODUCTION .........................................................................................................11 2. NON-ANGUIMORPH LIZARDS OF THE BROOKSVILLE 2 LOCAL FAUNA .....18 Introduction ................................................................................................................... 18 Anolis sp. ....................................................................................................................... 19 Referred Material........................................................................................................19 Description .................................................................................................................19 Dentary. ................................................................................................................. 19 Maxilla. ................................................................................................................. 23 Braincase. .............................................................................................................. 26 Frontal. .................................................................................................................. 30 Prefrontal............................................................................................................... 32 Postorbital. ............................................................................................................ 33 Quadrate. ............................................................................................................... 34 Remarks ......................................................................................................................35 cf. Anolis sp. .................................................................................................................. 44 Referred Material........................................................................................................44 Description .................................................................................................................44 4 Dentary. ................................................................................................................. 44 Remarks ......................................................................................................................47 Iguanidae gen. et sp. indet. ........................................................................................... 48 Referred Material........................................................................................................48 Description .................................................................................................................48 Dentary. ................................................................................................................. 48 Remarks ......................................................................................................................51 Iguaninae gen. et sp. indet. ........................................................................................... 52 Referred Material........................................................................................................52 Description .................................................................................................................52 Dentary. ................................................................................................................. 52 Remarks ......................................................................................................................54 Eublepharidae gen. et sp. indet. .................................................................................... 57 Referred Material........................................................................................................57 Description .................................................................................................................57 Maxilla. ................................................................................................................. 57 Dentary. ................................................................................................................. 62 Remarks ......................................................................................................................62 Rhineuridae gen. et sp. indet. ........................................................................................ 64 Referred Material........................................................................................................64 Description .................................................................................................................64 Dentary. ................................................................................................................
Load more

Recommended publications
  • Other Contributions
    Other Contributions NATURE NOTES Amphibia: Caudata Ambystoma ordinarium. Predation by a Black-necked Gartersnake (Thamnophis cyrtopsis). The Michoacán Stream Salamander (Ambystoma ordinarium) is a facultatively paedomorphic ambystomatid species. Paedomorphic adults and larvae are found in montane streams, while metamorphic adults are terrestrial, remaining near natal streams (Ruiz-Martínez et al., 2014). Streams inhabited by this species are immersed in pine, pine-oak, and fir for- ests in the central part of the Trans-Mexican Volcanic Belt (Luna-Vega et al., 2007). All known localities where A. ordinarium has been recorded are situated between the vicinity of Lake Patzcuaro in the north-central portion of the state of Michoacan and Tianguistenco in the western part of the state of México (Ruiz-Martínez et al., 2014). This species is considered Endangered by the IUCN (IUCN, 2015), is protected by the government of Mexico, under the category Pr (special protection) (AmphibiaWeb; accessed 1April 2016), and Wilson et al. (2013) scored it at the upper end of the medium vulnerability level. Data available on the life history and biology of A. ordinarium is restricted to the species description (Taylor, 1940), distribution (Shaffer, 1984; Anderson and Worthington, 1971), diet composition (Alvarado-Díaz et al., 2002), phylogeny (Weisrock et al., 2006) and the effect of habitat quality on diet diversity (Ruiz-Martínez et al., 2014). We did not find predation records on this species in the literature, and in this note we present information on a predation attack on an adult neotenic A. ordinarium by a Thamnophis cyrtopsis. On 13 July 2010 at 1300 h, while conducting an ecological study of A.
