Atlas of Yorkshire Coleoptera part 6 - Scarabaeoidea (Geotrupidae, Trogidae, Lucanidae and Scarabaeidae)
Introduction
This is Part 6 of the Atlas and covers the Scarabaeoidea, to include the Lucanidae, Trogidae, Bolboceratidae, Geotrupidae, and Scarabiidae.
101 species of Scarabaeoidea are known from Britain, of the roughly 35000 or so described worldwide taxa. Many species are fossorial and habitats range from dung and plant roots in the Scarabaeidae and Geotrupidae, rotting wood in the case of the Lucanidae , and carrion and dung in Trogidae. Please refer to the work of Freude, Harde and Lohse 8:266ff for the mid-European species, this publication covering most of the British species, and also the works of Jessop (1986), Britton (1956) and Joy (1932) for habitat requirements and distributions of the British species, although Britton and Joy are now much out of date.
The Scarabaeoidea is a rather poorly worked group in Yorkshire, and many species appear to be very scarce, especially of Aphodius, owing to the lack of fieldwork, with no records received for many species for many years.
Each species in the database is considered and in each case a distribution map representing records on the database (at April 2016) is presented. The number of records on the database for each species is given in the account in the form (a,b,c,d,e) where a to e are the number of records from VC61 to VC65 respectively. These figures include undated records (see comment on undated records in the paragraph below on mapping). As a recorder, I shall continue to use the vice-county recording system, as the county is thereby divided up into manageable, roughly equal, areas for recording purposes. For an explanation of the vice-county recording system, under a system devised in Watson (1883) and subsequently documented by Dandy (1969), Britain was divided into convenient recording areas ("vice- counties"). Thus Yorkshire was divided into vice-counties numbered 61 to 65 inclusive, and notwithstanding fairly recent county boundary reorganisations and changes, the vice-county system remains a constant and convenient one for recording purposes; in the text, reference to “Yorkshire” implies VC61 to VC65 ignoring modern boundary changes.
For some species there are many records, and for others only one or two. In cases where there are five species or less full details of the known records are given. Many common species have quite a high proportion of recent records. This is because the older county recorders tended not to record species they regarded as "common" quite as assiduously as we do today.
“Recent” records are those considered to be later than 1 January 1990 – this is also the break point used in the production of the distribution maps.
Habitat A brief statement on habitat is included where known and relevant.
Taxon status The national status of each taxon is given, this being the status as defined in JNCC’s Recorder6 data program. Status is included if Notable B or above. Please bear in mind that many of these statuses are now in need of revision, in the light of recent knowledge of distribution and frequency. The following status criteria are derived from Shirt (1987) and Hyman and Parsons (1992 and 1994):
RED DATA BOOK CATEGORY 1. RDB1 – ENDANGERED - Species which are known or believed to occur only as a single population within one 10km square of the OS National Grid.
RED DATA BOOK CATEGORY 2. RDB2 – VULNERABLE - Species which are declining throughout their range.
RED DATA BOOK CATEGORY 3. RDB3 – RARE - Species which are estimated to exist in only 15 or fewer 10km squares; this criterion may be relaxed where populations are likely to exist in more than 15 squares but occupy small areas of especially vulnerable habitat.
RED DATA BOOK CATEGORY I. RDB I – INDETERMINATE - Taxa considered to be endangered, Vulnerable or Rare, but where there is not enough information to say which of the three categories (RDB1 to 3) is appropriate.
RED DATA BOOK CATEGORY K. RDBK – INSUFFICIENTLY KNOWN - Taxa recently discovered or recognised in Great Britain, which may prove to be more widespread in the future (although some recent discoveries may be placed in other categories if the group to which they belong is thought to be not under-recorded).
PROVISIONAL RED DATA BOOK. Prdb - The prefix ‘p’ before any red Data Book category implies that the grading is provisional.
Notable A – species which do not fall within the RDB categories but are thought to occur in 30 or fewer 10km squares of the National Grid, or for less well-recorded groups, within seven or fewer vice-counties.
Notable B – species which do not fall within the RDB categories but thought to occur in between 31 and 100 10km squares of the National Grid.
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Nationally Notable - species which are estimated to occur within the range of 16 to 100 10km squares. (subdivision into Notable A and Notable B is not always possible because there may be insufficient information available). Now generally superseded by Nationally Scarce, and therefore no longer much used in modern literature.
The following status criteria are included in the species accounts and derive from those used in the Recorder 6 database. Please note that many of these statuses require reconsideration in the light of currently known records and distributions.
RedList_pre94-EN - Taxa in danger of extinction and whose survival is unlikely if the causal factors continue operating.. Superseded by new IUCN categories in 1994, so no longer in general use.
RedList_pre94-EX - Taxa which are no longer known to exist in the wild after repeated searches of their localities and other known likely places. Superseded by new IUCN categories in 1994, so no longer in general use.
RedList_pre94-NR - Taxa with small populations that are not at present Endangered or Vulnerable, but are at risk. (In GB, this was interpreted as species which exist in fifteen or fewer 10km squares). Superseded by new IUCN categories in 1994, so no longer in general use.
HabDir-A2* - Animal and plant species of Community interest (i.e. endangered, vulnerable, rare or endemic in the European Community) whose conservation requires the designation of special areas of conservation. Note that the contents of this annex have been updated in April 2003 following the Treaty of Accession.
A revised set of criteria governing taxon status is to be published shortly (DMa pers comm. to RJM), but for the time being the existing taxon status criteria and definitions included in the latest version of Recorder 6 will be used here.
Phenology Details are given for the range of months in the year for which there are records, with or without a peak month of occurrence, where the number and detail of records are sufficient to give a meaningful statement.
Maps Distribution maps have been produced using Dr Alan Morton’s DMAP software (version 7.0b). It may be worthwhile to note that as DMAP is asked to map pre- and post-1990 records in this application, any undated records will not be mapped. While efforts have been made to establish dates of records, some are difficult to ascertain, and I do not want to make guesses which may be wrong.
Nomenclature The YNU database is presently held in a copy of Recorder 6, which uses a new Coleoptera checklist (Duff, 2008) and this new list incorporates a very large number of taxon name changes since the checklist of Pope (1977); there is now a second edition of the latest Coleoptera checklist (Duff, 2012) which has not yet been incorporated into Recorder 6. Occasionally I receive records using “noddy names” (see Cooter, (1999) and Key (1999) for an excellent treatment of this subject), e.g. “Snail Hunter”, “Crucifix Beetle” and “Heath-glory ground beetle” (Cychrus caraboides, Panagaeus crux-major and Carabus nitens respectively). Such inventions were designed for use in political documents, and seem to have originated somewhere within Natural England (previously English Nature) or JNCC. For the purposes of serious science these terms are, in my opinion, vague, ridiculous, confusing and unhelpful. I do not accept records using such names only, unless accompanied by specimens or other convincing evidence.
The arrangement of species in the Atlas follows that of the latest Duff (2012) Checklist. There is however a new checklist of the Scarabaeidae shortly to be published by Darren Mann. Included in this all subgenera of Aphodius will be raised to generic rank. In the meantime for the purposes of this atlas the existing Duff 2012 checklist will be used.
Quality of the records A regrettable aspect of some of the records within the YNU database in particular (and I suspect sometimes within entomological datasets in general) is the lack of attention to detail accompanying the record, especially of accurate grid references, lack of pertinent comments as to the micro-habitat of the capture, its association with plants/animals, and sometimes, in the case of some modern records, a lack of care in determination, and the recording of the sex of the specimen. Old records, although sound, tend not to give much or indeed any of this detail. Old records on the database so rarely give habitat details but sometimes interpretation of the site name can give clues to the habitat type. Grid references could not be applied before 1930 anyway, and grid references do not figure in the paper archive of the YNU Coleoptera database until the late 1970s. Apparently it was not considered necessary to give that level of accuracy. Entering records onto Recorder 6 requires a grid reference, and with older records I have added a grid reference where I believe this has been possible. If the site name is a vague one I have used a 10km reference, and where a 1km reference is deemed accurate enough I have included that. There can be little excuse with modern records not to include some accurate comment with the record, and at least a six-figure grid reference.
