Ringing & Migration

VOLUME 24 2008 and 2009

Editor Chris Redfern

Managing Editor Jacquie Clark

Copy Editor John Marchant

Editorial Board

2008 & 2009

Guy Anderson, Franz Bairlein, Jon Coleman, Jennifer Gill, Hugh Insley, Åke Lindström, Stephen Norman, Jane Reid, Roger Riddington and Graham Scott

ISSN 0307-8698 Vol 24 Contents

Ringing & Migration Contents of Vol 24, 2008 and 2009

Part 1 (June 2008)

FULL PAPER Survival rates of hirundines in relation to British and African rainfall. Robert A. Robinson, Dawn E. Balmer and John H. Marchant...... 1

SHORT REPORTS From fledging to breeding: long-term satellite tracking of the migratory behaviour of a Lesser Black-backed Gull Larus fuscus intermedius. K. Pütz, A.J. Helbig, K.T. Pedersen, C. Rahbek, P. Saurola and R. Juvaste...... 7 On reading colour rings. Carl Mitchell and Mark Trinder...... 11

RINGING SCHEME REPORT ringing in Britain and Ireland in 2006. Liz Coiffait, Jacquie A. Clark, Robert A. Robinson, Jeremy R. Blackburn, Bridget M. Griffin, Kate Risely, Mark J. Grantham, John H. Marchant, Trevor Girling and Lee Barber...... 15

Part 2 (December 2008)

FULL PAPERS Sexual size dimorphism and moult in the Plain Swift Apus unicolor. Eduardo Garcia-Del-Rey, Andrew G. Gosler, Javier Gonzalez and Michael Wink...... 81 Ageing and sexing Marsh Tits Poecile palustris using wing length and moult. Richard K. Broughton, Shelley A. Hinsley, Paul E. Bellamy, Jane E. Carpenter and Peter Rothery...... 88 How far do fly during one migratory flight stage? K. Susanna S. Hall-Karlsson and Thord Fransson...... 95

SHORT REPORT Separation of Marsh Tit Poecile palustris from Willow Tit Poecile montana using a bill criterion. Richard K. Broughton, Shelley A. Hinsley and Paul E. Bellamy...... 101

RINGING SCHEME REPORT Bird ringing in Britain and Ireland in 2007. Liz Coiffait, Jacquie A. Clark, Robert A. Robinson, Jeremy R. Blackburn, Mark J. Grantham, John H. Marchant, Lee Barber, Diana De Palacio, Bridget M. Griffin and Dorian Moss...... 104

Part 3 (June 2009) SPECIAL CENTENARY ISSUE

EDITORIAL A century of ringing in Britain and Ireland. Chris P.F. Redfern, Jacquie A. Clark, Robert A. Robinson and Stephen R. Baillie...... 145

THE SCIENCE OF RINGING: DEVELOPMENT AND FUTURE 100 years of ringing in Britain and Ireland. Jeremy J.D. Greenwood...... 147 From individuals to flyways: the future of marking birds for conservation. Stephen R. Baillie, Robert A. Robinson, Jacquie A. Clark and Chris P.F. Redfern...... 155

MIGRATION AND MOVEMENTS: FOLLOWING THE BIRDS Ringing and the study of mechanisms of migration. Franz Bairlein and Michael Schaub...... 162 The use of intrinsic markers to study bird migration. Lisette Coiffait, Chris P.F. Redfern, Richard M. Bevan, Jason Newton and Kirsten Wolff...... 169 New technologies for monitoring bird migration and behaviour. Wolfgang Fiedler...... 175 Quantifying the movement patterns of birds from ring recoveries. Jacquie A. Clark, Kasper Thorup and David A. Stroud...... 180 Vol 24 Contents

AVIAN DEMOGRAPHY: QUANTIFYING THE UPS AND DOWNS Understanding changes in bird populations – the role of bird marking. Stephen R. Baillie and Michael Schaub...... 189 Constant effort: studying avian population processes using standardised ringing. Robert A. Robinson, Romain Julliard and James F. Saracco...... 199 The value of ringing for bird conservation. Guy Q.A. Anderson and Rhys E. Green ...... 205

