Polish Botanical Journal 57(2): 375–425, 2012

DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. ( BASIN) COMMON IN TURKISH INLAND WATERS

CÜNEYT NADIR SOLAK & AGATA Z. WOJTAL

Abstract. In a study of diatoms from ten springs and fi ve streams of Türkmen Mt. we identifi ed over 300 diatom species, 59 of which are among the most commonly reported diatoms from . Several of these diatoms are truly widespread but can also be accompanied or replaced by taxa of more limited distribution. Some commonly reported diatom species [e.g., Caloneis bacillum (Grunow) Cleve, Cymbella leptoceros (Ehrenberg) Kützing, Pinnularia viridis (Nitzsch) Ehrenberg] were not found in the material studied, but morphologically similar diatoms (e.g., P. viridiformis Krammer, Cymbella subleptoceros Krammer) were found in the springs and streams. We identifi ed some taxa included previously in the broad Caloneis bacillum concept [i.e., C. lancettula (Schulz) Lange-Bertalot & Witkowski, C. fontinalis (Grunow) Lange-Bertalot & Reichardt]. Previous records of several diatoms regarded as common but which we did not confi rm in the Türkmen Mt. area may very well be correct, as the reports from Turkey are from very diverse environments, including large rivers and lakes. However, the published data are largely unverifi able due to a general lack of collections and/or documentation. All the diatoms presented here are documented by light micrographs, and brief notes on their dimensions, distribution and ecology are provided. Caloneis lancettula, C. fontinalis, Cymbella affi niformis Krammer, C. subleptoceros and Pinnularia viridiformis are reported for the fi rst time from Turkey. Key words: Bacillariophyta, taxonomy, ecology, springs, streams, mountains, distribution, Turkey Cüneyt Nadir Solak, Department of Biology, Dumlupınar University, 43100, Kütahya, Turkey & Department of Palaeoceanology, Institute of Marine Sciences, University of Szczecin, Mickiewicza 18, 70-383, Szczecin, Poland; e-mail: [email protected] Agata Z. Wojtal, Department of Phycology, W. Szafer Institute of Botany, Polish Academy of Sciences, Lubicz 46, 31-512, Kraków, Poland; e-mail: [email protected]

INTRODUCTION

The structure of diatom assemblages refl ects en- shown, however, that the distribution of several vironmental conditions and the water quality of diatoms is infl uenced by geographic and climatic springs, streams (Battegazzore et al. 2004; Canto- factors (Vanormelingen et al. 2008). nati & Spitale 2009; Wojtal et al. 2009) and rivers Situated between large continents (Eurasia and (Vilbaste 2004; Blanco et al. 2006; Ács et al. 2009; Africa), Turkey comprises areas of very diverse Barinova & Tavassi 2009; Kelly et al. 2009; Angeli geological, climatic and environmental conditions, et al. 2010; Torrisi et al. 2010; Wojtal & Sobczyk including the Mediterranean region with many en- 2012). Diatoms are also one of the most common demic vascular plants (Anonymous 2003; Solak producers of organic matter in springs and small et al. 2012), so we should expect that other organ- streams (Cantonati et al. 2006; Wojtal 2009) and isms including diatoms will show high diversity one of the most important components of aquatic in this region. biodiversity (e.g., Gillett et al. 2011). The world- Although long lists of algal names from Turkey wide application of the most commonly used keys have been published (Gönülol et al. 1996; Aysel (i.e., Patrick & Reimer (1966); Krammer & Lange- 2005) there has been relatively little taxonomic study Bertalot 1986–1991b) has created the impression of the diatoms (Solak et al. 2012). Most publications that a large part of the European diatom fl ora is merely list the observed organisms and are chiefl y of cosmopolitan distribution, due to the common concerned with water quality (Barlas et al. 2001, practice of fi tting the taxon to the name (Van de 2002; Dalkıran et al. 2008; Gürbüz & Kıvrak 2002; Vijver et al. 2004). High-resolution taxonomy has Kalyoncu 2002; Karacaoğlu et al. 2008; Solak 2011; 376 POLISH BOTANICAL JOURNAL 57(2). 2012

