<<

Bulletin of the American Rock Society

Volume 50 Number 1 Winter 1992 Cover: hybrid "Queen of Hearts" by Rob Proctor of Denver, Colorado Bulletin of the American Society

Volume 50 Number 1 Winter 1992

Features

Hybrid Pitcher , by Lawrence Mellichamp 3

A Sphagnum Garden, by Roberta Case 11

Miniature Waterlilies, by Stephen Doonan and Phil Pearson 13

Harbinger of Spring, by Judy Glattstein 16

Building a Waterfall, by Ray Radebaugh 21

Bog and Bog Plants, by Frederick W. Case, Jr. 45

A Woodland Waterfall and Pool, by Judith Jones 47

Miniature Water Gardens, by Joseph V. Tomocik 50

Rock in Ontario, by Barrie Porteous 51 Departments

Plant Portrait 58

Books 59

Propagation 65 Sarracenia x catesbaei x Sarracenia alata

2 Bulletin of the American Rock Garden Society Vol. 50(1) Hybrid Pitcher Plants

by Lawrence Mellichamp

I collected my first pitcher in people have water gardens, in shade or 1968 in the coastal flat woods and pine sun; but the bog garden, which requires savannas of southeastern North Caroli• full sun and just the right level of mois• na. Sarracenias grew commonly in ture, remains a challenge. these characterized by moist, Possibly, too, bog gardening reminds highly organic, nutrient-poor soils and me of my first encounter with the beau• frequent fires that kept dense vegeta• tiful but fragile habitats that are so rich tion from forming. Sundews, bladder- in , without the pesky insects of worts, butterworts, and the world- the wild. famous Venus'-flytrap also grew in I can't deny, also, that a reason for abundance, before the massive land- growing bog plants is they always elicit drainage activities of the 1970s which a response of excitement, or awe, from left much of the terrain dry pine planta• visitors. Although different from the joy tions. North Carolina has more differ• of seeing a lotus or waterlily in bloom, ent genera of carnivorous plants than bog plants are very gratifying in their any other place in the world, and many abnormal behavior. of the species make interesting speci• The genus Sarracenia (named after mens for the home bog garden. 18th century Canadian botanist and My favorites have been the pitcher physician Dr. Michel Sarrazin) contains plants. I still have growing a piece of 11 species, all but one of which are the original one I collected over 20 endemic to the southeastern United years ago. The other carnivorous plants States. That one, Sarracenia seem to come and go for me in cultiva• purpurea, is remarkable in having tion, acting as annuals or short-lived spread during the 10,000 years since perennials. Perhaps they are just more glaciation to the Gulf Coast and north• particular about their growing condi• ward all the way west across Canada in tions. Nevertheless, I enjoy working peat and on wet lake shores. It with sarracenias and other bog species also seems to flourish as an introduced because they seem to represent a plant in northern European bogs. neglected area of gardening. Many There is one close Sarracenia relative,