BEHAVIOUR, LIFE-HISTORY AND CONDITION: THE STATE OF THE INDIVIDUAL Bird ringing as a tool for behavioural studies. Stuart P. Sharp...... 213 Moult and plumage. Ian Newton...... 220 The use of mass and fat reserve measurements from ringing studies to assess body condition. Will Cresswell...... 227

Part 4 (December 2009)

FULL PAPERS Sexing Black-legged Kittiwakes by measurement. J.C. Coulson...... 233 Ageing and sexing the Yellowhammer Emberiza citrinella caliginosa during the non-breeding season. Jenny C. Dunn and Chris Wright...... 240 Counts of spring passage Golden Plover Pluvialis apricaria in north Lewis. S. Rae, M.G.W. Jones, F. Stewart and R. Tingay...... 253 Migration of Mallards Anas platyrhynchos in Hungary: migration phenology, the origin of migrants, and long-term changes. András Liker and Lajos Nagy...... 259 Declining rates of ring recovery in British birds. Robert A. Robinson, Mark J. Grantham and Jacquie A. Clark...... 266

SHORT REPORTS Comparing the migration of Eurasian Teal Anas crecca from two main wintering areas of Western Europe: a long-term study from Essex, England, and the Camargue, France. Matthieu Guillemain, Richard Hearn, Roy King, Michel Gauthier-Clerc, Geraldine Simon and Alain Caizergues...... 273 Habitat, phenology and biometrics of the Aquatic Warbler Acrocephalus paludicola during autumn migration through a riverine wetland in Iberia. David Miguélez, Carlos Zumalacárregui, Benito Fuertes, Héctor Astiárraga, Rubén González-Jáñez, Isabel Roa and Fran de la Calzada...... 277 Ageing Black-headed Gulls on leg and bill colour – a cautionary note. Robin M. Sellers...... 280

RINGING REPORT Bird ringing in Britain and Ireland in 2008. Liz Coiffait, Jacquie A. Clark, Robert A. Robinson, Jeremy R. Blackburn, Mark J. Grantham, Dave I. Leech, John H. Marchant, Lee J. Barber, Diana De Palacio, Bridget M. Griffin, Dorian Moss and Miranda M. Shephard...... 281 Vol 24 Index INDEX TO VOLUME 24 Compiled by Hugh Insley

All species of bird mentioned in the text are included except those mentioned only in the 2006, 2007 and 2008 Ringing Reports. Page numbers given are those of the first mention of the species in each paper, and exclude the Ringing Reports.

Accipiter nisus 162, 222, 228, 271 Acrocephalus arundinaceus 97, 171, 221 A. paludicola 170, 277 A. palustris 97 A. schoenobaenus 95, 107, 156, 201, 216, 272, 283 A. scirpaceus 95, 107, 170, 201, 272, 283 Actitis hypoleucos 285 A. macularia 221 Aegithalos caudatus 107, 216, 281 Age determination 88 Alca torda 271 Alpine Swift 221 American Golden Plover 257 American Mink 282 American Redstart 171 Anas crecca 162, 183, 271, 273 A. formosa 170 A. penelope 149, 271 A. platyrhynchos 163, 183, 194, 259, 271 ANDERSON, GUY Q.A. and RHYS E. GREEN The value of ringing for bird conservation 205 Anguilla anguilla 208 Anser albifrons 182, 259 A. anser 271 A. brachyrhynchus 191, 271 A. fabalis 259 Anthus trivialis 97, 156 californica 214 Apus apus 81, 221, 272 A. melba 221 A. unicolor 81 Aquatic Warbler 170, 277 ASTIÁRRAGA, HÉCTOR see DAVID MIGUÉLEZ et al Atlantic Puffin 237, 271 Avian Influenza 16, 105, 156, 185, 291 Aythya ferina 105, 271 A. fuligula 105, 271

