Wisconsin Bee Identification Guide
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Honey Bee Immunity — Pesticides — Pests and Diseases
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Distance Master of Science in Entomology Projects Entomology, Department of 2017 A GUIDEBOOK ON HONEY BEE HEALTH: Honey Bee Immunity — Pesticides — Pests and Diseases Joey Caputo Follow this and additional works at: https://digitalcommons.unl.edu/entodistmasters Part of the Entomology Commons This Article is brought to you for free and open access by the Entomology, Department of at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Distance Master of Science in Entomology Projects by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. Photo by David Cappaert, Bugwood.org 1 A GUIDEBOOK ON HONEY BEE HEALTH Honey Bee Immunity — Pesticides— Pests and Diseases By Joey Caputo A graduate degree project submitted as partial fulfillment of the Option III requirements for the de- gree of Masters of Science in Entomology at the graduate school of the University of Nebraska- Lincoln, 2017. Last updated April 2017 — Version 1.2 i Contents Introduction 1 Honey Bee Immune System 2 Mechanical and Biochemical Immunity 2 Innate and Cell-Mediated Immunity 2 Humoral Immunity 2 Social Immunity 3 Detoxification Complexes 5 Problems in Beekeeping 5 Colony Collapse Disorder (CCD) 5 Bacterial, Fungal and Microsporidian Diseases 6 American foulbrood 6 European foulbrood 7 Nosemosis 8 Chalkbrood 10 Crithidia 10 Stonebrood 11 Varroa Mite and Viruses 11 Varroa Biology and Life Cycle 12 Varroa Mite Damage and Parasitic Mite -
Orchard Mason Bees
Orchard Mason Bees This page offers WWW links outside WSU as an option for finding further information. No endorsement of products or information is implied by including those sites. The orchard mason bee (Osmia lignaria) is a gentle beneficial insect that has potential as a pollinator of apples, cherries, and other tree fruits. It is found throughout most of North America, particularly in wooded areas but often around homes in towns and cities. Homeowners sometimes become concerned when they see the bee entering cavities under shake siding or investigating nail holes or other cavities in wood during March through early June. These are not destructive insects, since they do not excavate holes in the wood, though they will clean out loose debris. No controls are recommended, since no damage is done. To prevent the bee from nesting, holes may be filled with caulking. The orchard mason bee is usually slightly smaller than a honey bee and a shiny dark blue in color. The actual size of the bee depends largely upon the size of the hole in which it grew. Males (Fig. 1) are smaller than females, have longer antennae and an additional tuft of light colored hairs on the face. Females (Fig. 2) have hairs on the underside of the abdomen, called the "scopa", adapted for carrying pollen. Fig. 1 Male orchard bee. Note the long antennae. (The red coloring is caused by Chaetodactylus sp. mites riding on the bee.) Female orchard mason bee on blossom. Fig. 2 The female approaches the flower from the top, transferring pollen from other flowers in the process. -
Xhaie'ican%Mllsllm
XhAie'ican1ox4tate%Mllsllm PUBLISHED BY THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK 24, N.Y. NUMBER 2 244 MAY I9, I 966 The Larvae of the Anthophoridae (Hymenoptera, Apoidea) Part 2. The Nomadinae BY JEROME G. ROZEN, JR.1 The present paper is the second of a series that treats the phylogeny and taxonomy of the larvae belonging to the bee family Anthophoridae. The first (Rozen, 1965a) dealt with the pollen-collecting tribes Eucerini and Centridini of the Anthophorinae. The present study encompasses the following tribes, all of which consist solely of cuckoo bees: Protepeolini, Epeolini, Nomadini, Ammobatini, Holcopasitini, Biastini, and Neolarrini. For reasons presented below, these tribes are believed to represent a monophyletic group, and consequently all are placed in the Nomadinae. It seems likely that the cleptoparasitic tribes Caenoprosopini, Ammoba- toidini, Townsendiellini, Epeoloidini, and Osirini are also members of the subfamily, although their larvae have not