3 Darlingtonia californica, restricted to studied and described most of the natu• cool mountain streams and bogs in ral hybrids between pairs of species that northwestern California and adjacent grew together in the wild throughout the Oregon. Darlingtonia is difficult to Southeast. The genus was mono• grow in our hot southeastern summers graphed by Sidney McDaniel in 1971. and is not considered in this article. ("The Genus Sarracenia," Bulletin of Some growers may keep it alive, but I the Tall Timbers Research Station, have never seen plants grown well Tallahassee, 9:1-36.) In recent years, outside its native region. There are also Fred and Roberta Case have recognized distant relatives in southeastern two new distinct taxa from Alabama, Venezuela, in the genus Heliamphora, Sarracenia alabamensis (photo, p. 41) known as sun or pitchers. Since and S. alabamensis ssp. wherryi. In they are not hardy, they will not be addition, distinct forms have long been described here. known and cultivated, though not Most of the natural species of formally named, such as the giant form Sarracenia are well known as horticul• of S. minor from the Okefenokee tural subjects, having been highly in southeastern Georgia (photo, regarded in England and Europe since p. 43), a pink-flowered form of S. the Victorian era of the late 19th centu• purpurea from Mobile, and various albi• ry. The often-cited treatment by John no forms of most red-flowered species. MacFarlane in L.H. Bailey's Cyclope• Probably the most generally acknowl• dia of (1914) is still edged natural hybrid is S. x catesbaei required reading for anyone interested (photo, p. 43), a cross between S. in pitcher plants. flaua and S. purpurea. Wherever they I am especially proud of the fact that grow together, the hybrid may be found. an ancestor of mine, a Dr. Joseph H. Another common hybrid along the Gulf Mellichamp, who lived in the Bluffton Coast is S. x mitchelliana (S. leuco- area of southeastern South Carolina, phylla x S. purpurea). Both can be collected and studied pitcher plants in robust growers. Our research efforts the latter half of the 19th century. He now are to acquire—from the wild or published several scholarly papers man-made—specimens of every possi• describing his observations of Sarrace• ble cross between pairs of species. nia minor (photo, p. 44), and he is Sarracenias produce large, bee-polli• attributed with proving that sarracenias nated in spring. There is one catch and digest insects, i.e., that they nodding per stalk, and usually are truly insectivorous. Even Charles one stalk per growing lead (or "toe"). Darwin, who wrote the first book on Some smell sweet, others smell musty. insectivorous plants, never realized that A given individual plant blooms for sarracenias were carnivorous! about a week and the flowers are very One unusual trait of these species is showy, but it is the long-lasting pitchers that they hybridize freely among them• that make these plants famous and that selves, like tropical orchids, and the are mostly used to tell the different hybrids are fertile and capable of cross• species apart. Lately, individual pitchers ing with each other and back-crossing to of wild-collected S. leucophylla the species, often resulting in fantastic (photo, p. 41) are being sold in florist natural hybrid swarms in disturbed habi• shops as "cut flowers." All pitchers tats along the Gulf Coast. During the have hoods (mostly to keep out rain early 1950s, Ritchie Bell of North water), various hairs within the tubular Carolina and Fred Case of Michigan pitcher (to keep prey from

4 Bulletin of the American Rock Garden Society Vol. 50(1) escaping), and no moving parts to ing Co., Winston-Salem, NC), or catch insects. They are passive pitfall consult back issues of the Carnivorous traps. As in all carnivorous plants, it is Plant Newsletter, an excellent quarter• the that is modified to catch prey. ly guide to growing and understanding The pitchers will attract any number of all carnivorous plants, now in its 19th bees, moths, ants, flies, and wasps by year of publication (available from CPN, the sweet, sticky nectar secretions Biology Dept., California State Univer• around the mouth of the pitcher. The sity, Fullerton, CA 92634). insects slip into the tubes, can't get out, While the species are fairly easy to fall to the bottom and die, are digested grow, I feel that often they do not make by enzymes, and provide nutrients that the best ornamental plants because of are absorbed by the leaf to supplement their seasonal pitcher production, their their soil-based nutrition. We often use occasional dislike of all but the best cotton balls to keep moths out of the conditions of sun, soil, and water, and pitchers in the fall as they tend to over- the difficulty in obtaining propagated accumulate and "gum up" the pitchers (as opposed to wild-collected) speci• —as if a person tried to eat an mens. Hybrids are easier to maintain unplucked chicken. than species. All pitcher plants are very The species of pitcher plants are interesting and clearly intriguing to even described in Table 1, where brief distin• the most casual observer. Only the guishing features are given. Rowers are famous Venus'-flytrap ranks higher in either red or yellow; the pitchers are tall interest and that's because the trap or short and of various colors (influ• moves. Yet pitcher plants are larger, enced by the amount of sunlight); and more colorful, easier to grow, and more unless otherwise noted, the distributions readily propagated by division, and thus refer to the coastal plain region of the should be more widely known. In various states. The seasonal production response to an increasing interest in of pitchers can be distinctive: new pitcher plants on the part of the gener• pitchers may be produced starting in al public and based on comments from earliest spring and continuing until visitors to the University of North frost, or. there may be periods of no Carolina at Charlotte Botanical apparent growth. For example, some Gardens, where we have hundreds of species make new pitchers in spring specimens on display, I decided to only; others will have growing spurts in begin breeding man-made hybrids of spring and then again in late summer Sarracenia for ultimate distribution to or fall. Normally, fall pitchers are larger the horticulture trade. Only a few prop• and more colorful. Three species [S. agated species and hybrids are available fiaua (photo, p. 44), S. leucophylla now in the trade, and those only to a (photo, p. 41), and S. oreophila) very limited degree. produce distinctive, over-wintering, flat I started in 1984 experimenting with leaves that do not form tubes and are making crosses and germinating seeds. called phyllodia, in addition to their Cross pollinations are done in April and insect-trapping pitchers. These should May, as the potential parents come into not be cut off when grooming plants. bloom. There is much overlap of flow• For a good discussion of all the ering periods. Pollen can also be kept species and many of the forms and refrigerated or frozen for some time, hybrids, see Carnivorous Plants of allowing crosses between widely diver• the and Canada, by gent species even when there is no Donald E. Schnell (1976, Blair Publish• overlap in blooming times.

