DOCSLIB.ORG

Generieposition Pilsbry,1901

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Generieposition Pilsbry,1901 The malacologicalsocietymalacological society of Japan R# VENUS (Jup. Jour Malac.) Vol. 4S, No. 2 (1989):S5...95 r J< v; I; lf' a)}lfMtrcv)Li-( L ft ,im Generie Position of Kaliella･ yaeya･mensis Pilsbry, 1901 (Pulmonata: Helicarionidae) Rei UEsHIMA (Institute of Biolog'ical Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan 305) Abstract: Live materials of a helicarionid speeies Kaliellct yaeyamensis eollected from various localities <ineluding its type locality) were examined and its genital and radular features are deseribed. Although the present speeies has been assigned to the genus Para･kaliella of the tribe Eueonuli, subfamily Euconulinae, this species is transferred to the genus Liardetia (s,s.) of the tribe Mierocysti of the same subfamily on the basis ef genital and eon- chological features, L. yaeyameresis distributes at least from Ryukyu Islands to Caroline Islands through Formosa. As several related taxa, probably eonspecifie, are known from various loealities, its distribution range would be extended further. Local differentiation within this speeies is reeognized from the morphological variation in the penial stimulator. Introduction Katiella･ yaeyamensis Pilsbry, 1901 is a small helicarionid described from Yaeyama Islands and reported also from Formosa (Pilsbry 1905). This species, as well as other Japanese minute helicarionids, was first alloeated in the genus "widely KaLiella whieh was thought as a distributed" genus at that time (Pilsbry 1901). Habe's (1946,1957) anatomical studies revealed that Japanese heli- carionids formerly classified in Kalietta actually belonged to distinet groups. ``Kalielta" Following Habe's view, Kuroda (1958) divided -Japanese into two groups; Trochoehtamys Habe 1946 and ParakalieUa Habe 1946, and he pointed "Kalietta" out that yaeyamensis was conchologically distinguishable from those two groups. However, Kuroda (1963) later plaeed this speeies in Parakalielta without any explanation and this alloeation has been followed by subsequent authors (Azuma 1982, Minato 1988). During the review of Japanese Helicarionidae, many live speeimen$ of this species available for anatomieal studies were eollected not only from the type NII-Electronic Library Service The malacological society of Japan 86 VENUS: VoT. 48, No.2(1989) Ut 2 E PRM HG Ep × EC B PRM 1 PA/ VD -- Ut ."iEC---" PA - NII-Electronic Library Service The malacologicalsocietymalacological society ofJapanof Japan Ueshima: Generic position of Kalielta yaayamensis 87 loeality, but also from Minami-Daito Island and Formesa. Although the present species has been assigned to Parcticatiella which belongs to the Tribe Euconuli ef the subfamily Euconulinae, its genital features indicate that the present species is a member of the Tribe Microcysti of the same subfamily. Therefore, the present speeies eannot be retained in ParakalieZla, The following account presents this evidenee and provides anatomical features of this species and new generic assignment. Materials and Methods Materials dissected: Ishigaki Island, Yaeyama Islands: Mt. Omange, Hoshino (37 specimens, 30 Mareh 1984); Nagura (3 specimens, 31 Mareh 1984); Palm Park, Yonehara (2 specimens, 31 Mareh 1984). Iriomote Island, Yaeyama Islands: Nakano (26 specimens, 28 March 1984) ; Toyohara (2 specimens, 27 Mareh 1984). Minami-Daito Island, Daito Islands: Kyuto (12 specimens, 5 November 1986, collected by T. Kurozumi). Forrnosa (Taiwan): Ken-Ting Park (6 specimens, 22 August 1985); Cape Wolanpi (1 speeimen, 20 August 1985). General methods of dissection and scanning electron micrograph were carried out according to the methods described in Ueshima and Kurozumi (1988), except several specirnens collected