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(Leguminosae): Nomenclatural Proposals and New Taxa Great Basin Naturalist Volume 58 Number 1 Article 5 1-30-1998 Astragalus (Leguminosae): nomenclatural proposals and new taxa Stanley L. Welsh Brigham Young University Follow this and additional works at: https://scholarsarchive.byu.edu/gbn Recommended Citation Welsh, Stanley L. (1998) "Astragalus (Leguminosae): nomenclatural proposals and new taxa," Great Basin Naturalist: Vol. 58 : No. 1 , Article 5. Available at: https://scholarsarchive.byu.edu/gbn/vol58/iss1/5 This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. Great Basin Naturalist 58(1), © 1998, pp. 45-53 ASTRAGALUS (LEGUMINOSAE): NOMENCLATURAL PROPOSALS AND NEW TAXA Stanley L. Welsh! ABSTRACT.-As part of an ongoing summary revision of Astragalus for the Flora North America project, several nomenclatural changes are indicated. Nomenclatural proposals include A. molybdenus val'. shultziorom (Barneby) Welsh, comb. nov.; A. australis var. aboriginorom (Richardson) Welsh, comb. nov.; A. australis var. cattoni (M.E. Jones) Welsh, comb. nov.; A. aU8tralis var. lepagei (Hulten) Welsh, comb. nov; A. australis var. muriei (Hulten) Welsh, comb. nov.; A. subcinereus var. sileranus (M.E. Jones) Welsh, comb. nov.; A. tegetariaides val'. anxius (Meinke & Kaye) Welsh, comb. nov.; A. ampullarioides (Welsh) Welsh, comb. nov.; A. cutlen (Barneby) Welsh, comb. nov.; and A. laccaliticus (M.E. Jones) Welsh, comb. nov. Proposals of new taxa include Astragalus sect. Scytocarpi subsect. Micl'ocymbi Welsh, subsed. nov., and A. sabulosus var. vehiculus 'Velsh, var. nov. A lectotype is selected for Phaca australis L. Key words: Astragalus, nomenclature, new taxa. Astragalus, with more than 350 species and appearance of the entire manuscript within the a great many infraspecific taxa, is perhaps the FNA publication schedule. Format is as under­ largest genus of North American plants. Its stood for the FNA publication; order of treat­ complexity has long been recognized as evi­ ment is phylogenetic as per Barneby, or as per denced by its tangled nomenclatural history. present modification. Experts and others interested in this vast genus have encountered enormous problems in deal­ Astragalus molybdenus Barneby, Leafl. \ill Bot. ing with it, especially prior to 1964. In that 6:70. 1950. Leadville milkvetcb. year Rupert Barneby, in his classic account (in Low, loosely matted, shortly caulescent my opinion, the most impressive taxonomic perennials, 0.5-6 (14) cm long, from extensive­ work of tbe century), untied the Gordian knot ly branching subterranean caudex branches. of nomenclature, typification, and classifica­ Pubescence strigulose-pilosulous, basifixed. tion of Astragalus for North America. Regard­ Stems largely subterranean, the aerial tips less of ,,,hen a taxonomic work is attempted, prostrate or ascending. Stipules 2-5 mm long, there will be shortfalls in information avail­ all connate-sheathing. Leaves 1.5-7 cm long; ability, in adequacy ofspecimens, in confluence leaflets (9) 17-25, 2-10 mm long, ovate, ovate­ of data from disparate regions, and in overall oblong, or elliptic, obtuse, mostly crowded, understanding through time. Despite those folded or involute. Peduncles 1-3 (6.5) cm problems, the Atlas of North American Astra­ long; racemes loosely 3- to 6-flowered, the axis galus (Barneby 1964) will stand for all time as scarcely elongating, 3-10 (15) mm long in a remarkable attempt to understand this huge fruit; bracts 2.5-5 mm long; pedicels 0.5-2 genus and as a tribute to Barneby's genius. mm long: bracteoles 0-2. Calyx 5.2-7 mm It is hoped that the proposals discussed here­ long, the tube campanulate, 3-4.2 mm long, in represent some helpful minor additions to tbe teeth subulate, 2-3 mm long. Flowers the work by Barneby, whose treatment is re­ 10.7-12.5 mm long, pink-purple, lilac, or flected in a large manuscript now in prepara­ wbitisb, the banner veined and suffused with tion for the Flora Nortb America (FNA) project lilac, recurved through ca 45', the keel tip by S.L. Welsh and R. Spellenberg. Included maculate. Pods ascending, sessile or nearly so, below are sufficient portions of that treatment 7-11 (12) mm long, 3-3.5 mm thick, obliquely to allow the current proposals to be put into ovoid or ovoid-ellipsoid, somewhat incurved, perspective and to be used by workers prior to l-loculed, strigulose. Ovnles 6. IDepartment of Botany and Range Science, and M,L. Bean Life Science Museum, Brigham Young Univer~ity, Provo, liT 84602, 45 46 GREAT BASIN NATURALIST [Volume 58 1. LeaHets of upper leaves 17-25; racemes 3- to 6­ & G.M. Armstrong 3651," holotype POM!; iso­ flowered; plants of central Colorado and Teton types YlO, NY!, WYO. * Co., Montana val'. molybdenus Leaflets of upper leaves 9-17; racemes mostly l- Flowering July, August. Alpine tundra and or 2 (exceptionally 3)-flowered; plants of the Salt krummholz or on talus, at 2865-3150 m, in Hiver Range, Lincoln Co" Wyoming . Salt River Range, Lincoln Co., Wyoming. ............................... var. shultziorum Plants ofthe 2 varieties are essentially iden­ tical in aspect, bnt the features noted in the &tragalus molybdenus var. molybdenus key appear to be substantial diagnostic ones. [based on: A. plumbeus Barneby, LeaR w: Bot. 5:195. 