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Naturalist

Volume 43 Number 1 Article 6

1-31-1983

A vascular flora of the ,

James G. Harris Brigham Young University

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Recommended Citation Harris, James G. (1983) "A vascular flora of the San Rafael Swell, Utah," Great Basin Naturalist: Vol. 43 : No. 1 , Article 6. Available at: https://scholarsarchive.byu.edu/gbn/vol43/iss1/6

This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. A VASCULAR FLORA OF THE SAN RAFAEL SWELL, UTAH'

James G. Harris^

Abstract.— The vegetation of the San Rafael Swell in southeastern Utah is examined based on personal field col- lections and previously collected herbarium specimens in the Brigham Young University Herbarium (BRY). An anno- tated checklist includes information on frequency of occurrence and habitat preference for each entity. Treated are 491 vascular taxa from 59 families.

The San Rafael Swell is the eroded rem- (1981), Welsh (1978, 1980a, 1980b), Welsh massive in nant of a domal anticline, oval and Atwood (1981), Welsh and Moore (1973), shape, stretching along northeasterly axis a Welsh and Reveal (1977), Welsh et al. (1981); in from Capitol Reef National Park northern , Cronquist et al. (1977). Wayne County to the foot of the Tavaputs The checklist includes 478 Plateau in Carbon County. Folding of the taxa found in the study area and an addition- earth's crust, which formed the domal anti- al 13 taxa known to occur on lands immedi- cline, and the subsequent erosion of the cen- ately adjacent to the study area and to be ex- tral dome left a huge area of concentric stone pected in the San Rafael Swell. These 13 taxa rings, each progressively younger from the are listed below: innermost to the outermost. One of these Allionia incarnata L. rings, the Ferron Sandstone Member of the wrightii Gray Mancos Shale Formation, encloses approx- Descurainia pinnata (Walter) Britt. var. filipes (Gray) Peck imately L4 million acres of land and forms Eriogonum smithii Reveal the boundary of this study. In some cases, micromera Boiss. where the Ferron Sandstone has been com- Haplopappus drummondii (T. & G.) Blake pletely eroded from the structural confines of Helianthus petiolaris Nutt. Hymenopappus Hook. var. lugens (Greene) the swell, artificial boundaries have been fol- filifolius Jeps. lowed (Fig. 1). pusillus Pursh var. rubens (Rydb.) Welsh Throughout the field seasons of 1979 and Phacelia demissa Gray var. heterotricha Howell 1980, I attempted to collect representatives alternans (Jones) Howell of all vascular plant species growing in the Sphaeralcea munroana (Dougl.) Spach Sporobolus giganteus Nash San Rafael Swell. A voucher specimen from flora of the Rafael Swell is not each collection has been placed in the Brig- The San particularly rich in of species. This ham Young University Herbarium (BRY). In- numbers is part, the relatively nar- cluded in the checklist are species not found due, at least in to by me, but collected by others and vouched row elevational range in the area. Although rich in of species, the flora is for by specimens in the Brigham Young Uni- not numbers rather rich in numbers of unique and geo- versity Herbarium. The checklist is not defin- graphically restricted species. The following itive; additional taxa will undoubtedly be dis- to the San covered in the swell. eight species are strictly endemic Swell: The following sources were used for identi- Rafael fication and nomenclature: lower vascular rafaelensis Jones Cryptantha johnstonii Higgins and gymnosperms, Cronquist et al. Cryptantha pnesiana (Payson) Payson (1972); dicotyledons, Neese and Welsh maguirei Cronquist

'Part of a master's thesis, originally submitted to the Department of Botany and Range Science, Brigham Young University, Provo, Utah 84602. 'Department of Botany and Range Science, Brigham Young University, Prove, Utah 84602. Present address: Department of Botany, University of Alberta, Edmonton, Alberta T6G 2E9.

79 80 Great Basin Naturalist Vol. 43, No. 1

Emery County

Fig. 1. Map of the San Rafael Swell study area.

Pediocactus despainii Welsh & Goodrich Vegetative Zones wrightii Benson Sphaeralcea psoraloides Welsh The vegetation of the San Rafael Swell can Thelypodiopsis harnebyi Welsh & Atwood be divided into five major zones: the Sand An additional seven species are endemic to Desert Shrub Zone, the Salt Desert Shrub the San Rafael Swell and the adjacent lands Zone, the Mixed Desert Shrub Zone, the Pin- in close proximity to the swell. They are list- yon- Zone, and the Ponderosa Pine- ed as follows: Mountain Brush Zone. These zones are con- Astragalus pardalinus (Rydb.) Barneby Astragalus woodruffii Jones trolled principally by elevation, precipi- Hoffmanseggia repens (Eastw.) Cockerell tation, and local edaphic factors. Although in Hymenoxys depressa (T. & G.) Welsh & Reveal some areas the zones are clear and well de- Lomatium junceum Barneby & Holmgren fined, in most areas they interfuse consid- Psorothamnus polyadenius (Torr.) Rydb. var. jonesii Barneby erably and it becomes difficult to draw a apnea Welsh & Reveal sharp boundary line between them. January 1983 Harris: San Rafael Swell Flora 81

Sand Desert Shrub Zone.— The San Ra- 7800 ft (1766-2377 m). Within this altitudin-

fael Desert, in the southeastern region of the al range Juniperus osteosperma is dominant at

San Rafael Swell, ranges from an elevation of the lower elevations and Pinus edulis is domi- about 4200 ft (1280 m) at the lowest point to nant at the higher elevations. Intermediate about 5000 ft (1524 m) at the highest point. elevations yield a heterogeneous mixture of