as yet been collected. Although the interrelationships of the numerous taxa within the Nomadinae need to be re-evaluated, the tribal concepts used by Michener (1944) are employed here. Adjustments in the classifications will certainly have to be made in the future, however, for Michener (1954) has already indicated, for example, that characters of the adults in the Osirini, the Epeolini, and the Nomadini intergrade. The affinities of the Nomadinae with the other subfamilies of the Antho- phoridae will be discussed in the last paper of the series. Because of char- 1 Curator, Department of Entomology, the American Museum of Natural History. 2 AMERICAN MUSEUM NOVITATES NO. -
Rainfall and Parasitic Wasp (Hymenoptera: Ichneumonoidea
Agricultural and Forest Entomology (2000) 2, 39±47 Rainfall and parasitic wasp (Hymenoptera: Ichneumonoidea) activity in successional forest stages at Barro Colorado Nature Monument, Panama, and La Selva Biological Station, Costa Rica B. A. Shapiro1 and J. Pickering Institute of Ecology, University of Georgia, Athens, GA 30602-2602, U.S.A. Abstract 1 In 1997, we ran two Malaise insect traps in each of four stands of wet forest in Costa Rica (two old-growth and two 20-year-old stands) and four stands of moist forest in Panama (old-growth, 20, 40 and 120-year-old stands). 2 Wet forest traps caught 2.32 times as many ichneumonoids as moist forest traps. The average catch per old-growth trap was 1.89 times greater than the average catch per second-growth trap. 3 Parasitoids of lepidopteran larvae were caught in higher proportions in the wet forest, while pupal parasitoids were relatively more active in the moist forest. 4 We hypothesize that moisture availability is of key importance in determining parasitoid activity, community composition and trophic interactions. Keywords Barro Colorado Nature Monument, Ichneumonoidea, La Selva, parasitoids, precipitation, tropical moist forest, tropical wet forest. istics of each parasitoid species and abiotic factors. Seasonal Introduction patterns of insect activity are often correlated with temperature, One of the largest groups of parasitic Hymenoptera is the as processes such as development and diapause are often superfamily Ichneumonoidea, which consists of two families intimately associated with temperature change (Wolda, 1988). (the Ichneumonidae and the Braconidae), 64 subfamilies and an Fink & VoÈlkl (1995) gave several examples of small insects for estimated 100 000 species world-wide (Gauld & Bolton, 1988; which low humidity and high temperature have detrimental Wahl & Sharkey, 1993). -
Alien Dominance of the Parasitoid Wasp Community Along an Elevation Gradient on Hawai’I Island
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln USGS Staff -- Published Research US Geological Survey 2008 Alien dominance of the parasitoid wasp community along an elevation gradient on Hawai’i Island Robert W. Peck U.S. Geological Survey, [email protected] Paul C. Banko U.S. Geological Survey Marla Schwarzfeld U.S. Geological Survey Melody Euaparadorn U.S. Geological Survey Kevin W. Brinck U.S. Geological Survey Follow this and additional works at: https://digitalcommons.unl.edu/usgsstaffpub Peck, Robert W.; Banko, Paul C.; Schwarzfeld, Marla; Euaparadorn, Melody; and Brinck, Kevin W., "Alien dominance of the parasitoid wasp community along an elevation gradient on Hawai’i Island" (2008). USGS Staff -- Published Research. 652. https://digitalcommons.unl.edu/usgsstaffpub/652 This Article is brought to you for free and open access by the US Geological Survey at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in USGS Staff -- Published Research by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. Biol Invasions (2008) 10:1441–1455 DOI 10.1007/s10530-008-9218-1 ORIGINAL PAPER Alien dominance of the parasitoid wasp community along an elevation gradient on Hawai’i Island Robert W. Peck Æ Paul C. Banko Æ Marla Schwarzfeld Æ Melody Euaparadorn Æ Kevin W. Brinck Received: 7 December 2007 / Accepted: 21 January 2008 / Published online: 6 February 2008 Ó Springer Science+Business Media B.V. 2008 Abstract Through intentional and accidental increased with increasing elevation, with all three introduction, more than 100 species of alien Ichneu- elevations differing significantly from each other. monidae and Braconidae (Hymenoptera) have Nine species purposely introduced to control pest become established in the Hawaiian Islands. -