from Ishigaki Island which were dissected alive in a saline. Observations Genitat-ia (Figs. 1-4) : Hermaphroditic gland (ovotestis) is small, censisted of mainly two clusters of clavate alveoli (each cluster is sometimes divided further into two in large specimens as shown in Fig. 2), embedded in mid-gut gland. Hermaphroditie duct is slender, rather short, weakly convoluted. Talon is small, frequently albumen small embedded in gland. Carrefour is but prominent. AIbumen g}anq is triangular, rather small for heliearionids, eonsisted of large.loosely cemented cells. Uterus is well developed, extremely swollen by pregnancy. Upper region ttttt tttttt tttt t ttt t tt tttt-tt Figs. 1-3. Genital organs of Liardetia (s,s.) yaayamensis. =JK t; }= tfOthdeX 1. Mt, Omange, Hoshino, Ishigaki, Island (showing an individual dissected alive in a saline) ; 2. Cape Wolanpi, Formosa ; 3. Kyuto, Minami-Daito Island (A. showing the natural condition in which penial complex is compactly folded and surrounded by a thin sheath; B. penial complex with its sheath cut out). Scale line 1 mm, AG, albumen gland; At, atrium;BC, bursa copulatrix;C, carrefour;Em, emryo; EC, epiphallic caecum; Ep, epiphallus; HD, hermaphroditic duct ; HG, herma- phroditic gland or ovotestis; P, penis; PA, penial appendix; Pr, prostate; PRM, penial retractor muscle ; PUO, post-uterine oviduct ; T, talon ; Ut, uterus ; VD, vas deferens. NII-Electronic Library Service The malacologicalsocietymalacological society ofJapanof Japan 88 VENVSt VoL 48, No. 2 (1989) = =}' Fig. 4.Internal structures Qf penis of Liardetia (s.s.) yaayamensis. X t; ifDltgCDNutIYue A. a typical specimen collected from Ken-Ting Park, Formosa ; B, C. specimens with secondary horny stimulator collected from Kyuto, Minami-Daito Island ; D. a specimen with large beaky horny stimulator but no secondary stimulator collectedfrom Kyuto, Minami-Daito Island. Scale bar 250 ptm in A, 100 pm in B, C and D. BHS, beaky horny stimulator; EC, epiphallic caecum; PA, penial appendix; PPd, penial pad ; SHS, secondary horny stiTnulator ; VD, vas deferens. NII-Electronic Library Service The malacologicalsocietymalacological society of Japan Ueshima: Generic positien of Kolietla yaq),amensis 89 of uterus which attaches to prostate is small, slender, defiated, always empty, refiexed backwards at the end of prostate and then becomes abruptly swollen to contain eggs or embryos. This embryo-containing region (cerresponding to free oviduct of other helicarionids) is large, thin-walled, completely separated from sperm duct, and followed by empty regiQn called post-uterine oviduct (Baker 1938). Post-uterine oviduct is slender, thin-walled, does not have glandular zone nor internal pilaster. This species is ovoviviparous, All examined specimens ineluding young materials have one to ten eggs or embryos in uterus. Eggs in uterus do not have ealcareous shell nor outer layer, Embryos become progressively grown from top to bottom and the largest embryo found near the gonopore has about 2 whorls. Prostate is small for helicarionids, consisted of numerous acini, contjnues from carrefour to the beginning of vas deferens. Vas deferens is slender, simple, entering the apex of epiphallus beside the epiphallic caecum. Epiphallus is slender, s!ightly broader than vas deferens, 'is with a prominent epiphallic eaeeum. Basal region of epiphallus which re- cognizable from outside as opaque white region has much gland eells than other region and is probably homologous to eorona of Philonesiae (Baker 1938). Epiphallic caeeum is large, about twice the diameter of epiphallus, inwardly eurved, with rounded apex, having ne prominent strueture internally. Penial retraetor muscle arises from the diaphragm, and inserts on the apex of penis. Penial appendix loeating near the insert,ion of penial retracter muscle