GUIDE LEAFLET SERIES No. 70
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SMITHSONIAN MISCELLANEOUS COLLECTIONS [Vol
THE DINOSAUR TRACHODON ANNECTENS By R a. LUCAS The skeleton of TracJiodon, or Claosanrns, recently placed on exhi- bition in the U. S. National Museum, is an unusually perfect example of that group of extinct reptiles, the dinosaurs. It was included in the Marsh collection and was one of two nearly complete skeletons obtained by Mr. J. B. Hatcher some years ago on Lance creek, Wyoming. The completeness of the specimen is due to the fact that the animal was either engulfed in quicksand, and so came to his end, or that by some favorable accident, such as a cloudburst or a freshet, the body was otherwise covered with sand immediately after death, and before decomposition had set in. Whatever may have happened, the result was that the bones remained in place, the ribs being attached to their respective vertebrae and the great thigh bones re- maining in their sockets, the legs even having the position they would take in walking. This is shown in pi. lxxii, for in mounting the skeleton the ends of the thigh bones were left as found. Some ex- amples of Trachodon have been obtained in which the impression of the skin was preserved in the surrounding rock, and from these it is known that this animal was covered with small, irregularly six-sided, horny plates, somewhat like those covering portions of the bodies of crocodiles. Unfortunately the wearing away of the rock in which the present specimen was contained had exposed some of the bones, and portions of them had been damaged and the front of the skull weathered away before its discovery in the summer of 1891. -
Luis V. Rey & Gondwana Studios
EXHIBITION BY LUIS V. REY & GONDWANA STUDIOS HORNS, SPIKES, QUILLS AND FEATHERS. THE SECRET IS IN THE SKIN! Not long ago, our knowledge of dinosaurs was based almost completely on the assumptions we made from their internal body structure. Bones and possible muscle and tendon attachments were what scientists used mostly for reconstructing their anatomy. The rest, including the colours, were left to the imagination… and needless to say the skins were lizard-like and the colours grey, green and brown prevailed. We are breaking the mould with this Dinosaur runners, massive horned faces and Revolution! tank-like monsters had to live with and defend themselves against the teeth and claws of the Thanks to a vast web of new research, that this Feathery Menace... a menace that sometimes time emphasises also skin and ornaments, we reached gigantic proportions in the shape of are now able to get a glimpse of the true, bizarre Tyrannosaurus… or in the shape of outlandish, and complex nature of the evolution of the massive ornithomimids with gigantic claws Dinosauria. like the newly re-discovered Deinocheirus, reconstructed here for the first time in full. We have always known that the Dinosauria was subdivided in two main groups, according All of them are well represented and mostly to their pelvic structure: Saurischia spectacularly mounted in this exhibition. The and Ornithischia. But they had many things exhibits are backed with close-to-life-sized in common, including structures made of a murals of all the protagonist species, fully special family of fibrous proteins called keratin fleshed and feathered and restored in living and that covered their skin in the form of spikes, breathing colours. -
An Ankylosaurid Dinosaur from Mongolia with in Situ Armour and Keratinous Scale Impressions
An ankylosaurid dinosaur from Mongolia with in situ armour and keratinous scale impressions VICTORIA M. ARBOUR, NICOLAI L. LECH−HERNES, TOM E. GULDBERG, JØRN H. HURUM, and PHILIP J. CURRIE Arbour, V.M., Lech−Hernes, N.L., Guldberg, T.E., Hurum, J.H., and Currie P.J. 2013. An ankylosaurid dinosaur from Mongolia with in situ armour and keratinous scale impressions. Acta Palaeontologica Polonica 58 (1): 55–64. A Mongolian ankylosaurid specimen identified as Tarchia gigantea is an articulated skeleton including dorsal ribs, the sacrum, a nearly complete caudal series, and in situ osteoderms. The tail is the longest complete tail of any known ankylosaurid. Remarkably, the specimen is also the first Mongolian ankylosaurid that preserves impressions of the keratinous scales overlying the bony osteoderms. This specimen provides new information