Taxonomic knowledge and expertise have increased greatly since the days of Walsh, Bayford and Stainforth. Much good literature now exists for the taxonomist in the form of dichotomous keys, and when identifying specimens, especially with critical species pairs or groups, too much reliance on the outcome of keys can be placed on a determination, without apparently considering the habitat in which the specimen was taken, or the host plant/animal. Atlases of the national distributions of various orders of insect are now becoming available and should always be consulted. These often give a guide to the likelihood or otherwise of a determination, especially in scarce taxa. Museum collections containing correctly determined and labelled specimens are invaluable in accurate recording; museum collections are there for our use and should be consulted. If a species is ‘new’ to a vice-county or is very significant in some other respect, a second opinion should always be sought or an acknowledged specialist in the group consulted. A voucher should always be retained by the
2 | P a g e collector if possible. The recorder can only deal with the information he is given, and often has to beware of identifications made on photographic evidence only, and identifications of ‘difficult’ species by non-specialists. It takes many years to become competent in many areas of entomology, and the recorder often has to rely on the expertise of co-workers, especially when records covering large Orders are being documented.
I have adopted a policy in my county recording whereby any received record of a species afforded Notable B or above, especially if a member of a ‘difficult’ beetle group, should be confirmed by a specialist in that group before the record is entered onto the database.
Ultimately, the responsibility for verification of records on the YNU database rests with me. With modern records received from reputable coleopterists there is little problem. Some older records require vouchers for confirmation but have been retained in the database and flagged as requiring confirmation.
Contributors:
These are listed in alphabetical order of the first initial as these appear in the species accounts. Readers are requested to contact the author if initials in the text do not appear in the following list:-
AMR - AM Robertson MLT - ML Thompson
CJ - Colin Johnson NR - Neil Redgate
DA - D Applegarth PB - Paul Buckland
DM - David Maude PK - Peter Kendall
DMa - Darren Mann PS - Peter Skidmore
DS - Diane Stones RBA - Robert Angus (Huddersfield)
EGB - EG Bayford RGB - Roger Booth
EWA - Edward (Ted) W Aubrook RJM - Robert John Marsh
GBW – George Beckworth Walsh RL - Robert Lawson
HB - Harry Britten jnr. RSK - Roger S Key
HHC – Herbert Henry Corbett TS - T Stainforth
HP - H Pollard WAE - William A Ely
HWE - Herbert Willoughby Ellis WCH – William Croser Hey
ISR - Invertebrate Site Register WDH – Walter Douglas Hincks
JHF - John Flint WES – William Ernest Sharp
JM - J Melville WJF – William John Fordham
MLD - Mike Denton YNU – Yorkshire Naturalist Union reports in Naturalist.
0. Family GEOTRUPIDAE 462
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Subfamily BOLBOCERATINAE Mulsant, 1842
ODONTEUS Samouelle, 1819 ODONTAEUS Dejean, 1821 armiger (Scopoli, 1772) mobilicornis (Fabricius, 1775) testaceus (Fabricius, 1775)
Odonteus armiger (Scopoli, 1772)
3 WATSONIAN (0,0,1,0,0) RedList_pre94-NR. A rare species 2 YORKSHIRE in Yorkshire with only one record on the database: Huddersfield (SE11), 1892, JM, see 1 Naturalist 1892:37. A species associated with 0 NY NZ NZ OV SD SE SE TA subterranean fungi (see Jessop, 1986:15), 9 with a known distribution in southern 8 England, preferring unimproved grasslands 7 on chalk, limestone and sand. This record 6 requires verification. 5 4 3 2 1
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Subfamily GEOTRUPINAE Latreille, 1802
TYPHAEUS Leach, 1815 TYPHOEUS auctt. (misspelling) MINOTAURUS Mulsant, 1855 typhoeus (Linnaeus, 1758) pumilus (Marsham, 1802) vulgaris (Leach, 1815) thyphoeus (Leach, 1815)
Typhaeus typhoeus (Linnaeus, 1758)
3 WATSONIAN (14,12,3,1,0) An infrequent species which burrows under dung of various herbivores, 2 YORKSHIRE with a mainly eastern distribution in 1 Yorkshire. (i-x) NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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ANOPLOTRUPES Jekel, 1866 GEOTRUPES sensu auctt. partim non Latreille, 1796 stercorosus (Scriba, 1791) sylvaticus (Panzer, 1798)
Anoplotrupes stercorosus (Hartmann in Scriba, 1791)
3 WATSONIAN (0,6,6,0,3) An upland species generally, with records mainly from the west and north of 2 YORKSHIRE our recording area. (I have seen specimens in 1 collections confused with Geotrupes 0 NY NZ NZ OV SD SE SE TA stercorarius which is more generally 9 distributed in lowland sites). In herbivore 8 dung. (i-ix, peak vi). 7 6 5 4 3 2 1
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GEOTRUPES Latreille, 1797 mutator (Marsham, 1802)
Geotrupes mutator (Marsham, 1802)
3 WATSONIAN (2,0,0,0,0) Notable B. Rare with only two records on the database: Filey (TA18) 1871, 2 YORKSHIRE GBW; Cottingham (TA0534) 20/10/1948, 1 Anon. No further details are attached to 0 NY NZ NZ OV SD SE SE TA these records, but the species inhabits dung 9 of cow and horse. This species is often 8 misidentified and referable to 7 spiniger/stercorarius (DMa pers comm. to 6 RJM) and therefore vouchers would be 5 required to confirm our records. 4 3 2 1
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spiniger (Marsham, 1802) foveatus (Marsham, 1802) puncticollis Malinowsky, 1811 sublaevigatus Stephens, 1830
Geotrupes spiniger (Marsham, 1802)
3 WATSONIAN (11,1,13,3,1) Rather scarce with most 2 YORKSHIRE records from the south of the region. Horse dung has often been the habitat mentioned 1
NY NZ NZ OV in our records. (i v-xi, peak ix) 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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6 | P a g e stercorarius (Linnaeus, 1758) punctatostriatus Stephens, 1830
Geotrupes stercorarius (Linnaeus, 1758)
3 WATSONIAN (8,12,45,6,9) Widespread and the most 2 YORKSHIRE abundant of the Geotrupidae, and recorded from high moors (beware confusion with G. 1 stercorosus) down to sea level in the 0 NY NZ NZ OV SD SE SE TA southeast. Sheep dung is mentioned in 9 several of our records. (i-x, peaks vi,ix) 8 7 6 5 4 3 2 1
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TRYPOCOPRIS Motschulsky, 1860 GEOTRUPES sensu auctt. partim non Latreille, 1796
pyrenaeus (Charpentier, 1825)
NO RECORDS ON THE DATABASE
vernalis (Linnaeus, 1758) 21. Family 1819 465 Trypocopris v ernalis (Linnaeus, 1758)
3 WATSONIAN (1,5,0,0,1) Notable B. Very scarce with only one recent record. No habitat details are given 2 YORKSHIRE in any of our records except one - one 1 specimen in heather at a North York Moors 0 NY NZ NZ OV SD SE SE TA site. Sheep dung is known to be a favourite 9 habitat. 8 7 6 5 4 3 2 1
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TROX Fabricius, 1775 perlatus (Goeze, 1777) hispidus sensu auctt. Brit. non (Pontoppidan, 1763)
NO RECORDS ON THE DATABASE sabulosus (Linnaeus, 1758)
Trox sabulosus (Linnaeus, 1758)
3 WATSONIAN (0,0, 0,1,0) Notable A. There is a single 2 YORKSHIRE record on our database: Leeds (SE33) 1946, DA. No other details are given. This record 1 requires confirmation as sabulosus has a 0 NY NZ NZ OV SD SE SE TA strong preference for sandy soils and is 9 usually a misidentification of scaber away 8 from sandy soils (DMa pers. comm. to RJM). 7 Jessop (1986) and Freude, Harde & Lohse 6 8:270 make note of animal carcasses being a 5 habitat. 4 3 2 1