Baikal Teal 170 BAILLIE, STEPHEN R. see CHRIS P.F. REDFERN et al 145 BAILLIE, STEPHEN R., ROBERT A. ROBINSON, JACQUIE A. CLARK and CHRIS P.F. REDFERN From individuals to flyways: the future of marking birds for conservation 155 BAILLIE, STEPHEN R. and MICHAEL SCHAUB Understanding changes in bird populations – the role of bird marking 189 BAIRLEIN, FRANZ and MICHAEL SCHAUB Ringing and the study of mechanisms of migration 162 Bald Eagle 9 BALMER, DAWN E. see ROBERT A. ROBINSON et al BARBER, LEE see LIZ COIFFAIT et al 15, 104, 281 Barn Owl 269 Barn Swallow 1, 97, 162, 170, 221, 272 Barred Warbler 97, 221 Bar-tailed Godwit 176 Bean Goose 259

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Bee-eater 221 BELLAMY, PAUL E. see RICHARD K. BROUGHTON et al 88, 101 BEVAN, RICHARD M. see LISETTE COIFFAIT et al 169 Bill colour 101, 280; — depth 83; — length 81 Bittern 205 Blackbird 107, 150, 227, 272, 281 BLACKBURN, JEREMY R. see LIZ COIFFAIT et al 15, 104, 281 Blackcap 97, 107, 170, 184, 216, 272, 281 Black-capped Chickadee 88 Black-headed Gull 271, 280 Black-legged Kittiwake 233, 267 Black-tailed Godwit 171, 207, 213 Black-throated Blue Warbler 170 Bluethroat 215 Blue Tit 92, 107, 215, 272, 281 Body condition 207; — mass 81, 227 Botaurus stellaris 205 Brambling 222 Branta canadensis 11, 271 Brood patch 89 BROUGHTON, RICHARD K., SHELLEY A. HINSLEY, PAUL E. BELLAMY, JANE E. CARPENTER and PETER ROTHERY Ageing and sexing Marsh Tits Poecile palustris using wing length and moult 88 BROUGHTON, RICHARD K., SHELLEY A. HINSLEY and PAUL E. BELLAMY Separation of Marsh Tit Poecile palustris from Willow Tit Poecile montana using a bill criterion 101 Brown Rat 282 Brünnich’s Guillemot 177 Bubo virginianus 269 Bucephala clangula 259 Bullfinch 170, 224, 272, 281 Burhinus oedicnemus 193, 206 Buteo buteo 150 Buzzard 150

CAIZERGUES, ALAIN see MATTHIEU GUILLEMAIN et al Calidris alba 224 C. alpina 172, 186, 271, 283 C. canutus 186, 208, 214 C. ferruginea 221 C. fuscicollis 172 Canada Goose 11, 271 Cannon nets 191, 233 Caprimulgus ruficollis 221 Carduelis cannabina 107, 222, 272, 283 C. carduelis 225, 272 C. chloris 107, 272 C. flammea 221 C. spinus 269, 272 CARPENTER, JANE E. see RICHARD K. BROUGHTON et al 88 Carpodacus erythrinus 221 Catharus guttatus 176 C. ustulatus 172, 175 Ceratogymna bucinator 177 Certhia familiaris 107 Cerulean Warbler 170 CES see Constant effort sites Cettia cetti 156, 283 Cetti’s Warbler 156, 283 Chaffinch 221, 272 Charadrius alexandrinus 221