It is a sandy desert with moving dunes in both species. some areas. Common shrubs in this zone in- Dense stands have almost no understory, clude: Artemisia filifolia. torreyana. but more open or scattered stands will in- hatrimaniae, incana, Ti- clude almost all the species found in the quilia latior, and Coleogyne ramosissima. Mixed Desert Shrub Zone. PoNDEROSA Pine-Mountain Abronia elliptica, Oenothera pallida, Hoff- Brush Zone.— highest point in manseggia repens, and Oryzopsis hymenoides The the San Rafael Swell is the San Rafael Knob at 7921 ft are characteristic herbaceous plants of this (2414 m) in elevation. The area surrounding the zone. knob and the summit of Cedar Mountain rep- Salt Desert Shrub Zone.— In the San resent the highest vegetative zone in the Rafael Swell the Salt Desert Shrub Zone is swell. Pinus edulis continues to remain domi- widespread, occurring mainly on alkaline nant even at the highest elevations, but above clay between 4500 and 5800 ft 7500 ft (2286 m) plants characteristic of the (1372-1766 m) in elevation. Communities in Ponderosa Pine-Mountain Brush Zone be- this zone are characteristically dominated by come increasingly common. Some common Atriplex interspersed with any of confertifolia species in this zone are: Pinus ponderosa, Ar- the following subdominant species: Hilaria temisia nova, Cowania mexicana, Mahonia jamesii, Oryzopsis hymenoides, Enceliopsis fremontii, Populus tremuloides, Amelanchier nudicaulis, Aristida purpurea, Eriogonum in- utahensis, Cercocarpus montanus, Ribes lep- flatum, Salsola iherica or Ceratoides lanata. tanthum, Juniperus scopulorum, Hymenoxys Highly alkaline flood plains in this zone richardsonii, Symphoricarpos longiflorus and are often covered by nearly pure stands of Philadelphus microphyllus. Sarcobatus vermiculatus, which may at times In addition to these five major vegetative be accompanied by thick clumps of Tamarix zones, smaller communities dependent on ramosissima. unique local habitats are scattered through- Heavy clay soils of the Mancos Shale For- out the area. These include plants distributed mation are often dominated by Atriplex cu- along seeps and streams, along washes and neata and Atriplex corrugata. During wet roadsides where runoff collects, in hanging springs these areas may be blanketed by an- gardens, on rocky outcrops, and on disturbed nual species such as Cleome lutea, Lupinus sites. pusillus, Malcolmia africana, Camissonia sca- Annotated Checklist poidea, and Camissonia eastwoodiae. Mixed Desert Shrub Zone.— Deep, well- Following each species entry in the check- list (with the exception of the 13 taxa from drained soils between 4500 and 5800 ft area) a designa- (1372-1766 m) in elevation are often domi- lands adjacent to the study nated by one or several of the following tion is given concerning the frequency of oc- currence and habitat preference of the taxon. shrubby species: Artemisia tridentata, Arte- Plants of broad distribution that are not lim- misia bigelovii, Artemisia spinescens, Xantho- ited to a specific zone or community type are cephalum sarothrae or Chrysothamnus vis- listed as widespread. The following abbrevia- cidiflorus. These may be intermingled with tions are used: an understory of herbaceous plants including: C—common Hilaria jamesii, Stipa comata, Boutehua gra- O— occasional cilis. Astragalus spp., Sphaeralcea spp., and U—uncommon Leucelene ericoides. R— rare SnD— Sand Desert Shrub Zone Pinyon-Juniper Zone.— Pinus edulis and SID— Salt Desert Shrub Zone Juniperus osteosperma are conspicuously MD— Mixed Desert Shrub Zone dominant at elevations between 5800 and PJ—Pinyon-Juniper Zone 82 Great Basin Naturalist Vol. 43, No. 1