    [Show full text]
  • Xenosaurus Tzacualtipantecus. the Zacualtipán Knob-Scaled Lizard Is Endemic to the Sierra Madre Oriental of Eastern Mexico
    Xenosaurus tzacualtipantecus. The Zacualtipán knob-scaled lizard is endemic to the Sierra Madre Oriental of eastern Mexico. This medium-large lizard (female holotype measures 188 mm in total length) is known only from the vicinity of the type locality in eastern Hidalgo, at an elevation of 1,900 m in pine-oak forest, and a nearby locality at 2,000 m in northern Veracruz (Woolrich- Piña and Smith 2012). Xenosaurus tzacualtipantecus is thought to belong to the northern clade of the genus, which also contains X. newmanorum and X. platyceps (Bhullar 2011). As with its congeners, X. tzacualtipantecus is an inhabitant of crevices in limestone rocks. This species consumes beetles and lepidopteran larvae and gives birth to living young. The habitat of this lizard in the vicinity of the type locality is being deforested, and people in nearby towns have created an open garbage dump in this area. We determined its EVS as 17, in the middle of the high vulnerability category (see text for explanation), and its status by the IUCN and SEMAR- NAT presently are undetermined. This newly described endemic species is one of nine known species in the monogeneric family Xenosauridae, which is endemic to northern Mesoamerica (Mexico from Tamaulipas to Chiapas and into the montane portions of Alta Verapaz, Guatemala). All but one of these nine species is endemic to Mexico. Photo by Christian Berriozabal-Islas. amphibian-reptile-conservation.org 01 June 2013 | Volume 7 | Number 1 | e61 Copyright: © 2013 Wilson et al. This is an open-access article distributed under the terms of the Creative Com- mons Attribution–NonCommercial–NoDerivs 3.0 Unported License, which permits unrestricted use for non-com- Amphibian & Reptile Conservation 7(1): 1–47.
    [Show full text]
  • Of Vertebrate Fossils from the Middle Eocene Oil Shale of Messel, Germany: Implications for Their Taphonomy and Palaeoenvironment
    Palaeogeography, Palaeoclimatology, Palaeoecology 416 (2014) 92–109 Contents lists available at ScienceDirect Palaeogeography, Palaeoclimatology, Palaeoecology journal homepage: www.elsevier.com/locate/palaeo Isotope compositions (C, O, Sr, Nd) of vertebrate fossils from the Middle Eocene oil shale of Messel, Germany: Implications for their taphonomy and palaeoenvironment Thomas Tütken ⁎ Steinmann-Institut für Geologie, Mineralogie und Paläontologie, Universität Bonn, Poppelsdorfer Schloss, 53115 Bonn, Germany article info abstract Article history: The Middle Eocene oil shale deposits of Messel are famous for their exceptionally well-preserved, articulated 47- Received 15 April 2014 Myr-old vertebrate fossils that often still display soft tissue preservation. The isotopic compositions (O, C, Sr, Nd) Received in revised form 30 July 2014 were analysed from skeletal remains of Messel's terrestrial and aquatic vertebrates to determine the condition of Accepted 5 August 2014 geochemical preservation. Authigenic phosphate minerals and siderite were also analysed to characterise the iso- Available online 17 August 2014 tope compositions of diagenetic phases. In Messel, diagenetic end member values of the volcanically-influenced 12 Keywords: and (due to methanogenesis) C-depleted anoxic bottom water of the meromictic Eocene maar lake are isoto- Strontium isotopes pically very distinct from in vivo bioapatite values of terrestrial vertebrates. This unique taphonomic setting al- Oxygen isotopes lows the assessment of the geochemical preservation of the vertebrate fossils. A combined multi-isotope Diagenesis approach demonstrates that enamel of fossil vertebrates from Messel is geochemically exceptionally well- Enamel preserved and still contains near-in vivo C, O, Sr and possibly even Nd isotope compositions while bone and den- Messel tine are diagenetically altered.