British Trust for Ornithology, Ringing & Migration, 24 Vol 24 Index C. hiaticula 186, 223 Chiffchaff 107, 281 Chroicocephalus ridibundus 271, 280 Ciconia ciconia 8, 11, 162, 176, 180, 193 Cirl Bunting 193, 206 CLARK, JACQUIE A. see LIZ COIFFAIT et al 15, 104, 281 CLARK, JACQUIE A. see CHRIS P.F. REDFERN et al 145 CLARK, JACQUIE A. see STEPHEN R. BAILLIE et al 155 CLARK, JACQUIE A., KASPER THORUP and DAVID A. STROUD Quantifying the movement patterns of birds from ring recoveries 180 CLARK, JACQUIE A. see ROBERT A. ROBINSON et al 266 Clark’s 214 Coal Tit 214, 272 COIFFAIT, LIZ, JACQUIE A. CLARK, ROBERT A. ROBINSON, JEREMY R. BLACKBURN, BRIDGET M. GRIFFIN, KATE RISELY, MARK J. GRANTHAM, JOHN H. MARCHANT, TREVOR GIRLING and LEE BARBER Bird ringing in Britain and Ireland in 2006 15 COIFFAIT, LIZ, JACQUIE A. CLARK, ROBERT A. ROBINSON, JEREMY R. BLACKBURN, MARK J. GRANTHAM, JOHN H. MARCHANT, LEE BARBER, DIANA DE PALACIO, BRIDGET M. GRIFFIN and DORIAN MOSS Bird ringing in Britain and Ireland in 2007 104 COIFFAIT, LISETTE, CHRIS P.F. REDFERN, RICHARD M. BEVAN, JASON NEWTON and KIRSTEN WOLFF The use of intrinsic markers to study bird migration 169 COIFFAIT, LIZ, JACQUIE A. CLARK, ROBERT A. ROBINSON, JEREMY R. BLACKBURN, MARK J. GRANTHAM, DAVE I. LEECH, JOHN H. MARCHANT, LEE J. BARBER, DIANA DE PALACIO, BRIDGET M. GRIFFIN, DORIAN MOSS and MIRANDA M. SHEPHARD Bird ringing in Britain and Ireland in 2008 281 Collared Flycatcher 97, 170 Collared Pratincole 221 Colour fastness 11 Colour rings 11, 205, 213 Columba palumbus 268 Common Crossbill 170 Common Gull 271 Common Quail 221 Common Redpoll 221 Common Rosefinch 221 Common Sandpiper 285 Common Snipe Common Swift 81, 221, 272 Common Tern 177 Constant effort sites (CES) 1, 18, 107, 151, 158, 191, 199, 209, 283 Coot 271 Cormorant 13, 181, 266 Corn Bunting 208 Corncrake 193 caurinus 214 C. cornix 162 C. frugilegus 272 C. monedula 272 C. moneduloides 214 Coturnix coturnix 221 COULSON, J.C. Sexing Black-legged Kittiwakes by measurement 233 CRESSWELL, WILL The use of mass and fat reserve measurements from ringing studies to assess body condition 227 Crex crex 193 Cuckoo 156, 215 Cuculus canorus 156, 215 Curlew 268, 271 Curlew Sandpiper 221 Cyanistes caeruleus 92, 107, 215, 272, 281

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Cygnus cygnus 11 C. olor 271

DE LA CALZADA, FRAN see DAVID MIGUÉLEZ et al Delichon urbicum 1, 97, 268 Dendroica caerulescens 170 D. cerulea 170 D. petechia 172 DE PALACIO, DIANA see LIZ COIFFAIT et al 104, 281 Diomedea dabbenena 206 D. exulans 206 Dispersal 213 DNA 81, 208, 234 Dumetella carolinensis 201 Dunlin 172, 186, 271, 283 DUNN, JENNY C. and CHRIS WRIGHT Ageing and sexing the Yellowhammer Emberiza citrinella caliginosa during the non-breeding season 240 Dunnock 107, 215, 272

Eel 208 Eider 229, 271 Eleonora’s Falcon 229 Emberiza calandra 208 E. cirlus 193, 206 E. citrinella 240, 285 E. hortulana 97 E. schoeniclus 107, 201, 207, 215, 240, 272, 283 Erithacus rubecula 184, 228, 272, 283