Ponderosa Pine-Mountain Brush Zone PM— Chrysothamnus nauseosus (Pall.) Britt. var. junceus SS— Seeps and streams (Greene) Hall (U;WR) and roadsides WR—Washes Chrysothamnus nauseosus (Pall.) Britt. var. leio- HG—Hanging gardens spermus (Gray) Hall (R;WR) RO—Rocky outcrops Chrysothamnus pulchellus (Gray) Greene (U;WR) DS—Disturbed sites Chrysothamnus viscidiflorus (Hook.) Nutt. var. steno- W—Widespread phyllus (Gray) Hall (U;WR) ACERACEAE Chrysothamnus viscidiflorus (Hook.) Nutt. var. vis- Torr. (R;PM) Acer glabrum cidiflorus (C;WR) Anacardiaceae Cirsium pulchellum (Greene) Woot. & Standi. (U;WR) var. (Greene) Barkl. Rhus trilobata Nutt. simplicifolia Cirsium undulatum (Nutt.) Spreng. (C;WR) (C;WR) Dicoria canescens Gray (U;SnD) var. trilobata (C;WR) Rhus trilobata Nutt. Encelia frutescens Gray (U;WR) rydbergii (Small) Greene (C;SS,HG) Toxicodendron Enceliopsis nudicaulis (Gray) Nels. (C;S1D, MD) Apiaceae Enceliopsis nutans (Eastw.) Nels. (C;S1D,MD) (0;SnD,SlD) Cymopteris acaulis (Pursh) Raf. Erigeron argentatus Greene (0;S1D,MD,PJ) Nels. (C;W) Cyrnopteris bulbosus Erigeron compactus Blake var. consimilis Cronquist Cymopteris fendleri Gray (C;W) (0;MD,PJ) purpurascens (Gray) (0;S1D) Cymopteris Jones Erigeron divergens T. & G. (U;PJ) Cymopteris purpureus Wats. (0;W) Erigeron eatonii Gray (U;PJ,PM) Lomatium junceum Barneby & Holmgren Cronquist (R;WR) (R;S1D,MD,PJ) Erigeron pulcherrimus Heller var. pulcherrimus Stum suave Walt. (R;WR) (C;S1D,MD,PJ) Apocynaceae Erigeron pumilus Nutt. ssp. concinnoides Cronq. var. Amsonia eastwoodiana Rydb. (C;SnD) condensatus (Eaton) Cronq. (C;S1D,MD,PJ) Ainsonia jonesii Woodson (0;MD,PJ) Erigeron utahensis Gray (C;WR,RO) Apocynum cannabintim L. (C;SS,HG) Gaillardia pinnatifida Torr. (C;S1D,MD) Cycladenia humilis Benth. var. jonesii (Eastw.) Welsh Gaillardia spathulata Gray (C;S1D,MD,PJ) & Atwood (R;MD) Grindelia squarrosa (Pursh) Dunal. (0;WR) ASCLEPIADACEAE Haplopappus armerioides (Nutt.) Gray (C;MD,PJ,RO) Asclepias cryptoceras Wats. (C;W) Haplopappus drummondii (T. & G.) Blake Asclepias labriformis Jones (C;WR,SD,MD) Helianthella microcephala (Gray) Gray (C;PJ,PM) Asclepias macrosperma Eastw. (C;SnD,MD) Helianthus annutts L. (C;WR) Asclepias ruthiae Maguire & Woodson (0;W) Helianthus anomalus Blake (C;SnD) Asclepias speciosa Torr. (C;WR) Helianthus petiolaris Nutt. Heterotheca villosa (Pursh) Shinners (C;S1D,MD,PJ) Hook. var. lugens (Greene) Ambrosia acanthicarpa Hook. (0;S1D,MD,PJ) Hymenopappus filifolius Jeps. Ambrosia artemisiifolia L. (C;S1D,MD,PJ) Hymenopappus filifolius Hook. var. megacephalus Artemisia biennis Willd. (U;WR) Turner (C;S1D,MD) Artemisia bigelovii Gray (C;MD,PJ) Hymenopappus filifolius Hook. var. pauciflorus Artemisia campestris L. (U;SnD,MD) (Johnst.) Turner (0;S1D) Artemisia dracunculus L. (C;WR) Hymenoxys acaulis (Pursh) Parker var. arizonica Artemisia filifolia Torr. (C;SnD) (Greene) Parker (C;MD,PJ) Artemisia frigida Willd. (R; WR) Hymenoxys depressa (T. & G.) Welsh & Reveal Artemisia ludoviciana Nutt. (0;WR) (0;R0) Nels. (C;PJ,PM) Hymenoxys richardsonii (Hook.) Cockerell (C;PJ,PM) Artemisia pygmaea Gray (0;S1D,MD,PJ) Iva axillaris Pursh (0;SS,WR) Artemisia spinescens Eaton (0;S1D, MD) Iva xanthifolia Nutt. (0;WR) Artemisia tridentata Nutt. (C;MD,PJ,WR) Leucelene ericoides (Torr.) Greene (C;W) Aster glaucodes Blake (U;PM) Lygodesmia arizonica Tomb (C;S1D,MD,WR) Baccharis emoryi Gray (U;WR) Lygodesmia grandiflora (Nutt.) T. & G. (C;S1D) Brickellia longifolia Wats. (U;WR) Lygodesmia juncea (Pursh) D. Don (R;SnD) Brickellia microphylla (Nutt.) Gray (U;WR) Machaeranthera canescens (Pursh) Gray (R;SnD) Brickellia oblongifolia var. linifolia (Eaton) Robins Machaeranthera grindelioides (Nutt.) Shinners (C;W) (0;PJ,WR) Machaeranthera linearis Greene (U;WR) Brickellia scabra (Gray) Nels. (R;HG,WR) Machaeranthera tanacetifolia (H.B.K.) Nees (C;WR) Chaenactis stevioides Hook. & Arn. (C;S1D,MD) Malacothrix sonchoides (Nutt.) T. & G. (C;S1D,MD) Chamaechaenactis scaposa (Eastw.) Rydb. (0;PJ) Oxytena acerosa Nutt. (C;SS,WR) Chrysothamnus greenei (Gray) Greene (0;S1D) Parthenium ligulatum (Jones) Barneby (U;S1D,MD,PJ) Chrysothamnus linifolius Greene (0;WR) Petradoria pumila (Nutt.) Greene (C;PJ,WR,RO) Chrysothamnus nauseosus (Pall.) Britt. var. consimilis Platyschkuhria integrifolia (Gray) Rydb. var. oblongi- (Greene) Hall (C;WR) folia (Gray) Ellison (C;WR) Chrysothamnus nauseosus (Pall.) Britt. var. gnapha- Prenanthella exigua (Gray) Rydb. (C;WR,RO) loides (Greene) Hall (U;WR) Senecio multicapitatus Greenm. (U;PJ,PM,WR) January 1983 Harris: San Rafael Swell Flora 83

