Geologic History Regents Review
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Using Dental Enamel Wrinkling to Define Sauropod Tooth Morphotypes from the Cañadón Asfalto Formation, Patagonia, Argentina
RESEARCH ARTICLE Using Dental Enamel Wrinkling to Define Sauropod Tooth Morphotypes from the Cañadón Asfalto Formation, Patagonia, Argentina Femke M. Holwerda1,2,3*, Diego Pol4,5, Oliver W. M. Rauhut1,3 1 Staatliche Naturwissenschaftliche Sammlungen Bayerns (SNSB), Bayerische Staatssammlung für Paläontologie und Geologie, München, Germany, 2 GeoBioTec, Departamento de Ciências da Terra, Faculdade de Ciências e Tecnologia (FCT), Universidade Nova de Lisboa, Caparica, Portugal, 3 Department of Earth and Environmental Sciences and GeoBioCenter, Ludwig Maximilians Universität, München, Germany, 4 Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Buenos Aires, Argentina, 5 Museo Paleontológico Egidio Feruglio, Trelew, Argentina * [email protected] Abstract OPEN ACCESS The early Middle Jurassic is regarded as the period when sauropods diversified and be- Citation: Holwerda FM, Pol D, Rauhut OWM (2015) came major components of the terrestrial ecosystems. Not many sites yield sauropod mate- Using Dental Enamel Wrinkling to Define Sauropod Tooth Morphotypes from the Cañadón Asfalto rial of this time; however, both cranial and postcranial material of eusauropods have been Formation, Patagonia, Argentina. PLoS ONE 10(2): found in the Cañadón Asfalto Formation (latest Early Jurassic–early Middle Jurassic) in e0118100. doi:10.1371/journal.pone.0118100 Central Patagonia (Argentina), which may help to shed light on the early evolution of Academic Editor: Peter Dodson, University of eusauropods. These eusauropod remains include teeth associated with cranial and man- Pennsylvania, UNITED STATES dibular material as well as isolated teeth found at different localities. In this study, an assem- Received: September 1, 2014 blage of sauropod teeth from the Cañadón Asfalto Formation found in four different Accepted: January 7, 2015 localities in the area of Cerro Condor (Chubut, Argentina) is used as a mean of assessing sauropod species diversity at these sites. -
Constraints on the Timescale of Animal Evolutionary History
Palaeontologia Electronica palaeo-electronica.org Constraints on the timescale of animal evolutionary history Michael J. Benton, Philip C.J. Donoghue, Robert J. Asher, Matt Friedman, Thomas J. Near, and Jakob Vinther ABSTRACT Dating the tree of life is a core endeavor in evolutionary biology. Rates of evolution are fundamental to nearly every evolutionary model and process. Rates need dates. There is much debate on the most appropriate and reasonable ways in which to date the tree of life, and recent work has highlighted some confusions and complexities that can be avoided. Whether phylogenetic trees are dated after they have been estab- lished, or as part of the process of tree finding, practitioners need to know which cali- brations to use. We emphasize the importance of identifying crown (not stem) fossils, levels of confidence in their attribution to the crown, current chronostratigraphic preci- sion, the primacy of the host geological formation and asymmetric confidence intervals. Here we present calibrations for 88 key nodes across the phylogeny of animals, rang- ing from the root of Metazoa to the last common ancestor of Homo sapiens. Close attention to detail is constantly required: for example, the classic bird-mammal date (base of crown Amniota) has often been given as 310-315 Ma; the 2014 international time scale indicates a minimum age of 318 Ma. Michael J. Benton. School of Earth Sciences, University of Bristol, Bristol, BS8 1RJ, U.K. [email protected] Philip C.J. Donoghue. School of Earth Sciences, University of Bristol, Bristol, BS8 1RJ, U.K. [email protected] Robert J. -
The Oldest Platypus and Its Bearing on Divergence Timing of the Platypus and Echidna Clades