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scaber (Linnaeus, 1767) arenosus (Gmelin in Linnaeus, 1790) arenarius (Paykull, 1798) trisulcatus Curtis, 1845
Trox scaber (Linnaeus, 1767)
3 WATSONIAN (0,1,11,2,0) Very infrequent and mainly recorded from the south of the county. 2 YORKSHIRE Several records from old trees (possibly from 1 bird nests) and once in a light trap. (iv-viii). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1989 Map produced for Yorkshire Naturalists' Union by DMAP 17/4/2016 pre 31/12/1989 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 22. Family LUCANIDAE 1804 466
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Subfamily SYNDESINAE MacLeay, 1819
SINODENDRON Hellwig, 1792 cylindricum (Linnaeus, 1758)
Sinodendron cylindricum (Linnaeus, 1758)
3 WATSONIAN (9,29,126,25,5) Widespread in the county, and recorded from the dead wood of a wide range 2 YORKSHIRE of deciduous trees. (ii-xi, peak vi). 1
NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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Subfamily LUCANINAE Latreille, 1804
DORCUS MacLeay, 1819 parallelipipedus (Linnaeus, 1758) parallelopipedus auctt. (misspelling)
Dorcus parallelipipedus (Linnaeus, 1758)
3 WATSONIAN (30,3,40,10,0) Recorded widely from the southeastern quarter of the county, our 2 YORKSHIRE records include mention of beech, elm, ash 1 and poplar. (v-xii, peak vii) NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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LUCANUS Scopoli, 1763 cervus (Linnaeus, 1758) inermis (Marsham, 1802)
Lucanus cerv us (Linnaeus, 1758)
3 WATSONIAN (1,3,3,3,0) HabDir-A2* Rarely recorded - I 2 YORKSHIRE have received some records which clearly should be referrable to Dorcus. Where 1 records have come from reputable 0 NY NZ NZ OV SD SE SE TA coleopterists, the insects have been reported 9 from woodyards and factories, having been 8 imported from southern Britain or overseas. I 7 am not aware of this species breeding in 6 Yorkshire. 5 4 3 2 1
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PLATYCERUS Geoffroy, 1762 caraboides (Linnaeus, 1758)
NO RECORDS ON THE DATABASE 23. Family SCARABAEIDAE Latreille, 1802 467
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Subfamily AEGIALIINAE Laporte, 1840
AEGIALIA Latreille, 1807 Subgenus AEGIALIA Latreille, 1807 arenaria (Fabricius, 1787) globosa (Kugelann, 1794)
Aegialia (Aegialia) arenaria (Fabricius, 1787)
3 WATSONIAN (13,8,0,0,0) A coastal species of dunes and 2 YORKSHIRE beaches, and presumably associated with plant roots. (iv-ix) 1
NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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Subgenus PSAMMOPORUS Thomson, C.G., 1859 insularis Pittino, 2006 sabuleti sensu auctt. Brit. non (Panzer, 1797)
Aegialia (Psammoporus) insularis Pittino, 2006
3 WATSONIAN (1,6,2,3,3) Notable B. Scarce, widely and thinly distributed. Recorded from pitfalls 2 YORKSHIRE and beneath stones on sandy river banks. 1 (iv-ix). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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Subgenus RHYSOTHORAX Bedel, 1911 rufa (Fabricius, 1792) rufina Silfverberg, 1977
NO RECORDS ON THE DATABASE
Subfamily APHODIINAE Leach, 1815
Tribe APHODIINI Leach, 1815
APHODIUS Hellwig, 1798
Subgenus ACROSSUS Mulsant, 1842 depressus (Kugelann, 1792)
Aphodius (Acrossus) depressus (Kugelann, 1792)
3 WATSONIAN (11,11,35,11,4) Widespread and distributed 2 YORKSHIRE from high pastures in the north and west to sea level. Recorded from dung of sheep and 1 cow. (v-ix, peak vi) NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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12 | P a g e luridus (Fabricius, 1775) gagates (Geoffroy, 1764) variegatus (Herbst, 1783) nigrosulcatus (Marsham, 1802)
Aphodius (Acrossus) luridus (Fabricius, 1775)
3 WATSONIAN (4,5,13,2,0) Rather scarce in Yorkshire. 2 YORKSHIRE Widely distributed in sheep dung, wet straw 1 in pasture and in a pitfall trap in open fields.
NY NZ NZ OV Only a single recent record. (v-viii). 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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rufipes (Linnaeus, 1758) muticus (Stephens, 1830)
Aphodius (Acrossus) rufipes (Linnaeus, 1758)
3 WATSONIAN (219,19,107,56,241) Widespread and very 2 YORKSHIRE common, recorded from dung of horse, sheep and cow, from pitfall traps and in wet 1 straw in middens. The very large numbers of 0 NY NZ NZ OV SD SE SE TA records from VCs61, 63 and 65 result from 9 mv moth trap catches - this species is active 8 after dark. (v-x, peak vii-ix). 7 6 5 4 3 2 1
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Subgenus AGOLIINUS Schmidt, 1913
lapponum Gyllenhal, 1808 axillaris Stephens, 1839 subalpinus Hardy, 1847
Aphodius (Agoliinus) lapponum Gyllenhal, 1808
3 WATSONIAN (0,6,23,7,2) A boreo-montane and northern species which is widespread in suitable 2 YORKSHIRE habitats. There are few recent records 1 indicating a probable decline. Only recorded 0 NY NZ NZ OV SD SE SE TA from sheep dung in Yorkshire. (iii-x). 9 8 7 6 5 4 3 2 1
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nemoralis Erichson, 1848 Subgenus AGRILINUS Mulsant & Rey, 1870
NO RECORDS ON THE DATABASE
ater (De Geer, 1774) terrestris (Fabricius, 1775) obscurus (Marsham, 1802) non (Fabricius, 1792) terrenus Stephens, 1830
Aphodius (Agrilinus) ater (De Geer, 1774)
3 WATSONIAN (21,22,70,22,10) Widespread and fairly common throughout our recording area, on 2 YORKSHIRE farms, footpaths and in open pasture, and 1 noted from dung of cow, sheep and horse, 0 NY NZ NZ OV SD SE SE TA and occasionally in compost heaps. (iii-xii, 9 peak vi). 8 7 6 5 4 3 2 1
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14 | P a g e constans Duftschmid, 1805 nitidus Stephens, 1830
Aphodius (Agrilinus) constans Duftschmid, 1805
3 WATSONIAN (0,5,3,5,3) Very scarce. The only recent 2 YORKSHIRE records have come from the Little Don Valley area (SK1899) of VC63. Noted from 1 cow dung. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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rufus (Moll, 1782) scybalarius (Fabricius, 1781) rufescens (Fabricius, 1801) castaneus (Marsham, 1802) unicolor (Marsham, 1802) ochraceus Stephens, 1830
Aphodius (Agrilinus) rufus (Moll, 1782)
3 WATSONIAN (13,6,26,4,16) Widely distributed mainly in 2 YORKSHIRE the centre and south of the region. In dung of cow, sheep and horse, and often turning 1 up in light traps. Also recorded in a pitfall 0 NY NZ NZ OV SD SE SE TA trap in a stand of Eriophorum on large area 9 of milled peat on Hatfield Moor (SE6904). 8 (vi-xi, peak viii). 7 6 5 4 3 2 1
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15 | P a g e sordidus (Fabricius, 1775) Subgenus AMMOECIUS Mulsant, 1842
Aphodius (Agrilinus) sordidus (Fabricius, 1775)
3 WATSONIAN (0,1,4,0,0) Notable A. Very rare in our region with only five records on the YNU 2 YORKSHIRE database: Redcar area (NZ62) no date, MLT; 1 Blackmoorfoot (SE0912), 7/6/1992, MLD; 0 NY NZ NZ OV SD SE SE TA Dalton (SE11), 3/1963, EWA, at light; Honley 9 Wood (SE1111), 1984,DM; Richmond Park 8 (SK4092), 1987, WAE. The Redcar record is 7 undated (will be pre-1900) and therefore 6 not mapped. The Dalton record requires 5 confirmation as this species is diurnal and unlikely to come to light – may be a 4 misidentification of A. rufus (DMa pers. 3 comm. to RJM). 2 1
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NO RECORDS ON THE DATABASE
There was a record for Aphodius brevis on the YNU database (Arthington SE2745, 1956, JHF) but now removed. This has been shown to be a misidentification (see Jessop 1986:21) and is probably referrable to Aegialia insularis.