Senecio multilobatus T. & G. (0;PJ,WR) Chorispora tenella (Pall.) DC. (U;DS) Stephanomeria exigua Nutt. (0;WR) Descurainia pinnata (Walt.) Britt. var. filipes (Gray) Stephanomeria runcinata Nutt. (R;WR) Peck Stephanomeria spinosa (Nutt.) Tomb (U;WR) Descurainia pinnata (Walt.) Britt. var. intermedia Stephanomeria tenuifolia (Torr.) Hall (C;WR,RO) (Rydb.) Hitchc. (U;MD) Taraxacum officinale Weber (C;SS) Descurainia pinnata (Walt.) Britt. var. osmarium Tetradymia canescens DC. (0;PJ,PM) (Cockerell) Shinners (C;W) Tetradymia glabrata Gray (C;MD,PJ) Descurainia sophia (L.) Webb. (0;WR) Tetradymia spinosa Hook. & Am. (0;WR) Dithyrea wislizenii Engelm. in Wisliz. (0;SnD) Thelesperma subnudum Gray (C;W) Lepidium densiflorum Schrad. var. pubicarpum (Nels.) Townsendia annua Beaman (U;MD) Thell. (R;PJ) Welsh & Reveal (U;S1D,MD,PJ) Lepidium densiflorum Schrad. var. ramosum (Nels.) Townsendia incana Nutt. (C;S1D,MD,PJ) Thell. (0;MD,PJ) Tragopogon dubius Scop. (C;W) Lepidium montanum Nutt. var. jonesii (Rydb.) Hitchc. Vanclevea stylosa (Eastw.) Greene (C;SnD) (C;W) Wyethia scabra Hook. (C;WR) Lesquerella alpina (Nutt.) Wats. var. alpina Xanthium strumarium L. (0;SS) (0;PJ,PM) Xanthocephalum microcephalum (DC.) Shinners (R;PJ) Lesquerella intermedia (Wats.) Heller (0;PJ,PM) Xanthocephalum sarothrae (Pursh) Shinners Lesquerella kingii Wats. (R;PJ) (C;S1D,MD,PJ) Lesquerella rectipes Woot. & Standi. (C;W) Xylorhiza venusta (Jones) Heller (C;RO) Lesquerella subumbellata Rollins (U;PJ,PM) Berberidaceae Malcolmia africana (L.) R. Br. in Ait. (C;S1D,DS) Mahonia fremontii (Torr.) Fedde (C;MD,PJ) acutifolia Rydb. (C;PJ) Betulaceae Physaria chambersii Rollins (U;PJ) Betula occidentalis Hook. (R;SS) Schoencrambe linifolia (Nutt.) Greene (0;PJ,PM) Boraginaceae Sisymbrium altissimum L. (C;PJ,PM) Cryptantha cinerea Greene (R;PJ,PM) Stanleya pinnata (Pursh) Britt. (C;S1D) Cryptantha confertifolia (Greene) Payson (R;SnD) Stanleya viridiflora Nutt. in T. & G. (U;MD,PJ) Cryptantha crassisepala (T. & G.) Greene var. ela- Streptanthella longirostris (Wats.) Rydb. (C;W) chantha Johnst. (C;W) Streptanthus cordatus Nutt. ex T. & G. (C;PJ) Cryptantha flava (Nels.) Payson (C;W) Thelypodiopsis barnebyi Welsh & Atwood (R;MD) Cryptantha flavoculata (Nels.) Payson (C;PJ) Thelypodiopsis divaricata (Rollins) Welsh & Reveal Cryptantha fulvocanescens (Wats.) Payson var. ech- (C;S1D,MD,PJ) inoides (Jones) Higgins (0;S1D) Cactaceae vivipara Britt. Cryptantha fulvocanescens (Wats.) Payson var. ful- Coryphantha (Nutt.) & Rose (0;PJ) vocanescens (C;MD,PJ) Echinocereus triglochidiatus Engelm. var. melana- Cryptantha gracilis Osterh. (U;PJ) canthus (Engelm.) Benson (C;S1D,MD,PJ) Cryptantha humilis (Gray) Payson var. nana (Eastw.) Opuntia basilaris Engelm. & Bigel. (0;SnD,MD,PJ) Opuntia erinacea Engelm. (0;SnD) Higgins (0;PJ) Cryptantha jamesii (Torr.) Payson var. disticha Opuntia polyacantha Haw. (C;W) (Eastw.) Payson (U;MD,WR) despainii Welsh & Goodrich (R;PJ) Cryptantha jamesii (Torr.) Payson var. setosa (Jones) Pediocactus simpsonii (Engelm.) Britt. & Rose (R;PJ) Johnst. (C;PJ) Sclerocactus parviflorus Clover & Jotter var. inter^ Cryptantha johnstonii Higgins (U;S1D) medius (Peebles) Woodruff & Benson (C;W) Cryptantha jonesiana (Payson) Payson (U;S1D) Benson (U;S1D,MD,PJ) Cryptantha mensana (Jones) Payson (0;S1D) Capparidaceae Cleome lutea Hook. (C;S1D) Cryptantha paradoxa (Nels.) Payson (0;PJ) Cryptantha recurvata Coville (0;MD,PJ,WR) Cleome serrulata Pursh (0;PJ) Cryptantha rollinsii Johnst. (0;MD,PJ,WR) Cleomella palmerana Jones (0;PJ) Cryptantha tenuis (Eastw.) Payson (0;WR) Caprifoliaceae Cryptantha wetherillii (Eastw.) Payson (0;S1D) Sambucus coerulea Raf. (U;PJ,PM) Heliotropium convulvulaceum (Nutt.) Gray (U;SnD) Symphoricarpos longiflorus Gray (0;PJ,PM) Lappula occidentalis (Wats.) Greene var. cupulata Caryophyllaceae (Gray) Higgins (R;MD,PJ) Arenaria eastwoodiae Rydb. (C;PJ) Lappula occidentalis (Wats.) Greene var. occidentalis Arenaria fendleri Gray (C;S1D) (C;W) Arenaria hookeri Nutt. var. desertorum Maguire (R;PJ) Nutt. Lithospermum incisum Lehm. (C;PJ) Paronychia sessiliflora (0;PJ,WR) Tiquilia latior (Johnst.) Richardson (0;S1D,MD) Celastraceae Forsellesia meionandra (Koehne) Heller (0;PJ,RO) Arabis demissa Greene var. lanugida Rollins Chenopodiaceae (R;PJ,PM) Allenrolfea occidentalis (Wats.) Kuntze (0;S1D) Atriplex argentea Nutt. (U;S1D) Arabis perennans Wats. (0;PJ) Atriplex canescens (Pursh) Nutt. (C;S1D,MD) Arabis pulchra Jones var. pallens Jones (0;PJ) Frem.) Wats. (C;S1D) Arabis selbyi Rydb. (C;PJ) (Torr. & corrugata Wats. (C;S1D) Caulanthus crassicaulis (Torr.) Wats. (C;PJ) Atriplex 84 Great Basin Naturalist Vol. 43, No. 1