    [Show full text]
  • The World at the Time of Messel: Conference Volume
    T. Lehmann & S.F.K. Schaal (eds) The World at the Time of Messel - Conference Volume Time at the The World The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment and the History of Early Primates 22nd International Senckenberg Conference 2011 Frankfurt am Main, 15th - 19th November 2011 ISBN 978-3-929907-86-5 Conference Volume SENCKENBERG Gesellschaft für Naturforschung THOMAS LEHMANN & STEPHAN F.K. SCHAAL (eds) The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment, and the History of Early Primates 22nd International Senckenberg Conference Frankfurt am Main, 15th – 19th November 2011 Conference Volume Senckenberg Gesellschaft für Naturforschung IMPRINT The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment, and the History of Early Primates 22nd International Senckenberg Conference 15th – 19th November 2011, Frankfurt am Main, Germany Conference Volume Publisher PROF. DR. DR. H.C. VOLKER MOSBRUGGER Senckenberg Gesellschaft für Naturforschung Senckenberganlage 25, 60325 Frankfurt am Main, Germany Editors DR. THOMAS LEHMANN & DR. STEPHAN F.K. SCHAAL Senckenberg Research Institute and Natural History Museum Frankfurt Senckenberganlage 25, 60325 Frankfurt am Main, Germany [email protected]; [email protected] Language editors JOSEPH E.B. HOGAN & DR. KRISTER T. SMITH Layout JULIANE EBERHARDT & ANIKA VOGEL Cover Illustration EVELINE JUNQUEIRA Print Rhein-Main-Geschäftsdrucke, Hofheim-Wallau, Germany Citation LEHMANN, T. & SCHAAL, S.F.K. (eds) (2011). The World at the Time of Messel: Puzzles in Palaeobiology, Palaeoenvironment, and the History of Early Primates. 22nd International Senckenberg Conference. 15th – 19th November 2011, Frankfurt am Main. Conference Volume. Senckenberg Gesellschaft für Naturforschung, Frankfurt am Main. pp. 203.
    [Show full text]
  • REPTILIA: SQUAMATA: SCINCIDAE Eumeces Lagunensis Van Denburgh
    792.1 REPTILIA: SQUAMATA: SCINCIDAE EUMECES LAGUNENSIS Catalogue of American Amphibians and Reptiles. Beaman, K.R., J.Q. Richmond, and L.L. Grismer. 2004. Eumeces lagunensis. Eumeces lagunensis Van Denburgh San Lucan Skink Eumeces skiltonianus: Yarrow 1882:41 (part). Eumeces lagunensis Van Denburgh 1895:134. Type locality, “San Francisquito, Sierra Laguna, [Baja California Sur, México].” Holotype, California Academy of Sciences (CAS) 400, collected by Gustav Eisen on 28 March 1892 (examined by LLG). See Remarks. Plestiodon lagunensis: Van Denburgh and Slevin 1921:52. Plestiodon skiltonianus lagunensis: Nelson 1921:114–115. Eumeces skiltonianus lagunensis: Linsdale 1932:374. • CONTENT. The species is monotypic. • DEFINITION. Eumeces lagunensis is a small skink with a maximum total length of 147 mm. The scutellation is as fol- lows: 24 scale rows at midbody; 57–60 dorsal scale rows; 40– 46 ventral scale rows; 102 subcaudals; 4 supraoculars (three touching frontal); frontonasal in contact with frontal or not; large interparietal enclosed posteriorly by medial contact of large parietals; 7–8 supralabials; upper secondary temporal in broad 0 100 200 km contact ventrally with last supralabial; 2 postmentals; 6 infralabials; 2 postlabials (not superimposed); 2–2 nuchals, oc- casionally 1–1, 1–2, or 3–3, blending posteriorly with wide, MAP. Range of Eumeces lagunensis, the white circle marks the type cycloid, imbricate, dorsal scales of body and tail; 16 scales locality, the gray circle marks the neotype locality, and dots indicate around base of tail; and vent bordered by two large scales ante- other records. riorly. Granular axillary scales are not prominent and only 0–2 short rows are present and situated posterior to the medial mar- gin of the forelimb insertion.