fimetarius (Linnaeus, 1758)
Aphodius (Aphodius) fimetarius (Linnaeus, 1758)
3 WATSONIAN (9,17,45,5,5) Widespread in dung of horse, 2 YORKSHIRE cow and sheep. See comment under pedellus . Most records of 1 fimetarius/pedellus species agg. in Yorkshire 0 NY NZ NZ OV SD SE SE TA will be pedellus. Data so far suggest that 9 fimetarius is a southern insect mainly from 8 sand dune systems or areas where there are 7 locally higher temperatures (DMa pers. 6 comm. to RJM). (ii-x). 5 4 3 2 1
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16 | P a g e foetens (Fabricius, 1787) scrutator sensu (Marsham, 1802) non (Herbst,1789) aestivalis Stephens, 1839
Aphodius (Aphodius) foetens (Fabricius, 1787)
3 WATSONIAN (4,6 ,12,5,0) Rather scarce and very thinly distributed across most of the county. 2 YORKSHIRE Recorded from dung of horse, pony and cow, 1 also from straw debris from a stable, and 0 NY NZ NZ OV SD SE SE TA once in a water trap in a rural garden. (v-ix). 9 8 7 6 5 4 3 2 1
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foetidus (Herbst, 1783) scybalarius sensu auctt. non (Fabricius, 1781) coprinus (Marsham, 1802)
Aphodius (Aphodius) foetidus (Herbst, 1783)
3 WATSONIAN (5,3,0,0,1) Rare in Yorkshire with only nine known records. Noted from the dung of 2 YORKSHIRE sheep and cow. 1
NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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17 | P a g e pedellus (De Geer, 1774) fimetarius sensu auctt. partim non (Linnaeus, 1758) Subgenus BODILUS Mulsant & Rey, 1870
Aphodius (Aphodius) pedellus (De Geer, 1774)
3 WATSONIAN (0,0,1,0,0) There is a problem of nomenclature here. The checklist of Duff 2 YORKSHIRE (2012) notes that pedellus (De Geer) is a 1 species currently confused with fimetarius 0 NY NZ NZ OV SD SE SE TA (Linnaeus), the status of pedellus being 9 disputed among Continental workers. The 8 Duff (2012) checklist follows Lobl & Smetana 7 (2006) and includes pedellus as a valid 6 species. We have a single record of pedellus: 5 Blackmoorfoot (SE1012), 26/3/2012, MLD 4 teste DMa. Also see comments under fimetarius above. It may be that some or 3 indeed many of our 'fimetarius' records may 2 be pedellus. Without vouchers we will not 1 know. SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1989 Map produced for Yorkshire Naturalists' Union by DMAP 17/4/2016 pre 31/12/1989 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4
ictericus (Laicharting, 1781) nitidulus (Fabricius, 1792)
Aphodius (Bodilus) ictericus (Laicharting, 1781)
3 WATSONIAN (2,0,0,1,0) We have three records of this 2 YORKSHIRE county rarity: Copgrove (SE36) 1885, YNU; Hull (TA13), unknown date, TS; Spurn 1 (TA41), 1914, GBW. The Hull record is 0 NY NZ NZ OV SD SE SE TA undated and therefore not mapped. No 9 habitat details were given in these early 8 records, the species being associated with 7 dry grasslands, mostly in sheep dung, but 6 also of horse and cow (DMa pers. comm. to 5 RJM). 4 3 2 1
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18 | P a g e
Subgenus CALAMOSTERNUS Motschulsky, 1860 granarius (Linnaeus, 1767) emarginatus Stephens, 1830 haemorrhous Stephens, 1830 lucens Stephens, 1830 melanopus Stephens, 1830
Aphodius (Calomosternus) granarius (Linnaeus, 1767)
3 WATSONIAN (9,6,10,2,1) Widespread mainly in the 2 YORKSHIRE southern half of the county, but scarce. Recorded from dung (animal not noted), in 1 flood refuse, in ground litter in a bed of 0 NY NZ NZ OV SD SE SE TA Phragmites, on a rabbit carcass, in a manure 9 heap, in grass heaps, and on one occasion 8 numerous under a dead water vole on sand 7 at a roadside. (iii-ix). 6 5 4 3 2 1
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Subgenus CHILOTHORAX Motschulsky, 1860 conspurcatus (Linnaeus, 1758)
Aphodius (Chilothorax) conspurcatus (Linnaeus, 1758)
3 WATSONIAN (0,3,5,6,0) Notable B. Scarce with no recent records. Habitat details not noted in any of 2 YORKSHIRE our records, but the generally accepted 1 preference is for dung (mainly of horse) in 0 NY NZ NZ OV SD SE SE TA unexposed habitats such as woodland. 9 (iii-xii). 8 7 6 5 4 3 2 1
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19 | P a g e distinctus (Müller, O.F., 1776) inquinatus (Herbst, 1783) attaminatus (Marsham, 1802) foedatus (Marsham, 1802)
Aphodius (Chilothorax) distinctus (Müller, O.F., 1776)
3 WATSONIAN (5,4,8,2,0) Notable B. A scarce species of the 2 YORKSHIRE dung of various large herbivores and in compost heaps. Also noted from dung of 1 horse and badger, and once from a mole's 0 NY NZ NZ OV SD SE SE TA nest. (iii-x). Prefers sandy soils or calcareous 9 grassland. Records away from such areas 8 require confirmation (DMa pers. comm. to 7 RJM). 6 5 4 3 2 1
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paykulli Bedel, 1907 inquinatus sensu (Olivier, 1789) non (Herbst, 1783) tessulatus (Paykull, 1798) non (Laicharting, 1781)
Aphodius (Chilothorax) paykulli Bedel, 1907
3 WATSONIAN (1,0,7,0,0) Notable B. Very scarce with no recent records. Recorded from a manure 2 YORKSHIRE heap and once from a glasshouse. Habitat 1 details lacking in most of our records. Our 0 NY NZ NZ OV SD SE SE TA records are dated late autumn, winter and 9 early spring. Associated with well-drained 8 soils, calcareous grasslands etc.. (DMa pers. 7 comm. to RJM). (xi-iv). 6 5 4 3 2 1
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20 | P a g e
Subgenus COLOBOPTERUS Mulsant, 1842 erraticus (Linnaeus, 1758) Subgenus ESYMUS Mulsant & Rey, 1870
Aphodius (Colobopterus) erraticus (Linnaeus, 1758)
3 WATSONIAN (3,3,0,1,0) Very scarce in dung of horse and cow, and in flood refuse, mainly in open 2 YORKSHIRE situations. There have been no known 1 records received since 1951. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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merdarius (Fabricius, 1775)
Aphodius (Esymus) merdarius (Fabricius, 1775)
3 WATSONIAN (3,4,3,1,0) Very scarce and widely distributed except in the far west of the county. No 2 YORKSHIRE habitat details accompany any of the 1 Yorkshire records, of which there have been 0 NY NZ NZ OV SD SE SE TA none since 1948. We have a record from an 9 archaeological site of medieval date (Cowick 8 Moat SE652205). 7 6 5 4 3 2 1