Atriplex cuneata Nels. (C;S1D) Astragalus desperatus Jones var. petrophilus Jones Atriplex graciliflora Jones (U;S1D) (C;MD,SnD,PJ) Atriplex powellii Wats. {C;S1D) Astragalus episcopus Wats. (C;MD,PJ) Atriplex saccaria Wats. (U;MD) Astragalus flavus Nutt. var. argillosus (Jones) Bameby Bassia hysopifolia Pallas (U;WR) (C;S1D) Ceratoides lanata (Pursh) Howell (C;S1D) Astragalus flavus Nutt. var. flavus (C;S1D,MD,PJ) Chenopodium fremontii Wats. (C;W) Astragalus flexuosus (Hook.) Don. var. diehlii (Jones). Chenopodium glauciim L. (0;S1D) Barneby (0;S1D,MD,PJ) Chenopodium pratericola Rydb. (C;SS) Astragalus gey eri Gray (0;MD) Grayia brandegei Gray (U;MD) Astragalus kentrophyta Gray var. cohradoensis Jones Halogeton glomeratus (Bieb.) Meyer (C;DS) (U;WR,RO) Kochia iranica Bomm. (C;WR) Astragalus kentrophyta Gray var. elatus Wats. (R;S1D) Kochia americana Wats. (R;S1D,WR) Astragalus lentiginosus Dougl. var. araneosus (Sheld.) Monolepis nuttalliana (Schult.) Greene (0;WR) Barneby (G;MD,PJ) Sahola iberica Sennen & Pan (C;W) Astragalus lentiginosus Dougl. var. palans (Jones) Sarcobatus vermiculatus (Hook.) Torr. (C;S1D,SS,WR) Jones (C;MD,PJ) Suaeda torreyana Wats. (C;SS,WR) Astragalus lonchocarpus Torr. (C;S1D,MD,PJ) CONVOLVULACEAE Astragalus moencoppensis Jones (G;S1D,MD,PJ) Convoltmlus arvensis L. (C;WR) Astragalus mollissimus Torr. var. thompsonae (Wats.) Cuscuta cuspidata Engelm. (C;MD) Barneby (C;W) CuPRESSACEAE Astragalus musiniensis Jones (C;S1D,MD,PJ) Juniperus osteosperma (Torr.) Little (C;PJ,PM) Astragalus pardalinus (Rydb.) Barneby Juniperus scopulorum Sarg. (U;PM) (0;SnD,MD,PJ) Cyperaceae Astragalus praelongus Sheld. var. ellisiae (Rydb.) Carex filifolia Nutt. (C;SS) Barneby (C;S1D) Carex parryana Dewey (R;SS) Astragalus praelongus Sheld. var. praelongus (U;S1D) Eleocharis palustris (L.) Roemer & Schult. (0;SS) Astragalus preussii Gray var. preussii (C;S1D,MD,PJ) Eleocharis rostellata (Torr.) Torr. (U;SS) Astragalus rafaelensis Jones (0;S1D) Scirpus maritimus L. (0;SS) Astragalus sabulonum Gray (0;PJ,PM) Scirpus pungens Vahl. (C;SS) Astragalus spatulatus Sheld. (0;PJ,PM) Scirpus validus Vahl. (0;SS) Astragalus subcinereus Gray var. basalticus Welsh Elaeagnaceae (R;PJ) Shepherdia rotundifolia Parry (U;PJ) Astragalus tenellus Pursh (0;MD,PJ) Shepherdia argentea (Pursh) Nutt. (U;WR) Astragalus wingatanus Wats. (R;S1D,MD,PJ) Ephedraceae Astragalus woodruffii Jones (U;SnD,MD) Ephedra torreyana Wats. (C;W) Dalea flavescens (Wats.) Welsh (0;SnD,MD,PJ) Ephedra viridis Coville (C;W) Dalea oligophylla (Torr.) Shinners (0;WR,RO) Equisetaceae Glycyrrhiza lepidota Pursh (C;WR,SS) Equisetum laevigatum A. Br. (C;SS) Hedysarum boreale Nutt. (C;MD,PJ,PM) Hoffmanseggia repens (Eastw.) Cockerell (C;SnD) Euphorbia albomarginata T. & G. (U;PJ) Lathyrus brachycalyx Rydb. var. eucosmus (Butters & Euphorbia fendleri T. & G. (C;S1D,MD,PJ) St. John) Welsh (U;WR) Euphorbia micromera Boiss. Lupinus argenteus Pursh var. argenteus (R;WR) Euphorbia nephradenia Bameby (R;S1D,WR) Lupinus argenteus Pursh var. tenellus Dougl. (U;MD) Euphorbia palmeri Engelm. (R;S1D) Lupinus brevicaulis Wats. (U;S1D,MD) Euphorbia parryi Engelm. (G;SnD) Lupinus pusillus Pursh var. pusillus (C;S1D,MD,PJ) Euphorbia robusta (Engelm.) Small (0;PJ,PM) Lupinus pusillus Pursh var. rubens (Rydb.) Welsh Euphorbia serphyllifolia Pers. (C;PJ,PM) Medicago sativa L. (C;WR) Astragalus amphioxys Gray var. amphioxys Melilotus alba Medicus (0;SS,WR) (C;S1D,MD,PJ) Melilotus officinalis (L.) Lam. (0;SS,WR) Astragalus amphioxys Gray var. vespertinus (Sheld.) Oxytropis lambertii Pursh var. bigelovii Gray Jones (C;W) (R;PJ,PM) Astragalus argophyllus Nutt. var. martinii Jones Oxytropis sericea Nutt. (0;MD,PJ) (U;PJ,PM) Psoralea lanceolata Pursh var. lanceolata (U;MD,PJ) Astragalus asclepiadoides Jones (C;S1D) Psoralea lanceolata Pursh var. stenophylla (Rydb.) Astragalus brandegei Porter (U;S1D,MD,PJ) Toft & Welsh (0;WR) Astragalus calycosus Torr. (0;S1D,MD,PJ) Psorothamnus polyadenius (Torr.) Rydb. var. jonesii Astragalus ceramicus Sheldon (U;WR) Barneby (R;S1D,MD) Astragalus coltonii Jones var. coltonii (C;S1D,MD,PJ) Psorothamnus thompsonae (Vail) Welsh & Atwood Astragalus convallarius Greene var. convallarius (R;MD) (U;PJ,PM) Robinia pseudoacacia L. (R;WR) Astragalus cymboides Jones (C;S1D,MD,PJ) Sophora stenophylla Gray (0;MD) Astragalus desperatus Jones var. desperatus Fumariaceae (C;SnD,MD,PJ) Corydalis aurea Willd. (R;PJ) January 1983 Harris: San Rafael Swell Flora 85