on the shape, texture, and ar− rangement of osteoderms. Large flat, keeled osteoderms are found over the pelvis, and osteoderms along the tail include large keeled osteoderms, elongate osteoderms lacking distinct apices, and medium−sized, oval osteoderms. The specimen differs in some respects from other Tarchia gigantea specimens, including the morphology of the neural spines of the tail club handle and several of the largest osteoderms. Key words: Dinosauria, Ankylosauria, Ankylosauridae, Tarchia, Saichania, Late Cretaceous, Mongolia. Victoria M. Arbour [[email protected]] and Philip J. Currie [[email protected]], Department of Biological Sciences, CW 405 Biological Sciences Building, University of Alberta, Edmonton, Alberta, Canada, T6G 2E9; Nicolai L. Lech−Hernes [nicolai.lech−[email protected]], Bayerngas Norge, Postboks 73, N−0216 Oslo, Norway; Tom E. Guldberg [[email protected]], Fossekleiva 9, N−3075 Berger, Norway; Jørn H. -
External Anatomy
SALMONIDS IN THE CLASSROOM: SALMON DISSECTION EXTERNAL ANATOMY Shape l Salmon are streamlined to move easily through water. Water has much more resistance to movement than air does, so it takes more energy to move through water. A streamlined shape saves the fish energy. Fins l Salmon have eight fins including the tail. They are made up of a fan of bone- like spines with a thin skin stretched between them. The fins are embedded in the salmon’s muscle, not linked to other bones, as limbs are in people. This gives them a great deal of flexibility and manoeuverability. l Each fin has a different function. The caudal or tail, is the largest and most powerful. It pushes from side to side and moves the fish forward in a wavy path. l The dorsal fin acts like a keel on a ship. It keeps the fish upright, and it also controls the direction the fish moves in. l The anal fin also helps keep the fish stable and upright. l The pectoral and pelvic fins are fused for steering and for balance. They can also move the fish up and down in the water. l The adipose fin has no known function. It is sometimes clipped off in hatchery fish to help identify the fish when they return or are caught. Slime l Many fish, including salmon, have a layer of slime covering their body. The slime layer helps fish to: ¡ slip away from predators, such as bears. ¡ slip over rocks to avoid injuries ¡ slide easily through water when swimming ¡ protect them from fungi, parasites, disease and pollutants in the water Scales Remove a scale by scraping backwards with a knife. -
A Revised Taxonomy of the Iguanodont Dinosaur Genera and Species
ARTICLE IN PRESS + MODEL Cretaceous Research xx (2007) 1e25 www.elsevier.com/locate/CretRes A revised taxonomy of the iguanodont dinosaur genera and species Gregory S. Paul 3109 North Calvert Station, Side Apartment, Baltimore, MD 21218-3807, USA Received 20 April 2006; accepted in revised form 27 April 2007 Abstract Criteria for designating dinosaur genera are inconsistent; some very similar species are highly split at the generic level, other anatomically disparate species are united at the same rank. Since the mid-1800s the classic genus Iguanodon has become a taxonomic grab-bag containing species spanning most of the Early Cretaceous of the northern hemisphere. Recently the genus was radically redesignated when the type was shifted from nondiagnostic English Valanginian teeth to a complete skull and skeleton of the heavily built, semi-quadrupedal I. bernissartensis from much younger Belgian sediments, even though the latter is very different in form from the gracile skeletal remains described by Mantell. Currently, iguanodont remains from Europe are usually assigned to either robust I. bernissartensis or gracile I. atherfieldensis, regardless of lo- cation or stage. A stratigraphic analysis is combined with a character census that shows the European iguanodonts are markedly more morpho- logically divergent than other dinosaur genera, and some appear phylogenetically more derived than others. Two new genera and a new species have been or are named for the gracile iguanodonts of the Wealden Supergroup; strongly bipedal Mantellisaurus atherfieldensis Paul (2006. Turning the old into the new: a separate genus for the gracile iguanodont from the Wealden of England. In: Carpenter, K. (Ed.), Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. -
A Short History of Dinosaurian Osteocytes