The oldest platypus and its bearing on divergence timing of the platypus and echidna clades Timothy Rowe*†, Thomas H. Rich‡§, Patricia Vickers-Rich§, Mark Springer¶, and Michael O. Woodburneʈ *Jackson School of Geosciences, University of Texas, C1100, Austin, TX 78712; ‡Museum Victoria, PO Box 666, Melbourne, Victoria 3001, Australia; §School of Geosciences, PO Box 28E, Monash University, Victoria 3800, Australia; ¶Department of Biology, University of California, Riverside, CA 92521; and ʈDepartment of Geology, Museum of Northern Arizona, Flagstaff, AZ 86001 Edited by David B. Wake, University of California, Berkeley, CA, and approved October 31, 2007 (received for review July 7, 2007) Monotremes have left a poor fossil record, and paleontology has broadly affect our understanding of early mammalian history, been virtually mute during two decades of discussion about with special implications for molecular clock estimates of basal molecular clock estimates of the timing of divergence between the divergence times. platypus and echidna clades. We describe evidence from high- Monotremata today comprises five species that form two resolution x-ray computed tomography indicating that Teinolo- distinct clades (16). The echidna clade includes one short-beaked phos, an Early Cretaceous fossil from Australia’s Flat Rocks locality species (Tachyglossus aculeatus; Australia and surrounding is- (121–112.5 Ma), lies within the crown clade Monotremata, as a lands) and three long-beaked species (Zaglossus bruijni, Z. basal platypus. Strict molecular clock estimates of the divergence bartoni, and Z. attenboroughi, all from New Guinea). The between platypus and echidnas range from 17 to 80 Ma, but platypus clade includes only Ornithorhynchus anatinus (Austra- Teinolophos suggests that the two monotreme clades were al- lia, Tasmania). -
SUPPLEMENTARY INFORMATION: Tables, Figures and References
Samuels et al. Evolution of the patellar sesamoid bone in mammals SUPPLEMENTARY INFORMATION: Tables, Figures and References Supplementary Table S1: Mammals$ Higher taxa Genus sp. Estimated. age of Patellar Comments# (partial) specimen, location state 0/1/2 (absent/ ‘patelloid’/ present) Sinoconodonta Sinoconodon Jurassic 0 Patellar groove absent, suggests no rigneyi (Kielan- patella Jaworowska, Cifelli & Luo, Sinoconodon is included on our 2004) phylogeny within tritylodontids. Morganucodonta Megazostrodon Late Triassic, southern 0 rudnerae (Jenkins Africa & Parrington, 1976) Morganucodonta Eozostrodon sp. Late Triassic, Wales 0 Asymmetric patellar groove, (Jenkins et al., specimens disarticulated so it is hard 1976) to assess the patella but appears absent Docodonta Castorocauda 164 Mya, mid-Jurassic, 0 Semi-aquatic adaptations lutrasimilis (Ji, China Luo, Yuan et al., 2006) Docodonta Agilodocodon 164 Mya, mid-Jurassic, 0 scansorius China (Meng, Ji, Zhang et al., 2015) Docodonta Docofossor 160 Mya 0 brachydactylus (Luo, Meng, Ji et al., 2015) Docodonta Haldanodon 150-155 Mya, Late 0 Shallow patellar groove exspectatus Jurassic, Portugal (Martin, 2005b) Australosphenida Asfaltomylos Mid-Jurassic, South ? Postcranial material absent patagonicus America (Martin, 2005a) Australosphenida Ornithorhynchus Extant 2 Platypus, genome sequenced Monotremata anatinus (Warren, Hillier, Marshall Graves et (Herzmark, 1938; al., 2008) Rowe, 1988) Samuels et al. Australosphenida Tachyglossus + Extant 2 Echidnas Monotremata Zaglossus spp. (Herzmark, 1938; Rowe, 1988) Mammaliaformes Fruitafossor 150 Mya, Late Jurassic, 0 Phylogenetic status uncertain indet. windscheffeli (Luo Colorado & Wible, 2005) Mammaliaformes Volaticotherium Late Jurassic/Early ? Hindlimb material incomplete indet. antiquus (Meng, Cretaceous Hu, Wang et al., 2006) Eutriconodonta Jeholodens 120-125 Mya, Early 0 Poorly developed patellar groove jenkinsi (Ji, Luo Cretaceous, China & Ji, 1999) Eutriconodonta Gobiconodon spp. -
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Memoirs of Museum Victoria 74: 151–171 (2016) Published 2016 ISSN 1447-2546 (Print) 1447-2554 (On-line) http://museumvictoria.com.au/about/books-and-journals/journals/memoirs-of-museum-victoria/ Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians ROBIN M. D. BECK1,*, NATALIE M. WARBURTON2, MICHAEL ARCHER3, SUZANNE J. HAND4, AND KENNETH P. APLIN5 1 School of Environment & Life Sciences, Peel Building, University of Salford, Salford M5 4WT, UK and School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, NSW 2052, Australia ([email protected]) 2 School of Veterinary and Life Sciences, Murdoch University, 90 South Street, Murdoch, WA 6150, Australia ([email protected]) 3 School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, NSW 2052, Australia ([email protected]) 4 School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, NSW 2052, Australia ([email protected]) 5 National Museum of Natural History, Division of Mammals, Smithsonian Institution, Washington, DC 20013-7012, USA ([email protected]) * To whom correspondence should be addressed. E-mail: [email protected] Abstract Beck, R.M.D., Warburton, N.M., Archer, M., Hand, S.J. and Aplin, K.P. 2016. Going underground: postcranial morphology of the early Miocene marsupial mole Naraboryctes philcreaseri and the evolution of fossoriality in notoryctemorphians. Memoirs of Museum Victoria 74: 151–171. We present the first detailed descriptions of postcranial elements of the fossil marsupial mole Naraboryctes philcreaseri (Marsupialia: Notoryctemorphia), from early Miocene freshwater limestone deposits in the Riversleigh World Heritage Area, northwestern Queensland. -