GEI>rrtANACEAE Oenothera pallida Lindl. ssp. trichocalyx (Nutt.) Munz Swertia albomarginata (Wats.) Kuntze (R;PM) & Klein (C;S1D,MD) Swertih utahensis (Jones) St. John (C;SnD,MD) Orchidaceae Hydrophyllaceae Epipactis gigantea Dougl. ex Hook. (C;HG,SS) Phacelia constancei Atwood (R;S1D) Phacelia corrugata Nels. (C;W) Orobanche fasciculata Nutt. (0;W) Phacelia demissa Gray var. demissa (0;S1D,PJ) Papaveraceae Phacelia demissa Gray var. heterotricha Howell Argemone corymbosa Greene (C;SnD) Phacelia indecora Howell (R;S1D) Argemone munita Dur. & Hilg. (C;SnD) Phacelia ivesiana Torr. (C;S1D,MD,PJ) PiNACEAE Phacelia rafaelensis Atwood (C;MD,PJ) Pinus edulis Engelm. (C;PJ,PM) JUNCACEAE Pinus ponderosa Lawson (C;PM) Willd. (C;SS) Juncus arcticus Pseudotsuga menziesii (Mirb.) Franc. (C;PM) Juncus balticus Willd. (C;SS) Plantaginaceae Juncus nodosus L. (U;SS) Plantago patagonica Jacq. (C;S1D,MD) Juncus torreyi Coville (0;SS) POACEAE JUNCAGINACEAE Agropyron caninum (L.) Beauv. (U;MD,RO) Triglochin maritima L. (U;SS) Agropyron cristatum (L.) Gaertn. (C;MD,PJ) Agropyron spicatum (Pursh) Scribn. & Smith (U;PJ) (Torr.) Gray (C;SnD) Agropyron trachycaulum (Link) Malte (U;WR) Liliaceae Andropogon hallii Hackel (R;WR) Allium geyeri Wats. (C;PJ,PM) Aristida purpurea Nutt. (C;W) Allium macropetalum Rydb. (U;S1D) Bouteloua curtipendula (Michx.) Gray (R;MD) Allium textile Nels. & Macbr. (C;SnD) (H.B.K.) Lag. ex Steud. Androstephium breviflorum Wats. (C;S1D,MD,PJ) (C;S1D,MD,PJ) Asparagus officinalis L. (0;WR,SS) Bromus japonicus Thunb. (R;PJ) Cahchortus nuttallii T. & G. (C;S1D,MD) Bromus tectorum L. (C;WR,DS) albomarginatum (Jones) Jones (C;SnD) Calamagrostis scopuhrum Jones (U;WR) Smilacina stellata (L.) Desf. (C;HG) Dichanthelium lanuginosum (Elliott) Gould (U;SS) Yucca harrimaniae Trel. (C;SnD,MD,PJ) Distichlis spicata (L.) Greene var. striata (Torr.) LiNACEAE Scribn. (C;S1D,WR) Linum aristatum Engelm. (U;WR) Elymus canadensis L. (U;WR) Linum perenne L. (C;WR) Elymus cinereus Scribn. & Merr. (0;MD,PJ) Linum puberulum (Engelm.) Heller (0;S1D) Elymus junceus Fischer (R;S1D,MD) Linum subteres Winkler (0;WR,PJ) Elymus salina Jones LOASACEAE (C;PJ) Erioneuron pilosum (Buckley) Nash (C;MD,RO,WR) Mentzelia albicaulis Dougl. (C;S1D,MD,PJ) Erioneuron pulchellum (H.B.K.) Tateoka (R;PJ) Mentzelia humilis (Gray) DarL (U;WR) Festuca pratensis Huds. (U;SS) Mentzelia multiflora (Nutt.) Gray (C;SnD) Hilaria jamesii (Torr.) Benth. (C;W) Mentzelia pterosperma Eastw. (U;WR) Hordeum jubatum L. (C;WR) Mentzelia pumila (Nutt.) T. & G. (0;WR) Muhlenbergia asperifolia (Nees & Mey.) Parodi Malvaceae (0;WR) Sphaeralcea coccinea (Nutt.) Rydb. (C;W) Muhlenbergia pungens Thurber (C;SnD) Sphaeralcea grossulariifolia (Hook. & Arn.) Rydb. Munroa squarrosa (Nutt.) Torr. (0;SnD,MD) (0;PJ) Oryzopsis hymenoides (Roem. & Schult.) Ricker Sphaeralcea leptophylla (Gray) Rydb. (R;WR) (C;W) Sphaeralcea munroana (Dougl.) Spach Oryzopsis micrantha (Trin. & Rupr.) Thurber (U;PJ) Sphaeralcea parvifolia Nels. (C;W) Panicum virgatum L. (0;SS,WR) Sphaeralcea psoraloides Welsh (R;S1D,MD) Phragmites australis (Car.) Trin. ex Steud. (C;SS) Nyctaginaceae Poa fendleriana (Steud.) Vasey (0;PJ,PM) Abronia elliptica Nels. (C;SnD,MD,WR) Poa pratensis L. (U;SS,WR) Abronia nana Wats. (U;S1D,PJ) Poa sandbergii Vasey (U;PJ) Allionia incarnata L. Polypogon monspeliensis (L.) Desf. (U;SS,WR) Oxybaphus linearis (Pursh) Robins. (C;W) Schedonnardus paniculatus (Nutt.) Trel. (R;S1D) Tripterocalyx micranthus (Torr.) Hook. (0;SnD) Schizachyrium scoparium (Michx.) Nash (0;PJ,WR) Oleaceae Sitanion hystrix (Nutt.) Smith (C;W) Fraxinus anomala Torr. in Wats. (C;WR) Spartina gracilis Trin. (U;SS) Onagraceae Sporobolus airoides (Torr.) Torr. (C;S1D,MD) C^lylOphus lavandulaefolia (T. & G.) Raven Sporobolus contractus Hitchc. (0;MD,PJ,WR) (0;PT,RO) Sporobolus cryptandrus (Torr.) Gray (G;S1D,MD,PJ) Cafnissonia eastwoodiae (Munz) Raven (C;SnD,SlD) Sporobolus flexuosus (Thurb.) Rydb. (U;SnD,WR) CdviU^hhia scapoidea (T. & G.) Raven (C;SnD,SlD) Sporobolus giganteus Nash Cafnissonia walkeri (Nels.) Raven (U;S1D,MD) Stipa arida Jones (U;WR) Oenothera brachycarpa (Gray) Britt. (U;MD,PJ,WR) Stipa comata Trin. & Rupr. (C;MD,PJ) Oenothera caespitosa Nutt. (C;SnD,MD,WR) Stipa Columbiana Macoun. (U;PJ) Oenothera pallida Lindl. ssp. pallida (C;SnD,MD,WR) 86 Great Basin Naturalist Vol. 43, No. 1