Insects Parasitoids: Natural Enemies of Helicoverpa
Queensland the Smart State insects Parasitoids: Natural enemies of helicoverpa Introduction Helicoverpa caterpillars (often called heliothis) are serious pests of many crops in Australia. A range of parasitoid and predatory insects attack helicoverpa. Identifying and conserving these beneficial insects is fundamental to implementing pest management with a reduced reliance on chemical insecticides. This brochure describes the most important parasitoids of helicoverpa in Australian broadacre crops. Parasitoids versus parasites: What’s the difference? Parasitoids kill their hosts; parasites (such Figure 1. Netelia producta is one of the as lice and fleas) do not. All the insects most commonly encountered parasitoids in this brochure are parasitoids. Despite of helicoverpa. Females lay their eggs onto this difference, the terms parasitoid and caterpillars, and the hatching wasp larva parasite are often used interchangeably, if feeds on its host, eventually killing it. inaccurately. Parasitoids such as Netelia can be important biological control agents of helicoverpa in crops. (Photo: K. Power) All comments about parasitoid abundance in this publication are based on field observations in southern Queensland farming systems. These patterns may not occur in all parts of Australia. About parasitoids What is a parasitoid? How do parasitoids find their A parasitoid is an insect that kills (parasitises) hosts? its host — usually another insect — in Many adult parasitoids find their host by order to complete its lifecycle. In Australia, smell. They can detect the direct odour of helicoverpa are parasitised by many species the host itself, or odours associated with host of wasps and flies. All helicoverpa immature activity, such as plant damage or caterpillar stages are parasitised (that is, egg, caterpillar frass (dung). -
The Buzz About Bees: Honey Bee Biology and Behavior
4-H Honey Bee Leaders Guide Book I The Buzz About Bees: 18 U.S.C. 707 Honey Bee Biology and Behavior Publication 380-071 2009 To the 4-H Leader: The honey bee project (Books Grade 5 1 - 4) is intended to teach young people the basic biology and behavior of honey bees in addition to Living Systems 5.5 hands-on beekeeping management skills. The honey The student will investigate and understand that bee project books begin with basic honey bee and organisms are made up of cells and have distin- insect information (junior level) and advance to guishing characteristics. Key concepts include: instruction on how to rear honey bee colonies and • vertebrates and invertebrates extract honey (senior level). These project books are intended to provide in-depth information related Grade 6 to honey bee management, yet they are written for the amateur beekeeper, who may or may not have Life Science 5 previous experience in rearing honey bees. The student will investigate and understand how organisms can be classified. Key concepts include: Caution: • characteristics of the species If anyone in your club is known to have severe Life Science 8 allergic reactions to bee stings, they should not The student will investigate and understand that participate in this project. interactions exist among members of a population. The honey bee project meets the following Vir- Key concepts include: ginia State Standards of Learning (SOLs) for the • competition, cooperation, social hierarchy, and fourth, fifth, and sixth grades: territorial imperative Grade 4 Acknowledgments Authors: Life Processes 4.4 Dini M. -
Classification of the Apidae (Hymenoptera)