is short but more slender when the animal is living (Fig. 1), tapers towards its apex, lacking pilaster or stimulator int,ernally. In a single specimen eolleeted from !shigaki Island, penial appendix is found to be everted. Upper half of penis is rather slender, having weak internal pilasters in several speeimens. Mid portion of penis is swollen, thick-walled, internally containing horny stimulator and one or two large penial pads (Fig. 4). Beaky horny stimulator whieh locates just below the penial pad is small, inwardly curved and its tip is peinting towards gonopore. As this beaky stimulator is found in all specimen examined and attaehing to the thick wall of penis tightly, this structure is probably persistent. In most of the specimens collected from Minami-Daito Island, a secondary horny stimulater is found just above the beaky stimulator. Thi's secondary stimulator is large, recurved, with blunt or truncated tip. The phenb- type with the secondary horny stimulator is dominent in Minami-Daito Island but is never found in other localities. Basa} region of penis is rather slender, thin-walled, lacking pilaster or stimulator internally. A thin sheath attaching around the mid portion ef penis surrounds upper half of penis, penial appendix, epiphallus and epiphallic caecum that are eompactly folded (Fig. 3A). This sheath called penis sheath by Baker (1938) seems not to be homologous to thick muscular penis sheath of higher helicarionids whi ¢ h surrounds only a part of penis or epiphallus (this sheath is cut off and not shown in Figs. 1, 2, 3B, so NII-Electronic Library Service The malacologicalsocietymalacological society of Japan 90 VENUS: VoL 48, Ne. 2 (1989) Fig. 5. Radula ef Liardetia (s, s.) yaeyamensis: showing a specimen collected from Mt. Omange, Hoshino, Ishigaki Island, = A i} } tfOta= A. marginal teeth ; B. central teeth and latero-marginal transition showing the shape of basal plates ; C. outermost marginal teeth ; D. central teeth and latero- marginal transition in slightly worn rows. Scale
Load more

Recommended publications
  • On a New Speoies of Land Shells of the Genus Kaliella
    ON A NEW SPEOIES OF LAND SHELLS OF THE GENUS KALIELLA. BLANFORD FROM THE SIMLA HILLS (MOLLUSOA, GASTROPODA.: FAMILY ZONITIDAE). By A. S. RAJAGOPALAIENGAR, M.So., Zoological Survey of India, Oalcutta. INTRODUCTION. This paper deals with four tiny land shells of the family Zonitidae received for determination from Dr. M. L. RoonwaI, Entomologist, 'Foredt Research Institute, Dehra Dun with the following interesting note regarding topography of the place from where the specimens were collected :- "The shells were collected from the cups of Arcenthobium minut·i88imum Hook. F. (a parasitic plant belonging to the family Loranthaceae) found on the twigs of Abiea pindrow (Silver fir) in the Simla Hills. These twigs were collected from tops of trees about 50 feet high growing alongside the road from Khadrala to Nankhari about Ii mile from the Khadral& rest house (height 9700 feet above sea level) in the Lower Bashahr forest division, Himachal Pradesh. The material was collected on 21-5-51 by G. D. Bhasin, Assistant Forest Entomologist in this branch. " Unfortunately, nothing is known about the soft parts of the snails. Though very small in size, the specimens, on a closer study, were found to exhibit certain very well-defined and significant features in their shell characters which left no doubt as to their distinctness from any of the hitherto known forms of the genus Kaliella. So, I approached Dr. H. C. Ray, Officer in charge of the Mollusca Section for elucidation. He after examining the same very critically came to & similar conclusion. But in order to be more sure about the identity, he sent the best specimen in the lot to the British Museum (Natural History), Lttndon, for opinion.