Palaeont. afr., 34, 59-61 ( 1997) A SHORT HISTORY OF DINOSAURIAN OSTEOCYTES. by _ R.E.H. (Robin) Reid School of Geosciences, The Queen's University ofB elfast, Belfast BT7 INN, Northern Ireland. ABSRACT A recent supposed discovery of dinosauri an osteocytes by Fukuda and Obata ( 1993) ignored earli er records from more than 20 d inosaurs, dating back 150 years. Some of the bodies they identified as osteocytes are also more like ly to represent chondrocytes. KEYWORDS: Bone histology, osteocytes. INTRODUCTION Brontosaurus (pp. 302, 303), Diplodocus (pp. 304, In a recent paper, Fukuda and Obata (1993) claimed 306), Camarasaurus (p. 311, as "M orosaurus" pp. 322, the discovery of dinosaurian osteocytes, in the form of 324), Haplocanthosaurus (pp.313, 314), Allosaurus mineral casts of their lacunae and canaliculi seen in (pp. 315, 3 16), Stegosaurus (p. 321) Iguanodon (pp. hadrosaurid bones, and stated that, despite recent 326-328), Dryptosaurus (p. 344), Triceratops (p. 346), histological studies, " ... there is no information on and Anatosaurus (pp. 348-350), as "Trachodon". osteocytes in dinosaur bone ti ssues" (p. 99, para. 1). At Osteocyte Stegosaurus lacunae recorded as from that date, however, earlier studies outlined here Zanclodon (p. 262), which is now considered contained records from more than 20 dinosaurs, dating indeterminate, are probably from a prosauropod (c.f. back 150 years (Quelett 1849), and even electron Benton 1986, pp. 295-297). A preserved canalicular microscope photographs had been in print since 1966 network was noted in Brontosaurus (p. 302), though (Pawlicki, Korbel & Kubiak 1966). This note reviews a not illustrated, and lacunae affected by fungal range of relevant references, for readers not familiar enlargement, mineral infilling, loss of canaliculae or with them, and also shows that some bodies identified complete obliteration were reported from various as osteocytes by Fukuda and Obata in the articular genera. -
The Systematic Position of the Enigmatic Thyreophoran Dinosaur Paranthodon Africanus, and the Use of Basal Exemplifiers in Phyl
1 The systematic position of the enigmatic thyreophoran dinosaur Paranthodon africanus, 2 and the use of basal exemplifiers in phylogenetic analysis 3 4 Thomas J. Raven1,2 ,3 and Susannah C. R. Maidment2 ,3 5 61Department of Earth Science & Engineering, Imperial College London, UK 72School of Environment & Technology, University of Brighton, UK 8 3Department of Earth Sciences, Natural History Museum, London, UK 9 10Corresponding author: Thomas J. Raven 11 12Email address: [email protected] 13 14 15 16 17 18 19 20 21ABSTRACT 22 23The first African dinosaur to be discovered, Paranthodon africanus was found in 1845 in the 24Lower Cretaceous of South Africa. Taxonomically assigned to numerous groups since discovery, 25in 1981 it was described as a stegosaur, a group of armoured ornithischian dinosaurs 26characterised by bizarre plates and spines extending from the neck to the tail. This assignment 27that has been subsequently accepted. The type material consists of a premaxilla, maxilla, a nasal, 28and a vertebra, and contains no synapomorphies of Stegosauria. Several features of the maxilla 29and dentition are reminiscent of Ankylosauria, the sister-taxon to Stegosauria, and the premaxilla 30appears superficially similar to that of some ornithopods. The vertebral material has never been 31described, and since the last description of the specimen, there have been numerous discoveries 32of thyreophoran material potentially pertinent to establishing the taxonomic assignment of the 33specimen. An investigation of the taxonomic and systematic position of Paranthodon is therefore 34warranted. This study provides a detailed re-description, including the first description of the 35vertebra. Numerous phylogenetic analyses demonstrate that the systematic position of 36Paranthodon is highly labile and subject to change depending on which exemplifier for the clade 37Stegosauria is used. -
AMERICAN MUSEUM NOVITATES Published by of NATURAL HISTORY Dec