Solak et al. 2007a, b, 2009; Kalyoncu et al. 2009) and Western Mediterranean, Antalya, Burdur, Eastern fl oristic or seasonal changes in lakes, dam reservoirs Mediterranean, Seyhan, Asi, Ceyhan), and some large rivers (Yıldız 1987a; Altuner 1988; – Inner Anatolia zone (10.9°C, 429.7 mm; Şen 1988b; Şen & Çetin 1988; Altuner & Gürbüz Susurluk, Sakarya, Konya closed basin, part of 1989; Elmacı & Obalı 1992, 1998; Kolaylı et al. Seyhan, Akarçay and Kızılırmak), 1998; Şahin 1998; Aksın et al. 1999; Çevik et al. – zone (13.7°C, 1175.8 mm; Western 1999, 2007; Obalı & Atıcı 2000; Yavuz & Çetin Black Sea, Yeşilırmak, East Black Sea, part of 2000; Atıcı et al. 2003; Soylu & Gönülol 2003; Kızılırmak), Pala-Toprak & Çağlar 2008; Akbulut 2010). – Eastern Anatolia zone (7.2°C, 417.5 mm; Here we have prepared verifi able taxonomic Van closed basin, Fırat, Çoruh, ), records of the diatoms most commonly reported – Southeastern Anatolia zone (15.2°C, from Turkey, as the fi rst step toward a detailed 620.6 mm; Dicle). elaboration of diatom distribution and taxonomy The water of the river basins differs in quantity, in this very poorly recognized area (Solak et al. from lowest in the Akarçay basin to highest in the 2012). Fırat and Dicle basins (Akin & Akin 2007), and also quality, from most polluted in the Marmara and STUDY AREA Aegean basins to pristine in the Antalya basin (Solak et al. 2012). The Fırat and East Black Sea basins There are 26 river basins in seven climatic zones have drawn more attention because of their impor- in Turkey, listed here with their mean annual tem- tance in the nation’s water supply, with the largest perature and precipitation (Ünal et al. 2003): volumes of freshwater in the country. On the other – Marmara zone (13.7°C, 686.2 mm; Meriç- hand, the only published information about the algae Ergene, Marmara and part of Susurluk), of the Dicle basin is in one study of the Devegeçidi – Aegean zone (15.4°C, 682.0 mm; Northern reservoir (Baykal et al. 2004) and another about Aegean, Gediz, Küçük Menderes, Büyük Men- some nearby wetlands (Açıkgöz et al. 2007). deres), In large cities administered by metropolitan – Mediterranean zone (18.7°C, 797.2 mm; municipal authorities, domestic wastewater is

Table 1. Sampling stations of Türkmen Mountain (* – spring, ** – stream).

Longitude Latitude Altitude Stations E N m a.s.l. Darıpınar* 30°15′23,53″ 39°22′21,52″ 1298 Çobanlar plt.* 30°20′25,26″ 39°27′48,42″ 1512 Dümbüldek plt.* 30°13′18,12″ 39°21′51,60″ 1130 Gölcük plt.* 30°22′25,92″ 39°26′24,84″ 1627 Güllüdere plt.* 30°21′47,16″ 39°24′34,92″ 1366 İdris plt.* 30°24′44,94″ 39°24′00,96″ 1421 İnli plt.* 30°20′02,28″ 39°27′48,42″ 1572 Lütfi ye-Güllüdere** 30°19′00,30″ 39°23′42,90″ 1396 Söğüt-Lütfi ye** 30°20′09,42″ 39°24′14,76″ 1340 Güllüdere-Türkmenbaba** 30°21′53,76″ 39°23′42,30″ 1358 Söğüt plt.* 30°15′46,56″ 39°22′13,74″ 1361 Türkmenbaba plt.* 30°23′01,74″ 39°26′10,14″ 1710 Uluköy plt.* 30°09′45,03″ 39°29′13,99″ 954 Seydi** 30°15′57,20″ 39°22′06,02″ 990 Akındere** 30°21′42,34″ 39°27′12,26″ 1381 C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 377