    [Show full text]
  • 1994 IUCN Red List of Threatened Animals
    The lUCN Species Survival Commission 1994 lUCN Red List of Threatened Animals Compiled by the World Conservation Monitoring Centre PADU - MGs COPY DO NOT REMOVE lUCN The World Conservation Union lo-^2^ 1994 lUCN Red List of Threatened Animals lUCN WORLD CONSERVATION Tile World Conservation Union species susvival commission monitoring centre WWF i Suftanate of Oman 1NYZ5 TTieWlLDUFE CONSERVATION SOCIET'' PEOPLE'S TRISr BirdLife 9h: KX ENIUNGMEDSPEaES INTERNATIONAL fdreningen Chicago Zoulog k.J SnuicTy lUCN - The World Conservation Union lUCN - The World Conservation Union brings together States, government agencies and a diverse range of non-governmental organisations in a unique world partnership: some 770 members in all, spread across 123 countries. - As a union, I UCN exists to serve its members to represent their views on the world stage and to provide them with the concepts, strategies and technical support they need to achieve their goals. Through its six Commissions, lUCN draws together over 5000 expert volunteers in project teams and action groups. A central secretariat coordinates the lUCN Programme and leads initiatives on the conservation and sustainable use of the world's biological diversity and the management of habitats and natural resources, as well as providing a range of services. The Union has helped many countries to prepare National Conservation Strategies, and demonstrates the application of its knowledge through the field projects it supervises. Operations are increasingly decentralised and are carried forward by an expanding network of regional and country offices, located principally in developing countries. I UCN - The World Conservation Union seeks above all to work with its members to achieve development that is sustainable and that provides a lasting Improvement in the quality of life for people all over the world.
    [Show full text]
  • Mollusca: Gastropoda) from Islands Off the Kimberley Coast, Western Australia Frank Köhler1, Vince Kessner2 and Corey Whisson3
    RECORDS OF THE WESTERN AUSTRALIAN MUSEUM 27 021–039 (2012) New records of non-marine, non-camaenid gastropods (Mollusca: Gastropoda) from islands off the Kimberley coast, Western Australia Frank Köhler1, Vince Kessner2 and Corey Whisson3 1 Department of Environment and Conservation of Western Australia, Science Division, PO Box 51, Wanneroo, Western Australia 6946; and Australian Museum, 6 College Street, Sydney, New South Wales 2010, Australia. Email: [email protected] 2 162 Haynes Road, Adelaide River, Northern Terrritory 0846, Australia. Email: [email protected] 3 Department of Aquatic Zoology, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia. Email: [email protected] ABSTRACT – The coast of the Western Australian Kimberley boasts an archipelago that comprises several hundred large islands and thousands much smaller. While the non–marine gastropod fauna of the Kimberley mainland has been surveyed to some extent, the fauna of these islands had never been comprehensively surveyed and only anecdotal and unsystematic data on species occurrences have been available. During the Western Australian Department of Environment and Conservation’s Kimberley Island Survey, 2008–2010, 22 of the largest islands were surveyed. Altogether, 17 species of terrestrial non–camaenid snails were found on these islands. This corresponds to about 75% of all terrestrial, non–camaenid gastropods known from the entire Kimberley region. In addition, four species of pulmonate freshwater snails were found to occur on one or more of four of these islands. Individual islands harbour up to 15, with an average of eight, species each. Species diversity was found to be higher in the wetter parts of the region.
    [Show full text]
  • Molluscan Forum 2018
    Number 72 (February 2019) The Malacologist Page 1 NUMBER 72 FEBRUARY 2019 Contents Editorial ………………………………...………….…... 2 News and Notes…………………………………………. 2 Travel Grant Report Franziska S. Bergmeier, Molluscan Forum 2018 Abstracts…..…...…………8 to 27 15th Deep-Sea Biology Symposium in Monterey Bay, Research Grant Reports: California (USA) …………………………………………39 Rodrigo Brincalepe Salvador In Memoriam Palaeogene land snails of Europe ...……………….……….. 28 Charles F Sturm.......................................................................40 Robert Fernandez-Vilert Colin Redfearn ……………………………………………...40 Tylodinae species complex in the Mediterranean Sea and Eastern Atlantic ………………………………...……... ... 28 Forthcoming Meetings ………………………………..41 Sydney Lundquist Notice of the Annual General Meeting Freshwater mussels as environmental indicators in UK river And Nominations for Council………………....…..43. systems using a sclerochronological approach ………………...29 Society Awards and Grants …………………………...44 Kasper P. Hendriks et al Notices concerning Membership ……………………..45 Fieldwork to sample microsnails for diet and microbiome studies along the Kinabatangan River, Sabah, Malaysian Borneo ……..33 Molluscan Forum 2018 Last November young malacologists from across Europe took part in the 20th Malaco- logical Forum at the Natural History Museum. The abstracts of the twenty nine presentations start inside on page 8. The Malacological Society of London was founded in 1893 and registered as a charity in 1978 (Charity Number 275980) Number 72 (February 2019) The Malacologist Page 2 EDITORIAL David Reid was editor of the Journal of Molluscan Studies from 2002 to 2018, in which year he began the process of retiring from his editorial role. Before completely retiring, David managed an extended handover to the new editor Dr Dinazarde Raheem so that she would be fully au fait with all the complicated issues which underly the production of the Journal.