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21 | P a g e pusillus (Herbst, 1789) phaeopterus Stephens, 1830
Aphodius (Esymus) pusillus (Herbst, 1789)
3 WATSONIAN (2,2,8,4,2) Widely distributed but scarce with 2 YORKSHIRE only eighteen county records, only two of which are recent. In Yorkshire recorded from 1 sheep dung and a pile of wet straw in a 0 NY NZ NZ OV SD SE SE TA pasture. 9 8 7 6 5 4 3 2 1
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Subgenus EUORODALUS Dellacasa, G., 1983 coenosus (Panzer, 1798) tristis Panzer, 1833
Aphodius (Euorodalus) coenosus (Panzer, 1798)
3 WATSONIAN (0,0,1,0,0). Notable B. We have a single record on the YNU database: Rossington 2 YORKSHIRE (SK69), 1900, HHC. This record is on a card 1 index at Doncaster Museum. Specimens 0 NY NZ NZ OV SD SE SE TA exist in Doncaster Museum collections 9 labelled Rossington, May 1908 with no 8 collector (but presumably HH Corbett). The 7 species is associated with sandy soils (DMa 6 pers. comm. to RJM). 5 4 3 2 1
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22 | P a g e
Subgenus EUPLEURUS Mulsant, 1842 subterraneus (Linnaeus, 1758) Subgenus LABARRUS Mulsant & Rey, 1870
Aphodius (Eupleurus) subterraneus (Linnaeus, 1758)
3 WATSONIAN (1,1,4,2,0) RedList_pre94-NR. Very few records, the most recent being dated 2 YORKSHIRE 1951. No habitat details are provided with 1 these records except that from Spurn - "in 0 NY NZ NZ OV SD SE SE TA Phragmites". 9 8 7 6 5 4 3 2 1
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lividus (Olivier, 1789) bilituratus (Marsham, 1802) Subgenus LIMARUS Mulsant & Rey, 1870
NO RECORDS ON THE DATABASE
zenkeri Germar, 1813 Subgenus LIOTHORAX Motschulsky, 1860
Aphodius (Limarus) zenkeri Germar, 1813
3 WATSONIAN (1,0,3,0,0). Notable B. There are four records on the YNU database: Allerthorpe 2 YORKSHIRE Common (SE7647), 5/8/1989, RJM, in 1 horse dung; Scabba Wood (SE5201), 0 NY NZ NZ OV SD SE SE TA 5/7/2004, RJM, by general sweeping low 9 on the ground where roe deer were seen 8 regularly; Hatfield Moor (SE7006), 7/1991, 7 PS; Hatfield Moor (SE70) 8/1991, PS. A 6 deer dung specialist but can occur in other 5 dung where deer are present (Jessop 4 1986:23), viz. the Allerthorpe record. 3 2 1
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23 | P a g e niger (Illiger, 1798) niger (Panzer, 1797)
NO RECORDS ON THE DATABASE
plagiatus (Linnaeus, 1767) elongatus Stephens, 1830 Subgenus MELINOPTERUS Mulsant, 1842
Aphodius (Liothorax) plagiatus (Linnaeus, 1767)
3 WATSONIAN (4,0,0,0,0) Notable B. Very scarce with only 2 YORKSHIRE four records on the YNU database: Aughton Church (SE7038), 3/4/1993, PK; 1 Cherry Cob Sands (TA2121), 1980, RSK; 0 NY NZ NZ OV SD SE SE TA Spurn (TA4115), 20/6/1951, WDH, 9 Naturalist 1952:138); Spurn (TA4212), 8 9/7/2001, MLD, a single in a pitfall trap at 7 Narrow Neck. A coastal species associated 6 with temporary pool margins and streams, 5 the inland record may be a 4 misidentification of A. granarius (DMa pers. comm. to RJM). 3 2
1
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consputus Creutzer, 1799
NO RECORDS ON THE DATABASE
prodromus (Brahm, 1790)
Aphodius (Melinopterus) prodromus (Brahm, 1790)
3 WATSONIAN (19,20,70,15,5) A widespread and fairly 2 YORKSHIRE common species over much of Yorkshire. Noted from compost heaps, and the dung 1 of sheep, horse, dog and deer, and once 0 NY NZ NZ OV SD SE SE TA from dung of badger. We have no records 9 on the database that specify cow dung. 8 (iii-xii with a broad spring peak). 7 6 5 4 3 2 1
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24 | P a g e punctatosulcatus Sturm, 1805 sabulicola Thomson, C.G., 1868
NO RECORDS ON THE DATABASE
sphacelatus (Panzer, 1798) marginellus Stephens, 1830 punctatosulcatus sensu auctt. Brit. non Sturm, 1805 Subgenus NIMBUS Mulsant & Rey, 1870
Aphodius (Melinopterus) sphacelatus (Panzer, 1798)
3 WATSONIAN (18,,8,78,8,6) Widespread and common 2 YORKSHIRE throughout most of Yorkshire, and recorded from dung of horse, sheep and cow, from 1 piles of damp straw, once from a dead 0 NY NZ NZ OV SD SE SE TA pheasant, and from pitfall traps in pastures 9 and grassland. Sometimes very abundant 8 when found. (i-xii, peak iv,v). 7 6 5 4 3 2 1
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contaminatus (Herbst, 1783) ciliaris (Marsham, 1802)
Aphodius (Nimbus) contaminatus (Herbst, 1783)
3 WATSONIAN ( 11,12,48,10,4) Widespread and fairly 2 YORKSHIRE common across most of Yorkshire, and recorded from dung of cow, sheep, horse 1 and dog, often numerous when found. This 0 NY NZ NZ OV SD SE SE TA is an autumn species – June records are 9 very early and may require confirmation 8 with vouchers. (vi-xi, peak vi) 7 6 5 4 3 2 1
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25 | P a g e obliteratus Sturm, 1823
Aphodius (Nimbus) obliteratus Panzer, 1823
3 WATSONIAN ( 1,2,2,7,0 ) Very scarce with a distribution 2 YORKSHIRE across the centre of the county. Recorded from sheep dung. The species prefers 1 scrub/woodland and the dung of sheep 0 NY NZ NZ OV SD SE SE TA and horse, but rarely cow (DMa pers. 9 comm. to RJM). (vii-x). 8 7 6 5 4 3 2 1
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Subgenus OTOPHORUS Mulsant, 1842
haemorrhoidalis (Linnaeus, 1758) sanguinolentus (Marsham, 1802) Subgenus PHALACRONOTHUS Motschulsky, 1860
Aphodius (Otophorus) haemorrhoidalis (Linnaeus, 1758)
3 WATSONIAN (6,5,10,2,1) Rather scarce with most records coming from the central part of 2 YORKSHIRE the county. Our records indicate a 1 preference for horse dung, with dung of 0 NY NZ NZ OV SD SE SE TA cow on one occasion. This species is a 9 cattle dung specialist but also found in 8 horse/sheep dung but usually with cattle 7 nearby (DMa pers comm. to RJ M). (v-viii). 6 5 4 3 2 1
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quadrimaculatus (Linnaeus, 1761) Subgenus PLAGIOGONUS Mulsant, 1842
NO RECORDS ON THE DATABASE
26 | P a g e arenarius (Olivier, 1789) putridus (Geoffroy in Fourcroy, 1785) rhododactylus (Marsham, 1802)
Aphodius (Plagiogonus) arenarius (Oliv ier, 1789)