Geologic History a – 1 1. a Whalebone That Originally Contained 200 Grams of Radioactive Carbon-14 Now Contains 25 Grams of C
Geologic History A – B1 Base your answers to questions 7 and 8 on the graph 1. A whalebone that originally contained 200 grams of below, which shows the generalized rate of decay of radioactive carbon-14 now contains 25 grams of carbon- radioactive isotopes over 5 half-lives. 14. How many carbon-14 half-lives have passed since this whale was alive? (1) 1 (2) 2 (3) 3 (4) 4 2. The diagram below represents a sample of a radioactive isotope. Which diagram best represents the percentage of this radioactive isotope sample that will remain after 2 half- lives? 7. If the original mass of a radioactive isotope was 24 grams, how many grams would remain after 3 half-lives? (1)12 (2)24 (3)3 (4)6 3. The table below shows information about the 8. Which radioactive isotope takes the greatest amount radioactive decay of carbon-14. of time to undergo the change shown on the graph? (1) carbon-14 (3) uranium-238 (2) potassium-40 (4) rubidium-87 ~~~~~~~~~~~~~~~ 9. An igneous rock contains 10 grams of radioactive potassium-40 and a total of 10 grams of its decay products. During which geologic time interval was this rock most likely formed? What is the amount of carbon-14 remaining after 28,500 (1) Middle Archean years? (2) Late Archean (3) Middle Proterozoic (4) Late Proterozoic 10. Which geologic event occurred in New York State at approximately the same time that eurypterids were 4. How old is a fossil that has radioactively decayed becoming extinct? through 4 half-lives of carbon-14? (1) the opening of the Atlantic Ocean (1) 5,700 years (3) 22,800 years (2) the uplift of the Appalachian Mountains (2) 17,100 years (4) 28,500 years (3) the formation of the Catskill Delta (4) the intrusion of the Palisades Sill 5. -
Mammal Disparity Decreases During the Cretaceous Angiosperm Radiation
Mammal disparity decreases during the Cretaceous angiosperm radiation David M. Grossnickle1 and P. David Polly2 1Department of Geological Sciences, and 2Departments of Geological Sciences, Biology, and Anthropology, rspb.royalsocietypublishing.org Indiana University, Bloomington, IN 47405, USA Fossil discoveries over the past 30 years have radically transformed tra- ditional views of Mesozoic mammal evolution. In addition, recent research provides a more detailed account of the Cretaceous diversification of flower- Research ing plants. Here, we examine patterns of morphological disparity and functional morphology associated with diet in early mammals. Two ana- Cite this article: Grossnickle DM, Polly PD. lyses were performed: (i) an examination of diversity based on functional 2013 Mammal disparity decreases during dental type rather than higher-level taxonomy, and (ii) a morphometric analysis of jaws, which made use of modern analogues, to assess changes the Cretaceous angiosperm radiation. Proc R in mammalian morphological and dietary disparity. Results demonstrate a Soc B 280: 20132110. decline in diversity of molar types during the mid-Cretaceous as abundances http://dx.doi.org/10.1098/rspb.2013.2110 of triconodonts, symmetrodonts, docodonts and eupantotherians dimin- ished. Multituberculates experience a turnover in functional molar types during the mid-Cretaceous and a shift towards plant-dominated diets during the late Late Cretaceous. Although therians undergo a taxonomic Received: 13 August 2013 expansion coinciding with the angiosperm radiation, they display small Accepted: 12 September 2013 body sizes and a low level of morphological disparity, suggesting an evol- utionary shift favouring small insectivores. It is concluded that during the mid-Cretaceous, the period of rapid angiosperm radiation, mammals experi- enced both a decrease in morphological disparity and a functional shift in dietary morphology that were probably related to changing ecosystems. -
Physical Setting/Earth Science Examination