Falco eleonorae 229 F. peregrinus 172 F. tinnunculus 271 Farmland birds 193 Fat deposition 95, 163, 278; — score 227 Ficedula albicollis 97, 170 F. hypoleuca 97, 108, 170, 285 FIEDLER, WOLFGANG New technologies for monitoring bird migration and behaviour 175 Fieldfare 221 Flight range 95 FRANSSON, THORD see K. SUSANNA S. HALL-KARLSSON and THORD FRANSSON Fratercula arctica 237, 271 Fringilla coelebs 221, 272 F. montifringilla 222 FUERTES, BENITO see DAVID MIGUÉLEZ et al Fulica atra 271 Fulmar 181 Fulmarus glacialis 181

Gallinula chloropus 271 Gannet 183, 271, 282 GARCIA-DEL-REY, EDUARDO, ANDREW G. GOSLER, JAVIER GONZALEZ and MICHAEL WINK Sexual size dimorphism and moult in the Plain Swift Apus unicolor 81 Garden Warbler 97, 107, 164, 184 glandarius 272 GAUTHIER-CLERC, MICHEL see MATTHIEU GUILLEMAIN et al Geolocator loggers 175, 206 GIRLING, TREVOR see LIZ COIFFAIT et al 15 Glareola pratincola 221 Global System for Mobile Communication (GSM) 178

British Trust for Ornithology, Ringing & Migration, 24 Vol 24 Index Goldcrest 272 Goldeneye 259 Golden Plover 253 Goldfinch 225, 272 GONZALEZ, JAVIER see EDUARDO GARCIA-DEL-REY et al GONZÁLEZ-JÁÑEZ, RUBÉN see DAVID MIGUÉLEZ et al GOSLER, ANDREW G. see EDUARDO GARCIA-DEL-REY et al GRANTHAM, MARK J. see LIZ COIFFAIT et al 15, 104, 281 GRANTHAM, MARK J. see ROBERT A. ROBINSON et al 266 Grasshopper Warbler 97 Gray Catbird 201 Greater Black-backed Gull 271 Greater Flamingo 13 Great Horned Owl 269 Great Reed Warbler 97, 171, 221 Great Skua 282 Great Tit 92, 107, 150, 214, 230, 272, 281 GREEN, RHYS E. see GUY Q.A. ANDERSON and RHYS E. GREEN Greenfinch 107, 272 Green Sandpiper 221 GREENWOOD, JEREMY J.D. 100 years of ringing in Britain and Ireland 147 Greylag Goose 271 GRIFFIN, BRIDGET M. see LIZ COIFFAIT et al 15, 104, 281 GSM see Global System for Mobile Communication GUILLEMAIN, MATTHIEU, RICHARD HEARN, ROY KING, MICHEL GAUTHIER-CLERC, GERALDINE SIMON and ALAIN CAIZERGUES Comparing the migration of Eurasian Teal Anas crecca from two main wintering areas of Western Europe: a long-term study from Essex, England, and the Camargue, France 273 Guillemot 271

Haematopus ostralegus 271 Haliaeetus albicilla 206 H. leucocephalus 9 HALL-KARLSSON, K. SUSANNA S. and THORD FRANSSON How far do birds fly during one migratory flight stage? 95 Harlequin Duck 221 HEARN, RICHARD see MATTHIEU GUILLEMAIN et al HELBIG, K.T. see K. PÜTZ et al Hermit Thrush 176 Herring Gull 271 HINSLEY, SHELLEY see RICHARD K. BROUGHTON et al 88, 101 Hippolais icterina 97 H. polyglotta 221 Hirundo rustica 1, 97, 162, 170, 221, 272 Histrionicus histrionicus 221 Honey Buzzard 222 Hooded Crow 162 Horseshoe Crab 208 House Martin 1, 97, 268 House Sparrow 108, 207, 215, 228, 272, 285

Icterine Warbler 97 Integrated demographic analysis 106, 282 IPM (Integrated Population Monitoring) 151, 189, 282

Jackdaw 272 Jack Snipe 221 272 JONES, M.G.W. see S. RAE et al JULLIARD, ROMAIN see ROBERT A. ROBINSON et al 199

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JUVASTE, R. see K. PÜTZ et al