    [Show full text]
  • Studies on the Ecology and Systematics of the Terrestrial Molluscs of the Lake Sibaya Area of Zululand, South Africa
    STUDIES ON THE ECOLOGY AND SYSTEMATICS OF THE TERRESTRIAL MOLLUSCS OF THE LAKE SIBAYA AREA OF ZULULAND, SOUTH AFRICA by A. C. VAN BRUGGEN Dept. of Systematic Zoology and Evolutionary Biology of the University, c/o Rijksmuseum van Natuurlijke Historie, Leiden, Netherlands and C. C. APPLETON Bilharzia Field Research Unit of the South African Medical Research Council, Nelspruit, South Africa With 10 text-figures and 4 plates CONTENTS Ι. General introduction 3 2. Notes on the climate, ecology and isolation of the coastal dune forest at Lake Sibaya, pertinent to its molluscan fauna 5 2a. Introduction 5 2b. Climate 7 20 Description of the habitats 12 2d. Ecological observations on the Mollusca 13 2e. The origin and isolation of the dune forest environment 14 3. Systematic part 17 3a. Gastropoda Prosobranchia 18 3b. Gastropoda Pulmonata 19 3c. Zoogeographical conclusions 38 4. Summary 41 5. References 41 1. GENERAL INTRODUCTION Lake Sibaya (sometimes misspelt Sibayi) is a freshwater lake on the subtropical coastal plain of Zululand, the northeastern part of the province of Natal, South Africa. Northern Zululand, usually called Tongaland, a fairly narrow stretch of country wedged between the Indian Ocean and the eastern border of Swaziland and bounded to the north by Mozambique, is particularly interesting from biogeographical and ecological points of view. The area harbours a rich terrestrial malacofauna, which has been partly discussed by Van Bruggen (1969), while some early data are also recorded 4 ZOOLOGISCHE VERHANDELINGEN 154 (1977) in Connolly's monograph (1939).The junior author (Appleton) was privileged to be able to spend a year in 1972/3 at the Lake Sibaya Research Station of Rhodes University (Grahamstown) on the eastern shore of the lake, during which period as a sideline he studied the ecology of and generally surveyed the land molluscs of the wide surroundings of the station.
    [Show full text]
  • Madagascar's Biogeographically Most Informative Land-Snail Taxa
    Biogéographie de Madagascar,1996 :563-574 MADAGASCAR'S BIOGEOGRAPELICALLY MOST INFORMATIVE LAND- SNAIL TAXA Kenneth C. EMBERTON & Max F. RAKOTOMALALA Molluscar? Biodiversiiy Institute, 216-A Haddon Hills, Haddon$eld,NJ 08033, U.S.A. Departementd'Entomologie, Parc Botanique et Zoologiquede Tsimbazaza, Antananarivo 101, MADAGASCAR ABSTRACT.-Madagascar's known native land-snail faunais currently classifiedinto 540 species(97% endemic) in 68 genera (29% endemic)in 25 families (0% endemic). Recent survey work throughout the island may as much as double this number of species and should provide, for the first time, adequate material and distributional data for robust cladistic and biogeographic analyses. Preliminary analysisof existing cladograms and range maps suggestsareas of endenlism with recurrent patterns of vicariance. Which of the many Madagascan taxa will yieldthe most biogeographic information perunit of effort? Based on the criteria of species number, nzonophyly, vagility, character accessibility, and Gondwanan areas of endemism, the best candidates are (a) acavoids (giant, k-selected, (( bird's-egg snails D), (b) Boucardicus (minute, top-shaped shells with flamboyant apertures), (c) charopids (minute, discoid shells with complex microsculptures), and (d) streptaxids (small-to-medium-sized, white-shelled, high-spired carnivores). KEY-W0RDS.- Land-snail, Madagascar, Informative, Biogeography RESUME.- La faune connueà l'heure actuelle des escargots terrestres de Madagascar peut être classée dans 540 espèces (97% endémiques), 68 genres (29% endémiques) et 25 familles(0% endémiques). Un récent travail d'inventaire réalisé dans l'ensemble l'îlede pourra amener à doubler le nombre d'espèces et devra fournir pour la première fois un matériel et des données adéquates sur la distribution des espèces permettant des analyses cladistiques et biogéographiques robustes.