Fig. 1. Study area and the distribution of river basins in Turkey: 1 – Meriç-Ergene River Basin, 2 – Marmara River Basin, 3 – Susurluk River Basin, 4 – Northern Egean River Basin, 5 – Basin, 6 – Küçük Menderes River Basin, 7 – Büyük Menderes River Basin, 8 – Western Mediterranean River Basin, 9 – Antalya River Basin, 10 – Burdur River Basin, 11 – Akarçay River Basin, 12 – Sakarya River Basin, 13 – Western Black Sea River Basin, 14 – Yeşilırmak River Basin, 15 – Kızılırmak River Basin, 16 – Konya Closed River Basin, 17 – Eastern Mediterranean River Basin, 18 – Basin, 19 – Asi River Basin, 20 – Basin, 21 – Fırat River Basin, 22 – East Black Sea River Basin, 23 – Çoruh River Basin, 24 – Aras River Basin, 25 – Van Closed River Basin, 26 – Dicle River Basin (according to Akın & Akın 2007). treated before being released into nearby streams, Sakarya basin can be found in 44 publications, but many smaller towns have no or underutilized accounting for over 13% of all the papers men- water treatment facilities. Despite the presence of tioning diatoms from Turkey (Solak et al. 2012). sewage treatment plants, domestic pollution seems As in other basins, the main interest has focused on to be increasing in rivers that fl ow near large cities lakes and large rivers (Solak et al. 2012), whereas such as Ankara, Eskişehir and Kütahya (belonging springs and small streams have been much less to the Sakarya basin; Atıcı & Ahıska 2005; Solak studied. Due to that focus, there has been no phy- et al. 2009; Solak 2011), and the aquatic environ- cological study of Türkmen Mt. and Murat Mt., ment is increasingly threatened by wastewater and where the least polluted water bodies can still be nutrients from quickly developing urban areas and found. The localities we studied (Table 1) are situ- agriculture. ated at 954–1710 m a.s.l. and represent typical Türkmen Mt. (Aegean zone) is located in the rheocrene springs and small streams of the area. Sakarya basin, which is moderately rich in waters Some stations (Türkmenbaba, Güllüdere, Inli, (Akin & Akin 2007) and joins the Aegean and Inner Çobanlar) have near-natural conditions, and others Anatolian zones (Fig. 1). It is near the borders of (Darıpınar, Dümbüldek, Güllüdere-Türkmenbaba, the Irano-Turanean, Mediterranean and European- Lütfi ye-Güllüdere) are eutrophic due to agricul- Siberian phytogeographical areas (Anonymous tural and other human activities. 2004). The Sakarya basin is an agricultural and industrial area. Anthropogenic pollution of waters MATERIAL AND METHODS is caused mainly by domestic sewage and indus- trial wastewater, and from irrigation return water. The diatoms of ten mountain springs and fi ve small Türkmen Mt. and Murat Mt. are among the areas streams were investigated between May and December best preserved from anthropogenic changes in the 2007 (Table 1). The fi rst author collected samples from Sakarya basin. Information about diatoms in the the epilithon, periphyton and epipelon. Fifty samples 378 POLISH BOTANICAL JOURNAL 57(2). 2012 from submerged stones were collected monthly by lanceolatum (Brebisson ex Kützing) Round & brushing for epilithic samples and with a pipette aspi- Bukhtiyarova. Gönülol et al. (1996) prepared the rator for epipelon samples. The samples were boiled in fi rst Checklist of the Freshwater Algae of Turkey. H2O2 and HCl to remove calcium carbonate and organic Aysel (2005) updated the list, giving 63 Cosci- matter. After washing several times with distilled water nodiscophyceae, 102 Fragillariophyceae and 620 the material was air-dried on coverglasses and mounted in Naphrax®. The diatoms were observed with Nikon Bacillariophyceae and provided the list of 785 Eclipse 600 and Nikon Eclipse 80i light microscopes diatom taxa; more recent data (Baykal et al. 2009, (LM). Micrographs were taken with Nikon DS-5M and Solak & Wojtal 2010) increase the number of dia- Nikon DS-L1 cameras. Scanning electron microscopy toms reported from Turkey to 799 (not 631 as (SEM) observations used cleaned, gold-coated material. misquoted in Solak et al. 