Utah State University DigitalCommons@USU Mi Bee Lab 9-21-1990 Classification of the Apidae (Hymenoptera) Charles D. Michener University of Kansas Follow this and additional works at: https://digitalcommons.usu.edu/bee_lab_mi Part of the Entomology Commons Recommended Citation Michener, Charles D., "Classification of the Apidae (Hymenoptera)" (1990). Mi. Paper 153. https://digitalcommons.usu.edu/bee_lab_mi/153 This Article is brought to you for free and open access by the Bee Lab at DigitalCommons@USU. It has been accepted for inclusion in Mi by an authorized administrator of DigitalCommons@USU. For more information, please contact [email protected]. 4 WWvyvlrWryrXvW-WvWrW^^ I • • •_ ••^«_«).•>.• •.*.« THE UNIVERSITY OF KANSAS SCIENC5;^ULLETIN LIBRARY Vol. 54, No. 4, pp. 75-164 Sept. 21,1990 OCT 23 1990 HARVARD Classification of the Apidae^ (Hymenoptera) BY Charles D. Michener'^ Appendix: Trigona genalis Friese, a Hitherto Unplaced New Guinea Species BY Charles D. Michener and Shoichi F. Sakagami'^ CONTENTS Abstract 76 Introduction 76 Terminology and Materials 77 Analysis of Relationships among Apid Subfamilies 79 Key to the Subfamilies of Apidae 84 Subfamily Meliponinae 84 Description, 84; Larva, 85; Nest, 85; Social Behavior, 85; Distribution, 85 Relationships among Meliponine Genera 85 History, 85; Analysis, 86; Biogeography, 96; Behavior, 97; Labial palpi, 99; Wing venation, 99; Male genitalia, 102; Poison glands, 103; Chromosome numbers, 103; Convergence, 104; Classificatory questions, 104 Fossil Meliponinae 105 Meliponorytes, -
Anthidium Manicatum, an Invasive Bee, Excludes a Native Bumble Bee, Bombus Impatiens, from floral Resources
Biol Invasions https://doi.org/10.1007/s10530-018-1889-7 (0123456789().,-volV)(0123456789().,-volV) ORIGINAL PAPER Anthidium manicatum, an invasive bee, excludes a native bumble bee, Bombus impatiens, from floral resources Kelsey K. Graham . Katherine Eaton . Isabel Obrien . Philip T. Starks Received: 15 April 2018 / Accepted: 21 November 2018 Ó Springer Nature Switzerland AG 2018 Abstract Anthidium manicatum is an invasive pol- response to A. manicatum presence. We found that B. linator reaching widespread distribution in North impatiens avoided foraging near A. manicatum in both America. Male A. manicatum aggressively defend years; but despite this resource exclusion, we found no floral territories, attacking heterospecific pollinators. evidence of fitness consequences for B. impatiens. Female A. manicatum are generalists, visiting many of These results suggest A. manicatum pose as significant the same plants as native pollinators. Because of A. resource competitors, but that B. impatiens are likely manicatum’s rapid range expansion, the territorial able to compensate for this resource loss by finding behavior of males, and the potential for female A. available resources elsewhere. manicatum to be significant resource competitors, invasive A. manicatum have been prioritized as a Keywords Exotic species Á Resource competition Á species of interest for impact assessment. But despite Interspecific competition Á Foraging behavior Á concerns, there have been no empirical studies inves- Pollination tigating the impact of A. manicatum on North Amer- ican pollinators. Therefore, across a two-year study, we monitored foraging behavior and fitness of the common eastern bumble bee (Bombus impatiens) in Introduction With increasing movement of goods and people Electronic supplementary material The online version of around the world, introduction of exotic species is this article (https://doi.org/10.1007/s10530-018-1889-7) con- increasing at an unprecedented rate (Ricciardi et al. -
Towards Simultaneous Analysis of Morphological and Molecular Data in Hymenoptera
Towards simultaneous analysis of morphological and molecular data in Hymenoptera JAMES M. CARPENTER &WARD C. WHEELER Accepted 5 January 1999 Carpenter, J. M. & W. C. Wheeler. (1999). Towards simultaneous analysis of molecular and morphological data in Hymenoptera. Ð Zoologica Scripta 28, 251±260. Principles and methods of simultaneous analysis in cladistics are reviewed, and the first, preliminary, analysis of combined molecular and morphological data on higher level relationships in Hymenoptera is presented to exemplify these principles. The morphological data from Ronquist et al. (in press) matrix, derived from the character diagnoses of the phylogenetic tree of Rasnitsyn (1988), are combined with new molecular data for representatives of 10 superfamilies of Hymenoptera by means of optimization alignment. The resulting cladogram supports