    [Show full text]
  • The Subfamily Phaedusinae in Bhutan (Gastropoda, Pulmonata, Clausiliidae)
    The subfamily Phaedusinae in Bhutan (Gastropoda, Pulmonata, Clausiliidae) Edmund Gittenberger Naturalis Biodiversity Center, P.O. Box 9517, nl-2300 ra Leiden, The Netherlands; [email protected] [corresponding author] Pema Leda National Biodiversity Centre, Serbithang, Thimphu, Bhutan Sherub Sherub Ugyen Wangchuck Institute for Conservation and Environmental Research, Bumthang, Bhutan Choki Gyeltshen National Biodiversity Centre, Serbithang, Thimphu, Bhutan Gittenberger, E., Leda, P., Sherub, S. & Gyelt- (1914: 309) mentioned for the record only “Bhutan (Blan- shen, C., 2019. The subfamily Phaedusinae in Bhu- ford)”, without further details. According to Blanford & tan (Gastropoda, Pulmonata, Clausiliidae). – Bas- Godwin-Austen (1908: xxxi) nothing had been collected in teria 83 (4-6): 133-144. Leiden. “Bhutan east of longitude 89°”. West of that longitude there Published 9 November 2019 have been some changes in the political boundaries after the Anglo-Bhutan war of 1864. Localities that were once inside Bhutan, may no longer be Bhutanese territory now. Two genera of Clausiliidae, Phaedusinae, are reported from Despite the suggestive epithet, species like Alycaeus bhu- Bhutan. Cylindrophaedusa is represented with two species, tanensis Godwin-Austen, 1914, Dalingia bhutanensis God- viz. Cylindrophaedusa (Montiphaedusa) parvula Gitten- win-Austen, 1907, and Kaliella bhutanensis Godwin-Aus- berger & Leda, spec. nov. and C. (M.) tenzini Gittenberger ten, 1907, all of which described from the frontier areas of & Sherub, spec. nov. Phaedusa is represented with at least Bhutan, might no longer belong to the modern Bhutanese four species, viz. P. (P.) adrianae Gittenberger & Leda, spec. molluscan fauna. nov., P. (P.) bhutanensis Nordsieck, 1974, P. (P.) chimiae Git- tenberger & Sherub, spec. nov., and P. (P.) sangayae Gitten- berger & Leda, spec.
    [Show full text]
  • Diversity of Land Snails in the Karst Areas of Sta. Teresita, Cagayan Province, Luzon Island with Notes on New Distribution Records
    Philippine Journal of Science 150 (S1): 525-537, Special Issue on Biodiversity ISSN 0031 - 7683 Date Received: 04 Oct 2020 Diversity of Land Snails in the Karst Areas of Sta. Teresita, Cagayan Province, Luzon Island with Notes on New Distribution Records Julius A. Parcon1*, Ireneo L. Lit Jr.1,2, Ma. Vivian C. Camacho1,2, and Emmanuel Ryan C. de Chavez1,2 1Museum of Natural History 2Institute of Biological Sciences, College of Arts and Sciences University of the Philippines Los Baños, College 4031 Laguna, Philippines Malacofaunal research in a karst ecosystem is very limited not only in the northern region of Luzon Island but in the entire Philippines amidst extensive habitat disturbance and destruction. To address this, the diversity and abundance of land snails in the karst areas of Santa Teresita, Cagayan Province were determined. A total of 25 5 x 5 m2 quadrats were randomly set in five stations in the karst landscape. A total of 1206 land snails comprising 45 species under 36 genera representing 17 families were sampled. Camaenidae was the most represented family with 10 species. Luzonocoptis antennae constituted 25.1% of the total number of samples (303 individuals) and was the most abundant species in all stations. Of the 36 genera, five are new records in the Philippines. Several karst endemics and introduced species were recorded. Diversity indices showed diverse land snail fauna with Shannon-Weiner diversity index (H’) of 2.80, with evenness (J’) of 0.36 and dominance index of (D’) of 0.11. Species accumulation curve (SAC) showed late asymptote with a completeness ratio of 0.92.