1949, non A. plumbeus Gontscharow]. Astragalus australis (L.) Lam., F!. Fr. 2:637. Dwarfalpine plants, the caudex deeply sub­ 1778. Subarctic milk:vetch. terranean, the branches rhizomatous. Stipules [Phaca australis L., Mant. PI. 1:103. 1767J. 2-5 mm long. Leaves 1.5-7 mm long, (the Moderate, caulescent perennial, (10) 20-30 uppermost) with 17-25 leaflets, 2-10 mm long. cm tall, from a superficial caudex. Pubescence Peduncles 1-3 (6.5) cm long, the racemes 3- to silky-strigose, villous, or villous-tomentose, 6-flowered. Calyx 5.2-7 mm long, the tube basifixed. Stems erect or ascending, few to campanulate, 3-4.2 mm long, the teeth subu­ several. Stipules (1) 2-7 (11) mm long, often late, 2-3 mm long. Flowers 10.7-12.5 mm veined, semicoriaceous, at least the lowermost long, pink-purple, lilac, or whitish, the banner connate-sheathing. Leaves (1) 2-7 (10) cm recurved through ca 45'. Pods rather abruptly long; leaflets (5) 7-15, 3-28 (35) mm long, 1-7 contracted into a short beak. Ovules 6. *Type: (8) mm wide, oblong, linear-elliptic, elliptic, or "Colorado: about 4 miles east of Leadville, linear-oblong, acute, villous to glabrate on both Lake County Ripley & Barneby No. 9994 sides. Peduncles 2-10 (14) cm long; racemes ... west slope of Mosquito Pass, east of Lead­ 2- to 40-flowered, rather compact and ascend­ ville, No. 10045"; syntypes CAS!; isosyntypes ing at anthesis, the axis 1-15 em long in fruit; GH!, K, NY!, POM!, RM!, RSA, US, WTU.* bracts 1.2-5 mm long; pedicels 0.8-3.5 mm Flowering July, August. Alpine tundra com­ long; bracteoles O. Calyx 3.7-6.4 mm long, the munity at 3780-3965 m, along the Continental tube 2.1-5 mm long, campanulate, villous, the Divide, along the boundaries between Gunni­ teeth 1-3 mm long, subulate. Flowers 7.5­ son-Pitkin, Lake-Park, and Park-Summit coun­ 14.5 mm long, ochroleucous or snffnsed with ties, in central Colorado, and disjunct in Teton pink, the wing petals bilobed apically. the ban­ Co., Montana. ner recurved through 40-50°. Pods pendu­ lous, stipitate, the stipe 2.5-8 (10) mm long, &tragalus molybdenus var. shultziorum the body obliquely and narrowly elliptic in (Barneby) Welsh comb. nov. outline, 13-27 mm long, 3-9 (11) mm wide, [based on: A. shultziorum Barneby, Bdttonia 33:156. semibilocular, the septum 0-0.6 mm wide, 19.58]. glabrons or pubescent. Ovules 8-16; 2n ; 16, Dwarf alpine plants, the caudex deeply 32, 48. *Type: "Habitat in alpinis Helvetiae, subterranean, the branches rhizomatous. Stip­ ltaliae, Gallo Provinciae," lectotype here des­ ules 2-3 mm long. Leaves 1.5-7 (8) cm long, ignated, illustration of 'l\.stragaloides Alpina the uppermost with 9-15 (17) leaflets, 2-7 mm supina glabra, foliis auctioribus," in Tilli, Cat. long, lance- or ovate-elliptic, obtuse to acute, P!. Hart. Pisani 19. 5.14. f. 1. 1723!* mostly distant, flat or loosely folded. Pedun­ American materials of A. australis are por­ cles (0.5) 1-4 cm long, the racemes (usually 2-) tions of a vast circumboreal species complex 1- to 3-flowered. Calyx (5) 5.7-6.7 mm long, the demonstrating great variability, beginning tube campaoulate, (3) 3.4-4 mm long, the teeth with the typical material in southern Europe subulate, 1.8-3.3 mm long. Flowers 11-12 and extending eastward. Asiatic plants passing mm long, whitish lavender tinged, the banner under the names A. tugaranovii Basilevskaja, veined and snffused with lilac, recurved A. gorodkovii Jurtsev, A. tolmaczevii Jurtsev, through ca 50', the keel tip maculate. Pods and A. kolymensis Jurtsev (Korobkov et al. tapering to an elongate beak. Ovules 8 or 9. 1986) belong to this complex, with the entities *Type: "Wyoming. Lincoln County: stony hill­ having the same degree of morphological top, 9500 ft, mountains near Cottonwood integrity (or lack thereof) as the American Lake, E of Smoot, 31 Jul1923 (fl), E.B. Payson materials. Varietal segregation within the North 1998J ASTRAGALUS NOMENCLATURE 47 American variants has been based on differ­ long, whitish to purplish. Pod stipe 4-6 mm ences in pubescence, leaflet shape, and pod long, the body obliquely ellipsoid to narrowly size and shape. Several varieties have been oblong, 11-24 mm long, 4-7 mm wide, the proposed, with the best summary that of valves glabrous to occasionally strigose.
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