SHpa speciosa Trin. & Rupr. (U;PJ) Ranunculus cymbalaria Pursh (U;SS)

' Vulpia octoflora (Walt.) Rydb. (0;PJ,WR) Ranunculus testiculatus Grantz (0;DS)

. t'oLEMONIACEAE Gilia aggregata (Pursh) Spreng. (C;W) Amelanchier utahensis Koehne (C;WR,SS) Gilia congesta Hook. (0;S1D) Cercocarpus intricatus Wats. (G;PJ,RO) Gilia gunnisonii T. & G. (C;MD,WR) Cercocarpus montanus Raf. (G;PJ,PM,WR) Gilia inconspicua (Smith) Sweet (C;PJ) Coleogyne ramosissima Torr. (G;SnD) Gilia latifolia Wats. (R;RO,WR) Cowania mexicana D. Don (C;PJ) Gilia leptamena Gray (C;SnD,MD,PJ) Fallugia paradoxa (D. Don) Endl. (0;WR) Gilia polycladon Torr. in Emory (0;S1D) Physocarpus alternans (Jones) Howell Gilia pumila Nutt. (0;S1D) Physocarpus monogynus (Torr.) Gou. (U;PJ) Gilia roseata Rydb. (C;WR,RO) Purshia tridentata (Pursh) DC. (0;WR) Rosa woodsii Lindl. (R;PM) Gilia stenothyrsa Gray (0;PJ) Leptodactylon caespitosum Nutt. (U;WR) Rubiaceae Kellogg Leptodactylon pungens (Torr.) Nutt. (0;MD,WR,RO) Galium multiflorum (0;WR) Leptodactylon watsonii (Gray) Rydb. (U;WR,RO) Salicaceae Populus x acuminata Rydb. (U;WR,SS) Phlox austromontana Gov. (0;PJ) Popidus angustifolia James ex Torr. (0;WR) Phlox hoodii Rich. (C;MD,PJ) Populus fremontii Wats. (G;WR,SS) Phlox hngifolia Nutt. (0;S1D,MD) Populus tremuloides Michx. (0;PM) POLYGALACEAE Salix exigua Nutt. (G;SS) Polygala acanthoclada Gray (R;MD) Santalaceae Polygala subspinosa Wats. (G;SlD,PJ,RO) Comandra umbellata (L.) Nutt. var. pallida (DG.) POLYGONACEAE Jones (G;W,WR) Eriogonum alatum Torr. in Sitgr. (C;PJ) Saxifragaceae Eriogonum hatemanii Jones (C;PJ) Philadelphus microphyllus Gray (G;PJ,PM) Eriogonum. bicolor Jones (C;MD,PJ) Ribes cereum Dougl. (R;PJ,PM) Eriogonum Nutt. (C;S1D) cernuum Ribes leptanthum Gray (0;PM) Eriogonum corymbosum Benth. var. corymbosum Scrophulariaceae (G;S1D,MD) Castilleja chromosa Nels. (0;W) Eriogonum fasciculatum Benth. var. polifolium Castilleja linariaefolia Benth. (0;SS) (Benth. in DG.) T. & G. (R;MD) Castilleja scabrida Eastw. (G;W) Eriogonum (R;S1D) flexum Jones Cordylanthus kingii Wats. (0;PJ) Eriogonum gordonii Benth. (U;S1D,MD) Cordylanthus wrightii Gray Eriogonum hookeri Wats. (0;MD,PJ) Mimulus rubellus Gray (R;PM) Torr. var. Eriogonum inflatum & Frem. fusiforme Penstemon carnosus Pennell (G;PJ) (Small) Reveal (0;S1D,MD,PJ) Penstemon cyanocaulis Payson (U;WR) Eriogonum inflatum Torr. & Frem. var. inflatum Penstemon eatonii Gray (0;PJ,PM) (G;S1D,MD) Penstemon lentus Pennell (U;WR) jamesii Benth. var. Eriogonum flavescens Wats. Penstemon utahensis Eastw. (G;PJ) (G;MD,PJ,WR) Selaginellaceae Eriogonum leptocladon T. & G. var. leptocladon Selaginella mutica Eaton ex Underw. (U;HG) (C;S1D) Tamaricaceae Eriogonum microthecum Nutt. var. foliosum (T. & G.) Tamarix ramosissima Ledeb. (G;SS,WR) Reveal (0;S1D) Typhaceae var. Eriogonum ovalifolium Nutt. ovalifolium Typha latifolia L. (G;SS) (C;W,RO) Ulmaceae Eriogonum salsuginosum (Nutt.) Hook. (0;MD,PJ) Celtis reticulata Torr. (U;WR) Eriogonum var. shockleyi Wats. longilobum (Jones) Ulmus pumila L. (R;WR) Reveal (C;S1D,MD) Eriogonum smithii Reveal Eriogonum tumulosum (Bameby) Reveal (U;PJ) Literature Cited Eriogonum wetherillii Eastw. (0;W) Cronquist, a., a. Holmgren, N. Holmgren, and Polygonum aviculare L. (0;WR,DS) J. POLYPODIACEAE Reveal. 1972. Intermountain flora: vascular plants of the Intermountain West. vol. I. Hafner Adiantum capillus-veneris L. (U;HG) Publishing Gompany, Inc., New York. Cheilanthes feei Moore (0;HG) Gronquist, a., a. Holmgren, N. Holmgren, Reveal, Pellaea glabella Mett. ex Kuhn (R;HG) J. PORTULACEAE AND P. Holmgren. 1977. Intermountain flora: of the Intermountain West. vol. Portulaca oleracea L. (R;PJ,PM) vascular plants VI. Golumbia University Press, New York. Talinum validulum Greene (0;PJ) Ranunculaceae Neese, E., and S. L. Welsh. 1981. Astragalus section Aquilegia micrantha Eastw. (C;HG) Desperati (Leguminosae) and a new species from the of Utah. Rhodora 83:455-460. Clematis ligusticifolia Nutt. (G;WR) Welsh, S. L. 1978. Utah flora: Fabaceae (Leguminosae). Delphinium nuttallianum Pritz. ex Walp. (0;W) Great Basin Nat. 38:225-367. Delphinium scaposum Greene (0;PJ) January 1983 Harris: San Rafael Swell Flora 87

1980a. Utah flora: Malvaceae. Great Basin Nat. index of Utah vascular plant names. Great Basin 40:27-37. Nat. 41:1-108. 1980b. Utah flora: miscellaneous families. Great Welsh, S. L., and G. Moore. 1973. Utah plants: Basin Nat. 40:38-58. Tracheophyta. 3d ed. Brigham Young University 1981. New taxa of western plants— In tribute. Press, Provo, Utah. S. Brittonia 33:294-303. Welsh, L., and J. L. Reveal. 1977. Utah flora: Brassi- Welsh, S. L., N. D. Atwood, S. Goodrich, E. Neese, caceae (Cruciferae). Great Basin Nat. 37:297-364. K. H. Thorne, and B. Albee. 1981. Preliminary