    [Show full text]
  • Gittenberger Et Al
    JOURNAL OF CONCHOLOGY (2021), VOL.44, NO.1 11 RAHULA REVISITED (PULMONATA: EUCONULIDAE), WITH DATA FOR BHUTAN, INDIA (ASSAM), LAOS, VIETNAM AND INDONESIA, INCLUDING TWO NEW SPECIES 1 2 3 2 EDMUND GITTENBERGER , PEMA LEDA , SHERUB SHERUB & CHOKI GYELTSHEN 1National Biodiversity Center Naturalis, P.O. Box 9517, NL- 2300RA Leiden, The Netherlands 2National Biodiversity Centre, Serbithang, Ministry of Agriculture and Forests, Thimphu, Bhutan 3Ugyen Wangchuck Institute for Conservation and Environmental Research, Bumthang, Bhutan Abstract New data for Rahula species in Bhutan are given, including a new species and an extended description of R. trongsaensis based on a newly found fully grown shell. The genus is known from only the eastern half of the country. Two hitherto overlooked nominal taxa of Kaliella from Assam are regarded as Rahula species resembling R. trongsaensis; photographs of syntypes of these taxa are presented. Additions to the species list for Rahula that was published earlier are added, including a new species for Indonesia, Sulawesi. An updated distribution map for the genus is provided. Key words Euconulidae, Rahula, taxonomy, new species, Bhutan, Asia, distribution INTRODUCTION Supplementary data to the checklist of Rahula The genus Rahula Godwin- Austen, 1907 was species in Gittenberger, Leda & Sherub (2017b) classified with the family Helicarionidae are presented, including the description of Bourguignat, 1877, superfamily Helicarionoidea Rahula maasseni sp. nov. from Indonesia. The Bourguignat, 1877 by Zilch (1959: 306) and known range of the genus has expanded in Gittenberger et al. (2017a, c), whereas Vermeulen, southeastern direction in Indonesia, where R. Liew & Schilthuizen (2015), Gittenberger et al. maasseni sp. nov. now marks the distributional (2017b) and Foon & Marzuki (2020) adopted borderline in Sulawesi.
    [Show full text]
  • New Records of Non-Marine, Non-Camaenid Gastropods
    RECORDS OF THE WESTERN AUSTRALIAN MUSEUM 27 021–039 (2012) DOI: 10.18195/issn.0312-3162.27(1).2012.021-039 New records of non-marine, non-camaenid gastropods (Mollusca: Gastropoda) from islands off the Kimberley coast, Western Australia Frank Köhler1, Vince Kessner2 and Corey Whisson3 1 Department of Environment and Conservation of Western Australia, Science Division, PO Box 51, Wanneroo, Western Australia 6946; and Australian Museum, 6 College Street, Sydney, New South Wales 2010, Australia. Email: [email protected] 2 162 Haynes Road, Adelaide River, Northern Terrritory 0846, Australia. Email: [email protected] 3 Department of Aquatic Zoology, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia. Email: [email protected] ABSTRACT – The coast of the Western Australian Kimberley boasts an archipelago that comprises several hundred large islands and thousands much smaller. While the non–marine gastropod fauna of the Kimberley mainland has been surveyed to some extent, the fauna of these islands had never been comprehensively surveyed and only anecdotal and unsystematic data on species occurrences have been available. During the Western Australian Department of Environment and Conservation’s Kimberley Island Survey, 2008–2010, 22 of the largest islands were surveyed. Altogether, 17 species of terrestrial non–camaenid snails were found on these islands. This corresponds to about 75% of all terrestrial, non–camaenid gastropods known from the entire Kimberley region. In addition, four species of pulmonate freshwater snails were found to occur on one or more of four of these islands. Individual islands harbour up to 15, with an average of eight, species each.