    [Show full text]
  • The Sclerotic Ring: Evolutionary Trends in Squamates
    The sclerotic ring: Evolutionary trends in squamates by Jade Atkins A Thesis Submitted to Saint Mary’s University, Halifax, Nova Scotia in Partial Fulfillment of the Requirements for the Degree of Master of Science in Applied Science July, 2014, Halifax Nova Scotia © Jade Atkins, 2014 Approved: Dr. Tamara Franz-Odendaal Supervisor Approved: Dr. Matthew Vickaryous External Examiner Approved: Dr. Tim Fedak Supervisory Committee Member Approved: Dr. Ron Russell Supervisory Committee Member Submitted: July 30, 2014 Dedication This thesis is dedicated to my family, friends, and mentors who helped me get to where I am today. Thank you. ! ii Table of Contents Title page ........................................................................................................................ i Dedication ...................................................................................................................... ii List of figures ................................................................................................................. v List of tables ................................................................................................................ vii Abstract .......................................................................................................................... x List of abbreviations and definitions ............................................................................ xi Acknowledgements ....................................................................................................
    [Show full text]
  • Multi-National Conservation of Alligator Lizards
    MULTI-NATIONAL CONSERVATION OF ALLIGATOR LIZARDS: APPLIED SOCIOECOLOGICAL LESSONS FROM A FLAGSHIP GROUP by ADAM G. CLAUSE (Under the Direction of John Maerz) ABSTRACT The Anthropocene is defined by unprecedented human influence on the biosphere. Integrative conservation recognizes this inextricable coupling of human and natural systems, and mobilizes multiple epistemologies to seek equitable, enduring solutions to complex socioecological issues. Although a central motivation of global conservation practice is to protect at-risk species, such organisms may be the subject of competing social perspectives that can impede robust interventions. Furthermore, imperiled species are often chronically understudied, which prevents the immediate application of data-driven quantitative modeling approaches in conservation decision making. Instead, real-world management goals are regularly prioritized on the basis of expert opinion. Here, I explore how an organismal natural history perspective, when grounded in a critique of established human judgements, can help resolve socioecological conflicts and contextualize perceived threats related to threatened species conservation and policy development. To achieve this, I leverage a multi-national system anchored by a diverse, enigmatic, and often endangered New World clade: alligator lizards. Using a threat analysis and status assessment, I show that one recent petition to list a California alligator lizard, Elgaria panamintina, under the US Endangered Species Act often contradicts the best available science.
    [Show full text]
  • Cretaceous Fossil Gecko Hand Reveals a Strikingly Modern Scansorial Morphology: Qualitative and Biometric Analysis of an Amber-Preserved Lizard Hand
    Cretaceous Research 84 (2018) 120e133 Contents lists available at ScienceDirect Cretaceous Research journal homepage: www.elsevier.com/locate/CretRes Cretaceous fossil gecko hand reveals a strikingly modern scansorial morphology: Qualitative and biometric analysis of an amber-preserved lizard hand * Gabriela Fontanarrosa a, Juan D. Daza b, Virginia Abdala a, c, a Instituto de Biodiversidad Neotropical, CONICET, Facultad de Ciencias Naturales e Instituto Miguel Lillo, Universidad Nacional de Tucuman, Argentina b Department of Biological Sciences, Sam Houston State University, 1900 Avenue I, Lee Drain Building Suite 300, Huntsville, TX 77341, USA c Catedra de Biología General, Facultad de Ciencias Naturales, Universidad Nacional de Tucuman, Argentina article info abstract Article history: Gekkota (geckos and pygopodids) is a clade thought to have originated in the Early Cretaceous and that Received 16 May 2017 today exhibits one of the most remarkable scansorial capabilities among lizards. Little information is Received in revised form available regarding the origin of scansoriality, which subsequently became widespread and diverse in 15 September 2017 terms of ecomorphology in this clade. An undescribed amber fossil (MCZ Re190835) from mid- Accepted in revised form 2 November 2017 Cretaceous outcrops of the north of Myanmar dated at 99 Ma, previously assigned to stem Gekkota, Available online 14 November 2017 preserves carpal, metacarpal and phalangeal bones, as well as supplementary climbing structures, such as adhesive pads and paraphalangeal elements. This fossil documents the presence of highly specialized Keywords: Squamata paleobiology adaptive structures. Here, we analyze in detail the manus of the putative stem Gekkota. We use Paraphalanges morphological comparisons in the context of extant squamates, to produce a detailed descriptive analysis Hand evolution and a linear discriminant analysis (LDA) based on 32 skeletal variables of the manus.