AMERICAN MUSEUM NOVITATES Published by OF NATURAL HISTORY Dec. 4, 1933 Number 679 THE AMERICAN NewMUSEUMYork City 56.81, 9 (117:51.7) TWO NEW DINOSAURIAN REPTILES FROM MONGOLIA WITH NOTES ON SOME FRAGMENTARY SPECIMENS' BY CHARLES W. GILMORE2 CONTENTS PAGE INTRODUCTION......................................................... 1 DESCRIPTION OF GENERA AND SPECIES................................... 3 Order Orthopoda.................................................... 3 Family Nodosauridae............................................. 3 Pinacosaurus grangeri, new genus, new species. 3 Order Saurischia.................................................... 9 . Mongolosaurus haplodon, new genus, new species .. .. 11 NOTES ON OTHER DINOSAURIAN OCCURRENCES IN MONGOLIA................ 16 Family Deinodontidae.............................................. 16 Family Ornithomimidae............................................. 17 Family Hadrosauridae.............................................. 18 Family Ceratopsidae............................................... 19 BIBLIOGRAPHY............. 20 INTRODUCTION The present paper gives the results of a study of the residue of the dinosaur collections made in Mongolia by various Asiatic expeditions of The American Museum of Natural History. All of the material is frag- mentary, but diagnostic parts of two of the specimens are sufficiently preserved to be worthy of detailed description. One records the presence of a new genus and species of the Nodosauridae, the other a new genus and species of the Sauropoda. The -
Threat-Protection Mechanics of an Armored Fish
JOURNALOFTHEMECHANICALBEHAVIOROFBIOMEDICALMATERIALS ( ) ± available at www.sciencedirect.com journal homepage: www.elsevier.com/locate/jmbbm Research paper Threat-protection mechanics of an armored fish Juha Songa, Christine Ortiza,∗, Mary C. Boyceb,∗ a Department of Materials Science and Engineering, Massachusetts Institute of Technology, 77 Massachusetts Avenue, RM 134022, Cambridge, MA 02139, USA b Department of Mechanical Engineering, Massachusetts Institute of Technology, 77 Massachusetts Avenue, Cambridge, MA 02139, USA ARTICLEINFO ABSTRACT Article history: It has been hypothesized that predatory threats are a critical factor in the protective functional design of biological exoskeletons or “natural armor”, having arisen through evolutionary processes. Here, the mechanical interaction between the ganoid armor of the predatory fish Polypterus senegalus and one of its current most aggressive threats, a Keywords: toothed biting attack by a member of its own species (conspecific), is simulated and studied. Exoskeleton Finite element analysis models of the quadlayered mineralized scale and representative Polypterus senegalus teeth are constructed and virtual penetrating biting events simulated. Parametric studies Natural armor reveal the effects of tooth geometry, microstructure and mechanical properties on its ability Armored fish to effectively penetrate into the scale or to be defeated by the scale, in particular the Mechanical properties deformation of the tooth versus that of the scale during a biting attack. Simultaneously, the role of the microstructure of the scale in defeating threats as well as providing avenues of energy dissipation to withstand biting attacks is identified. Microstructural length scale and material property length scale matching between the threat and armor is observed. Based on these results, a summary of advantageous and disadvantageous design strategies for the offensive threat and defensive protection is formulated. -
EUOPLOCEPHALUS ANKYLOSAURUS TSAGANTEGIA GASTONIA GARGOYLEOSAURUS - -- PANOPLOSAURUS 7,8939, (6, - Edmontonla 1L,26,34)
THE UNIVERSITY OF CALGARY Skull Morphology of the Ankylosauria by Matthew K. Vickaryous A THESIS SUBMITED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE DEPARTMENT OF BIOLOGICAL SCIENCES CALGARY, ALBERTA JANUARY, 2001 O Matthew K. Vickaryous 2001 National Library Bibliothéque nationale I*l of Canada du Canada Acquisitions and Acquisitions et Bibliographie Services services bibliographiques 395 Wellington Street 395, rue Wellington Ottawa ON KIA ON4 Ottawa ON KIA ON4 Canada Canada Your Rie Votre rrlftimce Our fite Notre dfbrance The author has granted a non- L'auteur a accordé une licence non exclusive licence allowing the exclusive permettant à la National Library of Canada to Bibliothèque nationale du Canada de reproduce, loan, distribute or sell reproduire, prêter, distribuer ou copies of this thesis in microform, vendre des copies de cette thèse sous paper or electronic formats. la forme de microfiche/film, de reproduction sur papier ou sur format électronique. The author