3 WATSONIAN (1,0,4,2,1) Notable B. All our eight records were originally recorded under the name 2 YORKSHIRE of putridus (Fourcroy, 1785), and most are 1 recorded from upland areas. Sheep dung is 0 NY NZ NZ OV SD SE SE TA mentioned in only a single record. The 9 distribution shown on NBN Gateway 8 (https://data.nbn.org.uk/Taxa/NHMSYS00 7 01716971 - accessed 26/4/2016) is at odds 6 with the comments in Jessop 1986:21). [All 5 our arenarius records are referable either to borealis or fasciatus. Nomenclatural 4 errors in the past make most records of 3 this species difficult to assess without 2 vouchers. A. arenarius is an insect of 1 calcareous grasslands and chalk to sandy 0 SD SE SE TA SJ SK SK TF soils in the south of the country (DMa pers. 9 comm. to RJM)] 8 post 31/12/1989 Map produced for Yorkshire Naturalists' Union by DMAP 17/4/2016 pre 31/12/1989 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4
Subgenus PLANOLINUS Mulsant & Rey, 1870
borealis Gyllenhal, 1827 putridus sensu (Sturm, 1805) non (Geoffroy in Fourcroy, 1785)
Aphodius (Planolinus) borealis Gyllenhal, 1827
3 WATSONIAN (0,5,5,2,2) Few records, only two being recent, with a northern and western 2 YORKSHIRE distribution. There are no habitat details 1 attached to any of our records for this 0 NY NZ NZ OV SD SE SE TA species. (iv-viii). 9 8 7 6 5 4 3 2 1
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27 | P a g e fasciatus (Olivier, 1789) putridus sensu (Herbst, 1789) non (Geoffroy in Fourcroy, 1785) foetidus (Fabricius, 1792) non (Herbst, 1783) uliginosus Hardy, 1847 tenellus sensu auctt. non Say, 1823
Aphodius (Planolinus) fasciatus (Oliv ier, 1789)
3 WATSONIAN (0,8,4,1,3) Notable B. Recorded from the north and west of the county, often from 2 YORKSHIRE sheep dung. (v-x). 1
NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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Subgenus SIGORUS Mulsant & Rey, 1870 porcus (Fabricius, 1792) turpis (Marsham, 1802) ruficrus (Marsham, 1802)
Aphodius (Sigorus) porcus (Fabricius, 1792)
3 WATSONIAN (0,1,0,1,0) Notable B. Rare, with only two 2 YORKSHIRE records on the database: Hetchell Wood (SE3842), 1974, JHF; Strensall Common 1 (SE6559), 18/7/1971, JHF. No other details 0 NY NZ NZ OV SD SE SE TA are given with these records. 9 8 7 6 5 4 3 2 1
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28 | P a g e
Subgenus TEUCHESTES Mulsant, 1842 fossor (Linnaeus, 1758)
Aphodius (Teuchestes) fossor (Linnaeus, 1758)
3 WATSONIAN (20,21,44,8,9) Widespread and fairly 2 YORKSHIRE common in cow dung in pastures, but also recorded from dung of sheep, deer and 1 horse, and from pitfall traps in farmland. 0 NY NZ NZ OV SD SE SE TA (v-ix). 9 8 7 6 5 4 3 2 1
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Subgenus VOLINUS Mulsant & Rey, 1870 sticticus (Panzer, 1798) equestris (Panzer, 1798)
Aphodius (Volinus) sticticus (Panzer, 1798)
3 WATSONIAN (1,0,1,3,1) Very scarce with only six records 2 YORKSHIRE on the YNU database. Reported from horse dung and flood refuse. 1
NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1
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29 | P a g e
EUHEPTAULACUS Dellacasa, G., 1983 HEPTAULACUS sensu auctt. non Mulsant, 1842 sus (Herbst, 1783)
NO RECORDS ON THE DATABASE
villosus (Gyllenhal in Schönherr, 1806)
Euheptaulacus v illosus (Gyllenhal in Schönherr, 1806)
3 WATSONIAN (0,1,1,2,0) Notable A. We have four 2 YORKSHIRE records: Hetchell Crags (SE3742), 7/1958, JHF, on sandy ground; Hetchell Wood 1 (SE3742), 1960, NR; Redcar (NZ62), 0 NY NZ NZ OV SD SE SE TA 24/6/1921, MLT, on coastal sandhills; 9 Anston Stones Wood SSSI (SK5383), 8 7/1979, NR det CJ. 7 6 5 4 3 2 1
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HEPTAULACUS Mulsant, 1842 testudinarius (Fabricius, 1775)
Heptaulacus testudinarius (Fabricius, 1775)
3 WATSONIAN (0,2,0,2,0) RedList_pre94-NR. A scarce 2 YORKSHIRE species with only four, all very old, records known from Yorkshire: Adel dam (SE2740), 1 7/1876, HHC; Wetherby area (SE44), 1879, 0 NY NZ NZ OV SD SE SE TA YNU; Whitby (NZ81), 1879, HP; York 9 (SE65), unknown date, WCH (record 8 unmapped). Vouchers would be required 7 to confirm these records – this species is 6 sometimes misidentified for villosus (DMa 5 pers. comm. to RJM). 4 3 2 1
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30 | P a g e
OXYOMUS Dejean, 1833 sylvestris (Scopoli, 1763) silvestris auctt. (misspelling) porcatus (Fabricius, 1775) platycephalus (Marsham, 1802)
Oxyomus sylv estris (Scopoli, 1763)
3 WATSONIAN (0,2,2,0,0) Very scarce with four known records: Scarborough (TA08), c1900, RL; 2 YORKSHIRE Whitby (NZ81), 1936, HB; Thorne Moor 1 (SE71), 1910, Anon; Thorne Moor (SE71), 0 NY NZ NZ OV SD SE SE TA 6/1911, WES. No habitat details are given 9 in our records but the species is associated 8 with rotting vegetation, dung heaps and 7 carrion. 6 5 4 3 2 1
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Tribe EUPARIINI Schmidt, 1910
SAPROSITES Redtenbacher, 1858 mendax (Blackburn, 1892) parallelus sensu Champion, 1921 non Harold, 1867
NO RECORDS ON THE DATABASE natalensis (Peringuey, 1901)
NO RECORDS ON THE DATABASE
Tribe PSAMMODIINI Mulsant, 1842
Subtribe PSAMMODIINA Mulsant, 1842
BRINDALUS Landin, 1960 porcicollis (Illiger, 1803)
NO RECORDS ON THE DATABASE
DIASTICTUS Mulsant, 1842 vulneratus (Sturm, 1805)
NO RECORDS ON THE DATABASE
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PSAMMODIUS Fallén, 1807 PSAMMOBIUS Heer, 1841 asper (Fabricius, 1775) sulcicollis (Illiger, 1802)
Psammodius asper (Fabricius, 1775)
3 WATSONIAN (0,1,0,0,0) RedList_pre94-EN. There is a 2 YORKSHIRE single record on the YNU database: Scarborough (TA08) c.1900, RL. I have not 1 traced a voucher for this record. The 0 NY NZ NZ OV SD SE SE TA species is associated with coastal sandy 9 locations. 8 7 6 5 4 3 2 1
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TESARIUS Rakovic, 1981 PSAMMODIUS sensu auctt. partim non Fallén, 1807 caelatus (LeConte, 1857)
NO RECORDS ON THE DATABASE mcclayi (Cartwright, 1955)
NO RECORDS ON THE DATABASE
Subtribe RHYSSEMINA Pittino & Mariani, 1986
PLEUROPHORUS Mulsant, 1842 PSAMMOBIUS sensu auctt. partim non Heer, 1841 caesus (Creutzer in Panzer, 1796)
NO RECORDS ON THE DATABASE