Kentish Plover 221 Kestrel 271 KING, ROY see MATTHIEU GUILLEMAIN et al

Lanceolated Warbler 221 Lanius collurio 97 Laparotomy 81 Lapwing 221, 266, 271 Larus argentatus 271 L. canus 271 L. fuscus 7, 271 L. marinus 271 LEECH, DAVE I. see LIZ COIFFAIT et al 281 Lesser Black-backed Gull 7, 271 Lesser Whitethroat 97, 107 LIKER, ANDRÁS and LAJOS NAGY Migration of Mallards Anas platyrhynchos in Hungary: migration phenology, the origin of migrants, and long-term changes 259 Limosa lapponica 176 L. limosa 171, 207, 213 Limulus polyphemus 208 Linnet 107, 222, 272, 283 Locustella fluviatilis 221 L. lanceolata 221 L. luscinioides 171 L. naevia 97 Long-tailed Tit 107, 216, 281 Loxia curvirostra 170 Luscinia luscinia 97, 184, 221 L. svecica 215 Lymnocryptes minimus 221

Magpie 272 Mallard 163, 183, 194, 259, 271 Manx Shearwater 271, 282 MAPS see Monitoring Avian Productivity and Survivorship MARCHANT, JOHN H. see ROBERT A. ROBINSON et al MARCHANT, JOHN H. see LIZ COIFFAIT et al 15, 104, 281 Mark–recapture 5, 190 Marsh Sandpiper 221 Marsh Tit 88, 101, 285 Marsh Warbler 97 Melodious Warbler 221 Merops apiaster 221 Migration 273; — diurnal 96; — nocturnal 96; — phenology 259, 277; — spring 186, 253; — strategy 95, 165, 171 MIGUÉLEZ, DAVID, CARLOS ZUMALACÁRREGUI, BENITO FUERTES, HÉCTOR ASTIÁRRAGA, RUBÉN GONZÁLEZ-JÁÑEZ, ISABEL ROA and FRAN DE LA CALZADA Habitat, phenology and biometrics of the Aquatic Warbler Acrocephalus paludicola during autumn migration through a riverine wetland in Iberia 277 Milvus milvus 206 Mist net 200, 241 Mistle Thrush 272 MITCHELL, CARL and MARK TRINDER On reading colour rings 11 Monitoring Avian Productivity and Survivorship (MAPS) 201 Monteiro’s Storm Petrel 208 Moorhen 271 Mortality rate 150, 236 Morus bassanus 183, 271, 282

British Trust for Ornithology, Ringing & Migration, 24 Vol 24 Index MOSS, DORIAN see LIZ COIFFAIT et al 104, 281 Motacilla alba 272 M. flava 97, 156 Moult 220; — post-juvenile 88; — primary 81, 88, 93 Muscicapa striata 97 Mustela vison 282 Mute Swan 271

NAGY, LAJOS see ANDRÁS LIKER and LAJOS NAGY New Caledonian Crow 214 NEWTON, IAN Moult and plumage 220 NEWTON, JASON see LISETTE COIFFAIT et al 169 Northern Yellow Warbler 172 Northwestern Crow 214 Nucifraga columbiana 214 Numenius arquata 268, 271 N. phaeopus 221 N. tenuirostris 209

Oceanodroma monteiroi 208 Oenanthe oenanthe 97, 163 Ortolan Bunting 97 Osprey 9, 222 Otus scops 221 Oystercatcher 271

Pandion haliaetus 9, 222 Parasitism 215 Parus major 92, 107, 150, 214, 230, 272, 281 Passer domesticus 108, 207, 215, 228, 272, 285 P. montanus 206, 285 Passive Integrated Transponders (PIT) 175 PEDERSEN, K.T. see K. PÜTZ et al Peregrine 172 Periparus ater 214, 272 infaustus 214 Pernis apivorus 222 Phalacrocorax aristotelis 271 P. carbo 13, 181, 266 Phalaropus tricolor 221 Phenology 277 Phoenicopterus ruber 13 Phoenicurus phoenicurus 97 Phylloscopus collybita 107, 281 P. sibilatrix 97, 156 P. trochilus 97, 107, 171, 221, 272, 281 pica 272 Pied Flycatcher 97, 108, 170, 285 Pied Wagtail 272 Pink-footed Goose 191, 271 Plain Swift 81 Pluvialis apricaria 253 P. dominica 257 Pochard 105, 271 Poecile atricapilla 88 P. montana 88, 101, 107 P. palustris 88, 101, 285 Population change 201, 281; — dynamics 159, 189; — estimation 186, 189, 253