PS/EARTH SCIENCE The University of the State of New York REGENTS HIGH SCHOOL EXAMINATION PHYSICAL SETTING EARTH SCIENCE Wednesday, January 28, 2009 — 9:15 a.m. to 12:15 p.m., only This is a test of your knowledge of Earth science. Use that knowledge to answer all questions in this examination. Some questions may require the use of the Earth Science Reference Tables. The Earth Science Reference Tables are supplied separately. Be certain you have a copy of the 2001 Edition (Revised November 2006) of these reference tables before you begin the examination. Your answer sheet for Part A and Part B–1 is the last page of this examination booklet. Turn to the last page and fold it along the perforations. Then, slowly and carefully, tear off your answer sheet and fill in the heading. The answers to the questions in Part B–2 and Part C are to be written in your separate answer booklet. Be sure to fill in the heading on the front of your answer booklet. You are to answer all questions in all parts of this examination according to the directions provided in the examination booklet. Record your answers to the Part A and Part B–1 multiple-choice questions on your separate answer sheet. Write your answers to the Part B–2 and Part C questions in your answer booklet. All work should be written in pen, except for graphs and drawings, which should be done in pencil. You may use scrap paper to work out the answers to the questions, but be sure to record all your answers on your separate answer sheet and in your answer booklet. -
Morphological Evidence Supports Dryolestoid Affinities for the Living Australian Marsupial Mole Notoryctes
Reviewing Manuscript Morphological Evidence supports Dryolestoid affinities for the living Australian Marsupial Mole Notoryctes Federico Agnolin, Nicolas Roberto Chimento Recent discoveries demonstrated that the southern continents were a cradle for the evolutionary radiation of dryolestoid mammals at the end of the Cretaceous. Moreover, it becomes evident that some of these early mammals surpassed the K/T boundary in South America, at least. Notoryctes is a poorly known living mammal, currently distributed in the s t deserts of central Australia. Due to its extreme modifications to fossoriality and peculiar n i anatomy, the phylogenetic relationships of this genus were debated in the past, but most r P recent authors agree in its marsupial affinities. A comparative survey of the anatomy of e Notoryctes reveals the poorly sustained marsupial affinities for the genus and striking r P plesiomorphies for a living mammal. Surprisingly, Notoryctes exhibits similarities with dryolestoids. Dryolestoids were a diverse and mainly mesozoic mammalian group phylogenetically nested between the egg-lying monotremes and derived therians. In particular, Notoryctes share a number of shared features with the extinct dryolestoid Necrolestes, from the Miocene of Patagonia. Both taxa conform a clade of burrowing and animalivorous dryolestoids that survived other members of their lineage probably due to their peculiar habits. Accordingly, Notoryctes constitutes a “living-fossil” from the supposedly extinct dryolestoid radiation, extending the biochron of the group more than 20 million years to the present day. The intermediate phylogenetic position of Notoryctes has the pivotal potential to shed light on crucial anatomical, physiological, ecological, and evolutionary topics in the deep transformation from egg-lying to placental mammals. -
Download/4084574/Burrow Young1999.Pdf 1262 Burrow, C
bioRxiv preprint doi: https://doi.org/10.1101/2019.12.19.882829; this version posted December 27, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under 1aCC-BY-ND 4.0 International license. Recalibrating the transcriptomic timetree of jawed vertebrates 1 David Marjanović 2 Department of Evolutionary Morphology, Science Programme “Evolution and Geoprocesses”, 3 Museum für Naturkunde – Leibniz Institute for Evolutionary and Biodiversity Research, Berlin, 4 Germany 5 Correspondence: 6 David Marjanović 7 [email protected] 8 Keywords: timetree, calibration, divergence date, Gnathostomata, Vertebrata 9 Abstract 10 Molecular divergence dating has the potential to overcome the incompleteness of the fossil record in 11 inferring when cladogenetic events (splits, divergences) happened, but needs to be calibrated by the 12 fossil record. Ideally but unrealistically, this would require practitioners to be specialists in molecular 13 evolution, in the phylogeny and the fossil record of all sampled taxa, and in the chronostratigraphy of 14 the sites the fossils were found in. Paleontologists have therefore tried to help by publishing 15 compendia of recommended calibrations, and molecular biologists unfamiliar with the fossil record 16 have made heavy use of such works. Using a recent example of a large timetree inferred from 17 molecular data, I demonstrate that calibration dates cannot be taken from published compendia 18 without risking strong distortions to the results, because compendia become outdated faster than they 19 are published. The present work cannot serve as such a compendium either; in the slightly longer 20 term, it can only highlight known and overlooked problems. -