Apocrita and Aculeata as groups, and the superfamly Chrysidoidea, but not Chalcidoidea, Evanioidea, Vespoidea and Apoidea. James M. Carpenter, Department of Entomology, and Ward C. Wheeler, Department of Invertebrates, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024, U SA. E-mail: [email protected] Introduction of consensus techniques to the results of independent Investigation of the higher-level phylogeny of Hymenoptera analysis of multiple data sets, as for example in so-called is at a very early stage. Although cladistic analysis was ®rst `phylogenetic supertrees' (Sanderson et al. 1998), does not applied more than 30 years ago, in an investigation of the measure the strength of evidence supporting results from ovipositor by Oeser (1961), a comprehensive analysis of all the different data sources Ð in addition to other draw- the major lineages remains to be done. -
The “Houdini” Fly Jean R
Natter’s Notes A New Pest of Mason Bees: The “Houdini” Fly Jean R. Natter Early in 2020, a new pest of mason bee, Cacoxenus sluggishly, and are often seen near the entry to a nesting indagator, was identified in Washington State for the tunnel. first time. It’s often referred to as the Houdini Fly After the mother bee leaves the nesting tunnel, the because of the unique way it escapes from the mason Houdini fly enters the tube, lays eggs on the pollen ball, bee’s nesting cell. It’s also nicknamed the Red Devil due then quickly exits. After the nesting cell is closed by the to its large red eyes, or just Devil Fly. It’s presence in mother mason bee, the fly larvae hatch and eat the Oregon is suspected but not yet verified. pollen ball. As a result, the mason bee larva starves. The arrival of the Houdini fly is suspected to be an unfortunate example of moving bees without carefully inspecting them and their nests prior to the move. “In New York, the first two records were in 2011, although it may have arrived there earlier. It had presumably come there from Europe, probably someone moving an unclean nest block,” said Josh Vlach, from the Oregon Department of Agriculture; interviewed by Andony Melathopolous; PolliNation Podcast #154 (2020). The Houdini fly, Cacoxenus indagator, is a newly identified kleptoparasite of mason bees in Washington State. Its presence is not yet verified in Oregon. This fly resembles the Drosophila fruit fly; it’s the same size, has large red eyes, but is a drab brown overall and moves sluggishly. -
Seasonal and Spatial Patterns of Mortality and Sex Ratio in the Alfalfa
Seasonal and spatial patterns of mortality and sex ratio in the alfalfa leafcutting bee, Megachile rotundata (F.) by Ruth Pettinga ONeil A thesis submitted in partial fulfillment of the requirements for the degree of Master of Science in Entomology Montana State University © Copyright by Ruth Pettinga ONeil (2004) Abstract: Nests from five seed alfalfa sites of the alfalfa leafcutting bee Megachile rotundata (F.) were monitored over the duration of the nesting season in 2000 and 2001, from early July through late August. Cells containing progeny of known age and known position within the nest were subsequently analyzed for five commonly encountered categories of pre-diapause mortality in this species. Chalkbrood and pollen ball had the strongest seasonal relationships of mortality factors studied. Chalkbrood incidence was highest in early-produced cells. Pollen ball was higher in late-season cells. Chalkbrood, parasitism by the chalcid Pteromalus venustus, and death of older larvae and prepupae , due to unknown source(s) exhibited the strongest cell-position relationships. Both chalkbrood and parasitoid incidence were highest in the inner portions of nests. The “unknown” category of mortality was highest in outer portions of nests. Sex ratio was determined for a subset of progeny reared to adulthood. The ratio of females to males is highest in cells in inner nest positions. Sex ratio is female-biased very early in the nesting season, when all cells being provisioned are the inner cells of nests, due to the strong positional effect on sex ratio. SEASONAL AND SPATIAL PATTERNS OF MORTALITY AND SEX RATIO IN THE ALFALFA LEAFCUTTING BEE, Megachile rotundata (F.) by .