retains ownership of the L'auteur conserve la propriété du copyright in this thesis. Neither the droit d'auteur qui protège cette thèse. thesis nor substantial extracts fiom it Ni la thèse ni des extraits substantiels may be printed or otherwise de celle-ci ne doivent être imprimés reproduced without the author's ou autrement reproduits sans son permission. autorisation. Abstract The vertebrate head skeleton is a fundamental source of biological information for the study of both modern and extinct taxa. Detailed analysis of structural modifications in one taxon frequently identifies developmental and / or functional features widespread amongst a more inclusive clade of organisms. -
A Census of Dinosaur Fossils Recovered from the Hell Creek and Lance Formations (Maastrichtian)
The Journal of Paleontological Sciences: JPS.C.2019.01 1 TAKING COUNT: A Census of Dinosaur Fossils Recovered From the Hell Creek and Lance Formations (Maastrichtian). ______________________________________________________________________________________ Walter W. Stein- President, PaleoAdventures 1432 Mill St.. Belle Fourche, SD 57717. [email protected] 605-210-1275 ABSTRACT: A census of Hell Creek and Lance Formation dinosaur remains was conducted from April, 2017 through February of 2018. Online databases were reviewed and curators and collections managers interviewed in an effort to determine how much material had been collected over the past 130+ years of exploration. The results of this new census has led to numerous observations regarding the quantity, quality, and locations of the total collection, as well as ancillary data on the faunal diversity and density of Late Cretaceous dinosaur populations. By reviewing the available data, it was also possible to make general observations regarding the current state of certain exploration programs, the nature of collection bias present in those collections and the availability of today's online databases. A total of 653 distinct, associated and/or articulated remains (skulls and partial skeletons) were located. Ceratopsid skulls and partial skeletons (mostly identified as Triceratops) were the most numerous, tallying over 335+ specimens. Hadrosaurids (Edmontosaurus) were second with at least 149 associated and/or articulated remains. Tyrannosaurids (Tyrannosaurus and Nanotyrannus) were third with a total of 71 associated and/or articulated specimens currently known to exist. Basal ornithopods (Thescelosaurus) were also well represented by at least 42 known associated and/or articulated remains. The remaining associated and/or articulated specimens, included pachycephalosaurids (18), ankylosaurids (6) nodosaurids (6), ornithomimids (13), oviraptorosaurids (9), dromaeosaurids (1) and troodontids (1). -
Dinosaurs, by William Diller Matthew
The Project Gutenberg eBook of Dinosaurs, by William Diller Matthew http://www.gutenberg.org/files/19302/19302-h/19302-h.htm The[10][11][12][13][14][15][16][17][18][19][20][21][22][1][2][3][4][5][6][7][8][9] Project Gutenberg eBook, Dinosaurs, by T[100][103][104][105][106][108][109][110][111][112][113][115][117][118][119][120][121][123][124][125][126][127][128][129][130][131][132][133][134][135][136][137][138][139][140][141][142][143][144][145][146][147][148][149][150][151][152][153][154][155][156][157][158][159][161][162][10][11][12][13][14][15][17][18][19][20][21][22][23][24][26][27][28][29][30][31][34][35][36][38][39][40][41][42][43][44][45][46][47][48][49][50][51][52][53][54][55][56][57][58][61][63][64][65][66][67][68][69][70][71][72][73][74][76][77][78][79][80][81][83][84][85][86][87][88][90][91][93][94][95][96][97][98][99]OC William Diller Matthew This eBook is for the use of anyone anywhere at no cost and with almost no restrictions whatsoever. You may copy it, give it away or re-use it under the terms of the Project Gutenberg License included with this eBook or online at www.gutenberg.org Title: Dinosaurs With Special Reference to the American Museum Collections Author: William Diller Matthew Release Date: September 16, 2006 [eBook #19302] Language: English Character set encoding: ISO-8859-1 ***START OF THE PROJECT GUTENBERG EBOOK DINOSAURS*** E-text prepared by Brian Janes, Suzanne Lybarger, Jeannie Howse, and the Project Gutenberg Online Distributed Proofreading Team http://www.pgdp.net) Transcriber's Note: Click the image to see a larger version.