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Predation risk 214, 227 Productivity 207, 283 Prunella modularis 107, 215, 272 Puffinus puffinus 271, 282 PÜTZ, K., A.J. HELBIG, K.T. PEDERSEN, C. RAHBEK, P. SAUROLA and R. JUVASTE From fledging to breeding: long-term satellite tracking of the migratory behaviour of a Lesser Black-backed Gull Larus fuscus intermedius 7 Pyrrhula pyrrhula 170, 224, 272, 281

Radio tracking 175, 205 RAE, S., M.G.W. JONES, F. STEWART and R. TINGAY Counts of spring passage Golden Plover Pluvialis apricaria in north Lewis 253 RAHBEK, C. see K. PÜTZ et al RAS (see Retrapping Adults for Survival) Rattus norvegicus 282 Razorbill 271 Recovery rates 271 Red-backed Shrike 97 REDFERN, CHRIS P.F., JACQUIE A. CLARK, ROBERT A. ROBINSON and STEPHEN R. BAILLIE A century of ringing in Britain and Ireland 145 REDFERN, CHRIS P.F. see STEPHEN R. BAILLIE et al 155 REDFERN, CHRIS P.F. see LISETTE COIFFAIT et al 169 Red Kite 206 Red Knot 186, 208, 214 Red-necked Nightjar 221 Redshank 215, 230, 271, 283 Redstart 97 Redwing 272 Reed Bunting 107, 201, 207, 215, 240, 272, 283 Reed Warbler 95, 107, 170, 201, 272, 283 Regulus regulus 272 Reproduction 230 Retrapping Adults for Survival 1, 19, 108, 151, 158, 191, 209, 283 Ringed Plover 186, 223 Ringing effort 264 Ring reporting rates 176, 183, 195, 264, 266 Riparia riparia 1, 97, 148, 194, 272, 285 RISELY, KATE see LIZ COIFFAIT et al 15 Rissa tridactyla 233, 267 River Warbler 221 ROA, ISABEL see DAVID MIGUÉLEZ et al Robin 184, 228, 272, 283 ROBINSON, ROBERT A., DAWN E. BALMER and JOHN H. MARCHANT Survival rates of hirundines in relation to British and African rainfall 1 ROBINSON, ROBERT A. see CHRIS P.F. REDFERN et al 145 ROBINSON, ROBERT A., ROMAIN JULLIARD and JAMES F. SARACCO Constant effort: studying avian population processes using standardised ringing 199 ROBINSON, ROBERT A., MARK J. GRANTHAM and JACQUIE A. CLARK Declining rates of ring recovery in British birds 266 ROBINSON, ROBERT A. see LIZ COIFFAIT et al 15, 204, 281 ROBINSON, ROBERT A. see STEPHEN R. BAILLIE et al 155 Rook 272 Roseate Tern 207 ROTHERY, PETER see RICHARD K. BROUGHTON et al 88 Rudd 208

Sahel Region 1 Sanderling 224 Sand Martin 1, 97, 148, 194, 272, 285 SARACCO, JAMES F. see ROBERT A. ROBINSON et al 199 Satellite telemetry 7, 176, 206