RHYSSEMUS Mulsant, 1842
germanus (Linnaeus, 1767)
NO RECORDS ON THE DATABASE
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Subfamily SCARABAEINAE Latreille, 1802
Tribe COPRINI Leach, 1815
COPRIS Geoffroy, 1762 lunaris (Linnaeus, 1758)
NO RECORDS ON THE DATABASE
Tribe ONTHOPHAGINI Burmeister, 1846
ONTHOPHAGUS Latreille, 1802
Subgenus ONTHOPHAGUS Latreille, 1802 taurus (Schreber, 1759)
NO RECORDS ON THE DATABASE
Subgenus PALAEONTHOPHAGUS Zunino, 1979 coenobita (Herbst, 1783)
NO RECORDS ON THE DATABASE fracticornis (Preyssler, 1790) similis sensu auctt. non (Scriba, 1790)
NO RECORDS ON THE DATABASE (but see similis)
joannae Goljan, 1953 ovatus sensu auctt. partim non (Linnaeus, 1767)
Onthophagus (Paleonthophagus) j oannae Golj an, 1953
3 WATSONIAN (0,0,0,1,0) A single record exists on the 2 YORKSHIRE YNU database: Grass Wood Reserve (SD9865) 3/9/1985, RBA det CJ. A single 1 specimen was recorded from dung in 0 NY NZ NZ OV SD SE SE TA meadow on bank of the River Wharfe. 9 8 7 6 5 4 3 2 1
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medius (Kugelann, 1792) vacca sensu auctt. partim non (Linnaeus, 1767)
Onthophagus (Paleonthophagus) v acca (Linnaeus, 1767)
3 WATSONIAN (0,0,1,0,0) Notable B. There is a single record of this species on the YNU 2 YORKSHIRE database: Knottingley (SE42) 1918, 1 WJF. This record is a doubtful one in NY NZ NZ OV 0 SD SE SE TA the absence of the specimen. See 9 Jessop (1986:27) - known distribution 8 is south of London-Bristol line. The record is therefore unlikely given the 7 stated habitat, although the record 6 appears on the card index at Doncaster 5 Museum. The specimen was reported 4 from a manure heap: [an unlikely 3 habitat for an Onthophagus (DMa pers. comm. to RJM)]. 2 1
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nuchicornis (Linnaeus, 1758) verticicornis (Fabricius, 1775) xiphias (Fabricius, 1792) dillwynii (Stephens, 1830)
Onthophagus (Paleonthophagus) nuchicornis (Linnaeus, 1758)
3 WATSONIAN (0,1,0,0,0) Notable A. There is a single record of this species on the YNU database: 2 YORKSHIRE Sandburn Wood (SE6758), pre-1900, WCH. 1 [This record is old and will certainly be 0 NY NZ NZ OV SD SE SE TA referable to O. similis as the two names 9 similis and nuchicornis were mixed up in 8 early literature (DMa pers. comm. to RJM)]. 7 6 5 4 3 2 1
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34 | P a g e similis (Scriba, 1790) fracticornis sensu auctt. Brit. non (Preyssler, 1790)
Onthophagus (Paleonthophagus) similis (Scriba, 1790)
3 WATSONIAN (0,0,3,0,0) Two records appear in the YNU 2 YORKSHIRE database: Rossington (SK69), 18/4/1912, HHC; Thorne Moor (SE71), 14/5/1903, 1 HHC. Specimens are in the Doncaster 0 NY NZ NZ OV SD SE SE TA Museum. There is a third record from an 9 archaeological site of medieval date 8 (Cowick Moat, SE6520). 7 6 5 4 3 2 1
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verticicornis (Laicharting, 1781) nutans (Fabricius, 1787)
NO RECORDS ON THE DATABASE
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Subfamily MELOLONTHINAE Leach, 1819
Tribe HOPLIINI Latreille, 1829
HOPLIA Illiger, 1803
Subgenus DECAMERA Mulsant, 1842 philanthus (Füessly, 1775) farinosa sensu auctt. non (Linnaeus, 1761)
Hoplia philanthus (Füessly, 1775)
3 WATSONIAN (0,5,5,10,3) Widespread but scarce, not recorded from the far east or west of the 2 YORKSHIRE region. Recorded from grassland, 1 hawthorn blossom, in pitfalls and water 0 NY NZ NZ OV SD SE SE TA traps by rivers, in lawn turf and once in a 9 water butt in a suburban garden. 8 Apparently becoming more frequent. 7 6 5 4 3 2 1
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Tribe MELOLONTHINI Leach, 1819
Subtribe MELOLONTHINA Leach, 1819
MELOLONTHA Fabricius, 1775 hippocastani Fabricius, 1801
NO RECORDS ON THE DATABASE
melolontha (Linnaeus, 1758) vulgaris Fabricius, 1775
Melolontha melolontha (Linnaeus, 1758)
3 WATSONIAN (6,14,41,32,106) Widespread throughout our recording area from the coastal 2 YORKSHIRE lowlands to the higher ground in the west. 1 The high number of localised VC64 and 0 NY NZ NZ OV SD SE SE TA VC65 records reflects recording effort in 9 several locations where light trapping is 8 carried out regularly. Taken regularly in 7 light traps but occasionally in the sweep 6 net, seen by roadsides and in sandy soil. 5 Known to be declining in many areas of Britain. (iv-x). 4 3
2 1
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POLYPHYLLA Harris, 1842 fullo (Linnaeus, 1758)
Polyphylla fullo (Linnaeus, 1758)
3 WATSONIAN (0,0,1,0,0) RedList_pre94-EX. There is a single record on the YNU database: 2 YORKSHIRE Doncaster (SE50), 15/7/2000, DS det RJM, 1 a single specimen in a wooden crate used 0 NY NZ NZ OV SD SE SE TA to import a television set, specimen 9 provenance unknown. The species has an 8 association with conifers, the adults 7 feeding on needles and the larvae feeding 6 on roots of ground vegetation. Generally 5 recorded as a vagrant in Britain. 4 3 2 1
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Subtribe RHIZOTROGINA Burmeister, 1855
AMPHIMALLON Latreille, 1825 AMPHIMALLUS Mulsant, 1842 (misspelling) RHIZOTROGUS sensu auctt. partim non Berthold, 1827 fallenii (Gyllenhal, 1817) ochraceum sensu auctt. partim non (Knoch, 1801)
NO RECORDS ON THE DATABASE
38 | P a g e solstitiale (Linnaeus, 1758)
Amphimallon solstitiale (Linnaeus, 1758)
3 WATSONIAN (2,4,1,0,0) Very scarce in Yorkshire with 2 YORKSHIRE only 7 known records. Recorded from a light trap, once in an owl pellet, and in 1 flight at dusk. Only three recent records. 0 NY NZ NZ OV SD SE SE TA (vii-viii). 9 8 7 6 5 4 3 2 1
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Family EUCINETIDAE Lacordaire, 1857
Tribe SERICINI Kirby, 1837
OMALOPLIA Schönherr, 1817 HOMALOPLIA Erichson, 1847 ruricola (Fabricius, 1775) varius (Marsham, 1802)
NO RECORDS ON THE DATABASE
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SERICA MacLeay, 1819 brunnea (Linnaeus, 1758)
Serica brunnea (Linnaeus, 1758)