New Tribosphenic Mammals from the Mussentuchit Local Fauna (Cedar Mountain Formation, Cenomanian), Utah, Usa
NEW TRIBOSPHENIC MAMMALS FROM THE MUSSENTUCHIT LOCAL FAUNA (CEDAR MOUNTAIN FORMATION, CENOMANIAN), UTAH, USA RICHARD L. CIFELLI, JOSHUA E. COHEN, and BRIAN M. DAVIS Cifelli, R.L., Cohen, J.E., and Davis, B.M. 2016. New tribosphenic mammals from the Mussentuchit Local Fauna (Cedar Mountain Formation, Cenomanian), Utah, USA. Palaeontologia Polonica 67, 67–81. LSID urn:lsid:zoobank.org:pub:C7A0F8DD-F04A- 467C-8E45-3BFABDEEC6F5 Herein we describe two new tribosphenic mammals from the Cedar Mountain Formation of Emery County, Utah, USA. The fossils occur in a narrow stratigraphic interval near the top of the unit (Mussentuchit Member) and date to approximately the Early–Late Cretaceous boundary (early Cenomanian, ~97 Mya). The specimens were recovered from screen-wash- ing bulk rock samples. Both taxa are based on isolated teeth; from the limited morphology represented, both lack obvious specializations of the two major therian groups, Metatheria and Eutheria, and for present purposes they are regarded as basal tribosphenidans. Dakotadens pertritus sp. n. is the more abundant of the two and is closely related to a previously known species from the overlying (upper Cenomanian) Naturita Formation. Culicolestes kielanae gen. et sp. n. is notable in its large size (similar to Pariadens mckennai from the same fauna) relative to most other pre-Campanian tribosphenidans. The mammals of the Mussentuchit Local Fauna, now fully described on the basis of available fossils, represent one of the most diverse assemblages known from the Mesozoic, including some 22 taxa. Tribosphenidans, however, are neither abundant nor diverse, and include only six recognized species; most Late Cretaceous assemblages of North America include twice as many. -
Postcranial Anatomy of Haldanodon Exspectatus (Mammalia, Docodonta) from the Late Jurassic (Kimmeridgian) of Portugal and Its Bearing for Mammalian Evolution
Blackwell Science, LtdOxford, UKZOJZoological Journal of the Linnean Society0024-4082The Lin- nean Society of London, 2005? 2005 1452 219248 Original Article Postcranium of Haldanodont. martin Zoological Journal of the Linnean Society, 2005, 145, 219–248. With 16 figures Postcranial anatomy of Haldanodon exspectatus (Mammalia, Docodonta) from the Late Jurassic (Kimmeridgian) of Portugal and its bearing for mammalian evolution THOMAS MARTIN* Forschungsinstitut Senckenberg, Senckenberganlage 25, 60325 Frankfurt am Main, Germany Received October 2004; accepted for publication March 2005 The postcranium of the Late Jurassic docodont Haldanodon exspectatus Kühne & Krusat, 1972 is represented by a partial skeleton and isolated bones of other individuals from the Late Jurassic (Kimmeridgian) of the Guimarota coal mine in Portugal. Haldanodon exhibits adaptations for a fossorial lifestyle such as stout and short limb bones and humeri with greatly expanded distal joints and strong deltopectoral crests. Short first and second phalanges and moderately curved and laterally compressed terminal phalanges with lateral grooves suggest that Haldanodon was a scratch-digger. The dorso-ventrally elongated, triangular scapula has a convex transverse profile with strongly lat- erally reflected anterior and posterior scapula margins, enclosing a deep trough-like ‘infraspinous fossa’. A supra- spinous fossa is not developed. The saddle-shaped glenoid facet is mainly formed by the coracoid and orientated antero-ventrally indicating a sprawling gait. No epiphyses were detected and the wide size range of humerus and femur possibly indicate a lifelong growth. Haldanodon is more derived than Morganucodon by complete reduction of the procoracoid, absence of the procoracoid foramen, and a peg-like coracoid. It shares with monotremes a postscapular fossa that is absent in Morganucodon.