British Trust for Ornithology, Ringing & Migration, 24 Vol 24 Index SAUROLA, P. see K. PÜTZ et al Savi’s Warbler 171 Saxicola rubetra 97 S. torquatus 225 Scardinius erythrophthalmus 208 SCHAUB, MICHAEL see FRANZ BAIRLEIN and MICHAEL SCHAUB SCHAUB, MICHAEL see STEPHEN R. BAILLIE and MICHAEL SCHAUB Scolopax rusticola 147 Scops Owl 221 Sedge Warbler 95, 107, 156, 201, 216, 272, 283 SELLERS, ROBIN M. Ageing Black-headed Gulls on leg and bill colour – a cautionary note 280 Setophaga ruticilla 171 Sex determination 88, 233, 240; dimorphism 81, 88, 240 Shag 271 SHARP, STUART P. Bird ringing as a tool for behavioural studies 213 SHEPHARD, MIRANDA M. see LIZ COIFFAIT et al 281 Siberian Jay 214 SIMON, GERALDINE see MATTHIEU GUILLEMAIN et al SIRs (see Stable isotopes) Siskin 269, 272 Slender-billed Curlew 209 Sociable Lapwing 207 Social behaviour 215 Somateria mollissima 229, 271 Song Thrush 151, 266, 281 Sparrowhawk 162, 222, 228, 271 Spotted Flycatcher 97 Spotted Redshank 221 Spotted Sandpiper 221 Stable isotopes 169, 209 Starling 162, 180, 214, 224, 266 Stercorarius skua 282 Sterna dougallii 207 S. hirundo 177 STEWART, F. see S. RAE et al Stonechat 225 Stone-curlew 193, 206 Strix aluco 271 STROUD, DAVID A. see JACQUIE A. CLARK et al Sturnus vulgaris 162, 180, 214, 224, 266 Survival rate 1, 17, 105, 155, 193, 207, 266 Swainson’s Thrush 172, 175 Swallow Roost Project 19 Sylvia atricapilla 97, 107, 170, 184, 216, 272, 281 S. borin 97, 107, 164, 184 S. communis 97, 107, 272, 283 S. curruca 97, 107 S. nisoria 97, 221

Tail length 81, 241 Tarsus length 241 Tawny Owl 271 Teal 162, 183, 271, 273 THORUP, KASPER see JACQUIE A. CLARK et al Thrush Nightingale 97, 184, 221 TINGAY, R. see S. RAE et al Total head length 233, 241 Treecreeper 107

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Tree Pipit 97, 156 Tree Sparrow 206, 285 TRINDER, MARK see CARL MITCHELL and MARK TRINDER Tringa erythropus 221 T. ochropus 221 T. stagnatilis 221 T. totanus 215, 230, 271, 283 Tristan Albatross 206 Trumpeter Hornbill 177 Tufted Duck 105, 271 Turdus iliacus 272 T. merula 107, 150, 227, 272, 281 T. philomelos 151, 266, 281 T. pilaris 221 T. viscivorus 272 Turnover 186, 282 Tyto alba 269

Uria aalge 271 U. lomvia 177

Vanellus gregarius 207 V. vanellus 221, 266, 271

Wandering Albatross 206 Weight change 228, 236, 278 Western Scrub-jay 214 Wheatear 97, 163 Whimbrel 221 Whinchat 97 White-fronted Goose 182, 259 White-rumped Sandpiper 172 White Stork 8, 11, 162, 176, 180, 193 White-tailed Eagle 206 Whitethroat 97, 107, 272, 283 Whooper Swan 11 Wigeon 149, 271 Willow Tit 88, 101, 107 Willow Warbler 97, 107, 171, 221, 272, 281 Wilsonia pusilla 171 Wilson’s Phalarope 221 Wilson’s Warbler 171 Wing length 81, 88, 233, 241 Wing tags 206 WINK, MICHAEL see EDUARDO GARCIA-DEL-REY et al Wintering areas 170, 273 WOLFF, KIRSTEN see LISETTE COIFFAIT et al 169 Woodcock 147 Woodpigeon 268 Wood Warbler 97, 156 WRIGHT, CHRIS see JENNY C. DUNN and CHRIS WRIGHT

Yellowhammer 240, 285 Yellow Wagtail 97, 156

ZUMALACÁRREGUI, CARLOS see DAVID MIGUÉLEZ et al

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