3 WATSONIAN (10,9,42,26,65) Widespread and fairly common, especially at light on warm 2 YORKSHIRE evenings after hot weather during the day. 1 Many records from VCs 64 and 65 are from 0 NY NZ NZ OV SD SE SE TA moth traps. Also recorded from beneath 9 stones, around house lights and by 8 sweeping in grassland. The larvae feed on 7 plant roots and may have a two-year 6 development period. (iv-ix). 5 4 3 2 1
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Subfamily RUTELINAE MacLeay, 1819
Tribe ANOMALINI Streubel, 1839
ANOMALA Samouelle, 1819 EUCHLORA MacLeay, 1819 dubia (Scopoli, 1763) aenea (De Geer, 1774) frischii (Fabricius, 1775)
Anomala dubia (Scopoli, 1763)
3 WATSONIAN (19,4,1,0,0) A scarabaeid with a very 2 YORKSHIRE limited distribution in the Spurn area and in the area around Coatham Sands. Found 1 in the roots of plants in the sand and 0 NY NZ NZ OV SD SE SE TA sometimes beaten and swept from foliage. 9 Also taken in pitfall traps. There is a single 8 inland record for "Doncaster" dated 1921 7 (EGB), which would require confirmation 6 as the species is associated with sandy 5 soils (of which there are plenty of sites on 4 the east side of Doncaster). Nationally, nearly all records are coastal or estuarine. 3 (vi-viii). 2 1
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PHYLLOPERTHA Stephens, 1830 horticola (Linnaeus, 1758) arvicola sensu (Marsham, 1802) non (Fabricius, 1775)
Phyllopertha horticola (Linnaeus, 1758)
3 WATSONIAN (13,4,16,1,0) Widespread but not common, mainly in the south and east of 2 YORKSHIRE our recording area. The species feeds at 1 the roots of plants, particularly in turf. Also 0 NY NZ NZ OV SD SE SE TA recorded from nettle beds and by beating 9 the foliage of various trees. (iv-vii). 8 7 6 5 4 3 2 1
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Subfamily CETONIINAE Leach, 1815
Tribe CETONIINI Leach, 1815
CETONIA Fabricius, 1775 aurata (Linnaeus, 1761)
Cetonia aurata (Linnaeus, 1758)
3 WATSONIAN (1,3,1,0,0,) There are five records for this 2 YORKSHIRE species on the YNU database: Helmsley (SE68) 1884, HWE; Langdale End (SE9492) 1 date unknown, recorder noted as "Pittam"; 0 NY NZ NZ OV SD SE SE TA West Ayton (SE9884) 6/1952, AMR; 9 Hunmanby (TA07) GBW; Conisbrough 8 (SK5198) date unknown, PB, specimen 7 fragments from an archaeological site of 6 medieval date. Recorded from roses and 5 various fruit trees. 4 3 2 1
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PROTAETIA Burmeister, 1842 CETONIA sensu auctt. partim non Fabricius, 1775
Subgenus POTOSIA Mulsant & Rey, 1870
metallica (Herbst, 1782) cuprea sensu auctt. Brit. non (Fabricius, 1775) floricola (Herbst, 1790)
Protaetia (Potosia) metallica (Herbst, 1782)
3 WATSONIAN (0,1,0,0,0) Notable B. There is a single 2 YORKSHIRE record on our database: Ellers Wood (SE5292) 1984, RGB, see Naturalist 1 1988:71. Apparently a northern species 0 NY NZ NZ OV SD SE SE TA with a peculiar life history, the larva being 9 associated with ant nests and the adult on 8 various fruit trees. 7 6 5 4 3 2 1
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Tribe TRICHIINI Fleming, 1821
GNORIMUS Lepeletier & Audinet-Serville, 1828 ALEUROSTICTUS Kirby, 1827 nobilis (Linnaeus, 1758)
Gnorimus nobilis (Linnaeus, 1758)
3 WATSONIAN (0,0,1,0,0) RedList_pre94-EX. There is a 2 YORKSHIRE single record on the YNU database: Norton 1 Common (SE5615) 7/1970, PS, sub-fossil fragments found in a rot hole in a buried 0 NY NZ NZ OV SD SE SE TA bog oak. See Naturalist 1971:111. 9 8 7 6 5 4 3 2 1
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43 | P a g e variabilis (Linnaeus, 1758) octopunctatus (Fabricius, 1775)
NO RECORDS ON THE DATABASE
TRICHIUS Fabricius, 1775 fasciatus (Linnaeus, 1758)
NO RECORDS ON THE DATABASE gallicus Dejean, 1821 rosaceus (Voët, 1769) zonatus sensu auctt. Brit. non Germar, 1831 abdominalis sensu auctt. Brit. non (Ménétries, 1832)
NO RECORDS ON THE DATABASE
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References
Britton, E.B., 1956 - Coleoptera: Scarabaeoidea. Handbk. Ident. Br. Insects, 5(11):1-29. Royal Entomological Society, London.
Cooter, J., (1999) On “English‟ names for British Coleoptera. Coleopterist 8 (1): 43-45.
Dandy J.E. 1969. Watsonian Vice-counties of Great Britain. The Ray Society, London
Duff, A.G. (Ed.) 2008 - Checklist of the Beetles of the British Isles. A.G. Duff, Wells.
Duff, A.G., (Ed.) 2012 - Checklist of the Beetles of the British Isles, 2nd Edition. Pemberley Books, Iver.
Freude, H.,, Harde, K.W., & Lohse, G.A., 1969. Die Kafer Mitteleuropas. Band 8. Goecke & Evers, Krefeld.
Hyman P.S. and Parsons M.S. 1992. A review of the scarce and threatened Coleoptera of Great Britain, part 1. JNCC, Peterborough.
Hyman P.S. and Parsons M.S. 1994. A review of the scarce and threatened Coleoptera of Great Britain, part 2. JNCC, Peterborough.
Jessop, L., 1986 - Coleoptera: Scarabaeoidea. Handbk. Ident. Br. Insects 5(11)1-53. Royal Entomological Society, London.
Joy, N. H., 1932. A Practical Handbook of British Beetles. (2 volumes). Witherby, London. (Reprinted 1972, E.W. Classey, Faringdon).
Key, R.S., 1999. More on “noddy names‟ for British beetles. Coleopterist 8 (2): 94.
Lobl & Smetana 2006. Catalogue of Palaearctic Coleoptera. Vol. 3.Scarabaeoidea - Scirtoidea - Dascilloidea -Buprestoidea - Byrrhoidea. Stenstrup: Apollo Books, 690pp.
Pope, R.D. 1977. Kloet & Hincks. A Check List of British Insects. Part 3: Coleoptera and Strepsiptera. Second revised edition. Handbks. Ident. Brit. Insects 11(3), pp. xiv+105. Royal Entomological Society, London.
Watson, H.C., 1883. 2nd. ed. Topographical Botany. Quaritch, London.
Shirt, D.B. (Ed.) 1987. British Red Data Books: 2. Insects. Nature Conservancy Council, Peterborough.
Naturalist. Various issues of the journal of the Yorkshire Naturalists' Union.
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Acknowledgements
Thanks are due to Alan Morton for permission to use his DMAP mapping software; Paul Buckland for his inventory of the British material in the Doncaster Museum Coleoptera collection which has proved invaluable in tracking down voucher specimens; and also special thanks to Darren Mann of Oxford University Museum who offered help and advice and gave his time with aspects of the species descriptions in a draft of this part of the Atlas.
RJM 161130
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