    [Show full text]
  • U.S. Fish & Wildlife Service June 14, 2016 Biological Opinion Revised
    U.S. Fish & Wildlife Service June 14, 2016 Biological Opinion ON Revised Land and Resource Management Plan Amendment to increase Florida Scrub- Jay Management Areas on the Ocala National Forest (Amendment 12) Prepared by: U.S. Fish and Wildlife Service Jacksonville, Florida Biological Opinion U.S. Forest Service Southern Region FWS Log No. 04EF1000-2016-F-0215 2 The Service concurs with your determination that the effects from activities under the proposed amendment on the Florida bonamia, scrub buckwheat, and Lewton’s polygala are within the scope of effects described in the September 18, 1998 BA for the LRMP and evaluated in the Service’s 1998 Opinion. In addition, effects of implementing the LRMP (including the proposed amendment) on the scrub pigeon wings were recently disclosed in your Biological Assessment (BA) of Nov 24, 2015 were evaluated in the Service’s Opinion of December 17, 2015. Therefore, these plant species will not be addressed further in the amended Opinion below. This amended Opinion is based on information provided to the Service through a BA, telephone conversations, e-mails, field investigation notes, and other sources of information. A complete administrative record of this consultation is on file at the Jacksonville Ecological Services Office. Consultation History September 21, 1998: NFF initiated formal consultation on revision of the LRMP December 18, 1998: The Service provided a non-jeopardy combined Biological and Conference Opinion on the LRMP to NFF concluding formal consultation. From March 2014 to November of 2015, the Service and staff from the NFF supervisor’s office and ONF participated in several meetings and conference calls to discuss how to address Forest Service Section 7(a)(1) obligations under the Act and the proposed amendment to the NFF LRMP.
    [Show full text]
  • Checklist Reptile and Amphibian
    To report sightings, contact: Natural Resources Coordinator 980-314-1119 www.parkandrec.com REPTILE AND AMPHIBIAN CHECKLIST Mecklenburg County, NC: 66 species Mole Salamanders ☐ Pickerel Frog ☐ Ground Skink (Scincella lateralis) ☐ Spotted Salamander (Rana (Lithobates) palustris) Whiptails (Ambystoma maculatum) ☐ Southern Leopard Frog ☐ Six-lined Racerunner ☐ Marbled Salamander (Rana (Lithobates) sphenocephala (Aspidoscelis sexlineata) (Ambystoma opacum) (sphenocephalus)) Nonvenomous Snakes Lungless Salamanders Snapping Turtles ☐ Eastern Worm Snake ☐ Dusky Salamander (Desmognathus fuscus) ☐ Common Snapping Turtle (Carphophis amoenus) ☐ Southern Two-lined Salamander (Chelydra serpentina) ☐ Scarlet Snake1 (Cemophora coccinea) (Eurycea cirrigera) Box and Water Turtles ☐ Black Racer (Coluber constrictor) ☐ Three-lined Salamander ☐ Northern Painted Turtle ☐ Ring-necked Snake (Eurycea guttolineata) (Chrysemys picta) (Diadophis punctatus) ☐ Spring Salamander ☐ Spotted Turtle2, 6 (Clemmys guttata) ☐ Corn Snake (Pantherophis guttatus) (Gyrinophilus porphyriticus) ☐ River Cooter (Pseudemys concinna) ☐ Rat Snake (Pantherophis alleghaniensis) ☐ Slimy Salamander (Plethodon glutinosus) ☐ Eastern Box Turtle (Terrapene carolina) ☐ Eastern Hognose Snake ☐ Mud Salamander (Pseudotriton montanus) ☐ Yellow-bellied Slider (Trachemys scripta) (Heterodon platirhinos) ☐ Red Salamander (Pseudotriton ruber) ☐ Red-eared Slider3 ☐ Mole Kingsnake Newts (Trachemys scripta elegans) (Lampropeltis calligaster) ☐ Red-spotted Newt Mud and Musk Turtles ☐ Eastern Kingsnake
    [Show full text]
  • Eumeces Gilberti Van Denburgh Gilbert's Skink