5 Seed pods mature in the late One of the most interesting aspects summer and fall and contain up to 500 of my research has been obtaining the seeds, depending on the degree of parental breeding stock. I collected typi• fertility of the cross. (In the wild, seeds cal specimens from the wild in North are apparently dispersed by water Carolina and especially the Mobile during the fall hurricane season, region of the Gulf Coast where most of according to reports by George Folkerts the species are found. Rare species at Auburn University.) Seeds may be were obtained from long-time growers stored dry in the refrigerator for several (before CITES permits were required). I years but must be stratified (given a cold was also most interested in finding and and moist period) to germinate. I did selecting unusually attractive natural experiments with stratification times to hybrids from the wild, as well as from find the optimal germination rates. I other collectors, including Fred Case of found that placing the seeds on top of a Michigan, Clyde Bramblett of Miami moist, peat-based medium (1 part (photo of plant selection, p. 42), John milled Canadian peat: 1 part perlite or Hummer of Arlington, Virginia, and vermiculite) and then refrigerating for Donald Schnell of Pulaski, Virginia. In four weeks at 40°F was acceptable for this way, I was able to gather together good germination. Under normal some of the finest breeding stock in conditions the seeds sprout in about existence. two weeks after refrigeration. I grow If there is need to justify my collect• the seedlings in a peat and perlite medi• ing from the wild, it is to say that I um, keeping them moist and in full sun, believe it is necessary for botanists to fertilizing twice a month with 1/4 know their plants first-hand. It also teaspoon per gallon 18N-18P-18K (or ensures knowledge of the and 9-45-15) water-soluble fertilizer during geographical origin of the material the summer. being utilized. Furthermore, it may have Even at that, the seedlings take been the only way to obtain certain about three years to reach flowering interesting and new breeding material maturity, though they begin showing from the wild. Pitcher plants are very some adult characteristics during the prolific species and will colonize any second good growing season. (Do not suitable habitat. Selective collecting fertilize adult plants.) Right now I have from the wild does no harm as long as hundreds of one- and two-year-old the habitat is not destroyed. Some seedlings waiting to be evaluated, of growers, in fact, have in cultivation which only a small handful will be kept specimens from now extinct popula• for further consideration for naming as tions of rare pitcher plants because they cultivars or use as breeding stock. The collected back before alteration. While I majority will be sold at our spring plant abhor mass collecting for profit, I see sale. Breeding is a slow and tedious no problem with knowledgeable process, too much work to simply people taking horticulturally interesting discard even the less spectacular plants. material of plants from the wild for While everyone who buys may not get purposes of propagation and breeding. a named cultivar, every seedling is a In fact, I encourage it, because that is potentially interesting plant. So you how many new forms become available see, they are not as easy to grow as for general distribution. annuals, but rather present a challenge I worked on hybridizing and selecting to the as great as that of some with Rob Gardner, curator of carnivo• of the choice alpines. rous plants at the North Carolina

6 Bulletin of the American Rock Garden Society Vol. 50(1) % u z

co NC , £ "o 3 TIT >

T X aT 3 A L o GJ s C td _D 6 3 o (0 a) oT < o -*-" CO o u •4-* (0 D ALt o VAt o ver y r < U z < U o z

>> j> C to 5 => O 3 2 -5 o - CO CO p CO CO cp to G 3 a c >- c c •C _D •c •c •c •S ~ w a a a a Q. (A a o s •c •c 3 o o *9 JG -C 0) w -p

o i-, CO oc

-C O) •C.S3 Qo) .<£> o T3 0) •=3 c •=6 c *5 S ...J! -o -C -TD * U -o a 2! & n J= O-S o-S •s * O CO _2 o ^3 c a c a C | I c g CO

0) J- | 2 -2 TJ TD T3 13 T3 -T3 o o -o <*> u c o £ _o 3 ^ 3 -Q O 3 .2 o .S! «3 "5 a