    [Show full text]
  • Check-List of Land Pulmonate Molluscs of Vietnam (Gastropoda: Stylommatophora)
    Ruthenica, 2011, vol. 21, No. 1: 1-68. © Ruthenica, 2011 Published April 2011 http: www.ruthenica.com Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora) A.A. SCHILEYKO A.N. Severtzov Institute of Problems of Evolution, Russian Academy of Sciences, Leninsky Prospect 33, Moscow 119071, RUSSIA. E-mail: [email protected] ABSTRACT. Critical review of stylommatophoran mol- have been changed since the time of description. luscs of the fauna of Vietnam. The Check-list includes So, in the text of the Check-list the spellings of the 477 species and subspecies (96 genera, 20 families). For type localities are indicated exactly as in the original every species (subspecies) references to original de- descriptions; in the rubric “Distribution” the names scription, synonymy, type locality (in original spelling) including synonyms, shell dimensions (for slugs – body of places are given in their current spellings (or length) and distributional data are given. In the end the current names). In the end I placed the list of some index of mentioned molluscan names and the list of localities where for ancient (or used by the authors) localities with geographic coordinates is placed. names, the current names are indicated, and for some of them the geographical coordinates are Introduction given. Similarly the dimensions are cited according to the original description. In XIX – beginning of XX centuries numerous It should be also taken into consideration that in species of terrestrial molluscs have been described XIX and beginning of XX century the demarcation from the territory of Vietnam. The most eminent of boundaries between southern parts of China, researchers of the fauna of Indochina are A.
    [Show full text]
  • Checklist of the Mollusca of Cocos (Keeling) / Christmas Island Ecoregion
    RAFFLES BULLETIN OF ZOOLOGY 2014 RAFFLES BULLETIN OF ZOOLOGY Supplement No. 30: 313–375 Date of publication: 25 December 2014 http://zoobank.org/urn:lsid:zoobank.org:pub:52341BDF-BF85-42A3-B1E9-44DADC011634 Checklist of the Mollusca of Cocos (Keeling) / Christmas Island ecoregion Siong Kiat Tan* & Martyn E. Y. Low Abstract. An annotated checklist of the Mollusca from the Australian Indian Ocean Territories (IOT) of Christmas Island (Indian Ocean) and the Cocos (Keeling) Islands is presented. The checklist combines data from all previous studies and new material collected during the recent Christmas Island Expeditions organised by the Lee Kong Chian Natural History Museum (formerly the Raffles Museum of Biodiversty Resarch), Singapore. The checklist provides an overview of the diversity of the malacofauna occurring in the Cocos (Keeling) / Christmas Island ecoregion. A total of 1,178 species representing 165 families are documented, with 760 (in 130 families) and 757 (in 126 families) species recorded from Christmas Island and the Cocos (Keeling) Islands, respectively. Forty-five species (or 3.8%) of these species are endemic to the Australian IOT. Fifty-seven molluscan records for this ecoregion are herein published for the first time. We also briefly discuss historical patterns of discovery and endemism in the malacofauna of the Australian IOT. Key words. Mollusca, Polyplacophora, Bivalvia, Gastropoda, Christmas Island, Cocos (Keeling) Islands, Indian Ocean INTRODUCTION The Cocos (Keeling) Islands, which comprise North Keeling Island (a single island atoll) and the South Keeling Christmas Island (Indian Ocean) (hereafter CI) and the Cocos Islands (an atoll consisting of more than 20 islets including (Keeling) Islands (hereafter CK) comprise the Australian Horsburgh Island, West Island, Direction Island, Home Indian Ocean Territories (IOT).
    [Show full text]