    372.1 REPTILIA: SQUAMATA: SAURIA: SCINCIDAE EUMECES GILBERTI Catalogue of American Amphibians and Reptiles. Rodgers (1944) describes E. g. placerensis, and Lowe and Shannon (1954) E. g. arizonensis. Stebbins (1966) and Behler and King JONES,K. BRUCE. 1985. Eumeces gilberti. (1979) provide brief descriptions of the species. Eumeces gilberti Van Denburgh • ILLUSTRATIONS.Stebbins (1966) and Behler and King (1979) Gilbert's Skink provide color illustrations and color photographs of juveniles and adults, respectively. Black and white photographs appear in Van Denburgh (1922), Taylor (1935), and Smith (1946). Rodgers (1944) Eumeces gilberti: Van Denburgh, 1896:350. Type.locality, "Yo· provides a photograph of the type.specimen E. g. placerensis. Van semite Valley, Mariposa County, California." Holotype, Cali• Denburgh (1922), Taylor (1935), Smith (1946), and Rodgers and fornia Acad. Sci.-Stanford Univ. 4139, collected by Charles Fitch (1947) provide black and white illustrations with the latter H. Gilbert and James M. Hyde on 10-15 June 1896 (not the most detailed. examined by author). Eumeces skiltonianus: Cope, 1900:643 (part, by inference). • DISTRIBUTION.The species is distributed through central Cal• Eumeces skiltonianus: Camp, 1916:72-73 (part). ifornia, north approximately to the Yuba River, east through the Plestiodon skiltonianum: Grinnell and Camp, 1917:175, 176 (part). San Joaquin Valley to the Sierra Nevada, and west to the San Eumeces gilberti: Taylor, 1935:438. Resurrected name. Francisco Bay area. Its range extends southward along the Califor· nia coast (but at least 20 km inland) to San Diego, and into the • CONTENT.Five subspecies are recognized: gilberti, cancel• chaparral vegetation association of the San Pedro Martir of Baja losus, placerensis, rubricaudatus, and arizonensis.
    [Show full text]
  • Analysis of Diet Composition and Morphological Characters of The
    Official journal website: Amphibian & Reptile Conservation amphibian-reptile-conservation.org 13(1) [General Section]: 111–121 (e172). Analysis of diet composition and morphological characters of the little-known Peruvian bush anole Polychrus peruvianus (Noble, 1924) in a northern Peruvian dry forest 1Antonia Beuttner and 2,*Claudia Koch 1Universität Tübingen, Geschwister-Scholl-Platz, 72074 Tübingen, GERMANY 2Zoologisches Forschungsmuseum Alexander Koenig (ZFMK), Adenauerallee 160, 53113 Bonn, GERMANY Abstract.—Analyses of diet composition are an important element of studies focusing on biological diversity, ecology, and behavior of animals. Polychrus peruvianus was found to be a quite abundant lizard species in the northern Peruvian dry forest. However, our knowledge of the ecology of this species remains limited. Herein, we analyze the species dietary composition and the morphological features that may be related to the feeding behavior. Our results show that P. peruvianus is a semi-herbivorous food generalist, which also consumes faunistic prey. All age groups prefer mobile prey as sit-and-wait predators. However, during ontogenesis, plant material becomes the main component in the diet of adult specimens. Keywords. Iguania, bite force, ontogenetic change, foraging strategy, stomach contents, food niche, feeding behavior, Marañón river Citation: Beuttner A, Koch C. 2019. Analysis of diet composition and morphological characters of the little-known Peruvian bush anole Polychrus peruvianus (Noble, 1924) in a northern Peruvian dry forest. Amphibian & Reptile Conservation 13(1) [General Section]: 111–121 (e172). Copyright: © 2019 Beuttner and Koch. This is an open access article distributed under the terms of the Creative Commons Attribution License [At- tribution 4.0 International (CC BY 4.0): https://creativecommons.org/licenses/by/4.0/], which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
    [Show full text]