2012). Inclusion of an SEM micrographs were taken in the Laboratory of Field additional publication by Brunnthaler (1903) and Emission Microscopy, Scanning Electron Microscopy our identifi cation of Caloneis fontinalis, C. lancet- and Microanalysis (Institute of Geological Sciences of tula, Cymbella affi niformis, C. subleptoceros and the Jagiellonian University and additionally at Goethe Pinnularia viridiformis, not previously reported University in Frankfurt). from Turkey, extend the list of diatom taxa re- Species reported from Turkey in at least 10% of the literature by Solak et al. (2012) are noted here as diatoms ported from Turkish inland waters in the course most commonly reported from Turkey. The distribution of more than 150 years, raising it to 807. Many of of diatoms in Turkey is given according to Gönülol et al. the diatoms commonly reported from very diverse (1996) and Aysel (2005), separately for the given river bodies of water in Turkey are taxa known from basins. The literature used for identifi cation, dimensions, European and Asiatic areas (Solak et al. 2012). distribution and ecology (Hustedt 1930; Patrick & Reimer This indicates their broad range of tolerance and 1966, 1975; Krammer & Lange-Bertalot 1986–1991b; distribution but it also points to a problem. Because Krammer 1997, 2002, 2003; Lange-Bertalot 2001; of the lack of taxonomic monographs on the whole Håkansson 2002; Levkov 2009; Hofmann et al. 2011) diatom fl ora of Turkey or regional fl oras, Turkish is indicated for every species separately. Species and in- fraspecifi c taxa are arranged alphabetically in the text and phycologists have had to rely on various European plates. The material studied is deposited in the collection or other non-native fl oras. This may be why mainly of the Department of Biology of Dumlupınar University diatoms known from Europe have been identifi ed (Turkey), the Department of Phycology of the W. Szafer and reported from Turkey. Institute of Botany, Polish Academy of Sciences (Poland), Some of the commonly reported diatom names and the Department of Geological Science of Szczecin (e.g., Achnanthidium minutissimum, Nitzschia University (Poland). palea, Sellaphora pupula) probably concern species complexes such as Achnanthidium minutissimum RESULTS AND DISCUSSION complex, Nitzschia palea complex, or Sellaphora pupula complex (Mann et al. 2007, Hlúbiková In this study we identifi ed 304 diatom species in et al. 2009, Wojtal et al. 2011). These diatoms were 10 springs and 5 streams of Türkmen Mt. The reported from very diverse water bodies, implying most common were representatives of the genera broad ecological tolerance ranges. Cymbella, Fragilaria, Gomphonema, Navicula Unfortunately, the materials used for most of and Nitzschia; the distribution of Chamaepinnu- the previous studies in Turkey cannot be re-inves- laria, Eunotia, Neidium, Pinnularia species was tigated because of the lack of collections where the much more limited to the near-natural localities materials should have been deposited. Of the 129 (Çobanlar, Gölcük, Türkmenbaba). In terms of diatom species classifi ed previously as common in commonness and abundance the most important Turkey in the Checklist of the Freshwater Algae diatoms in the springs and streams of Türkmen of Turkey and other sources (Solak et al. 2012) Mt. were Cymbella neocistula Krammer, C. hel- we identifi ed 59 in the material collected from vetica Kützing, Navicula cryptocephala Kützing, Türkmen Mt. These 59 are known from very di- N. cryptotenella Lange-Bertalot and Planothidium verse water bodies sometimes quite different from C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 379 the springs and streams of Turkey. It is impossible 1990a, b, 1991, 1994, 1996; Altuner & Pabucçu 1993, to state defi nitively that a given taxon is not present 1994; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz in Turkey’s many diverse aquatic environments; 2000; Gürbüz & Altuner 2000), Gediz (Gezerler et al. some that we did not fi nd in the study area may 1996), Kızılırmak (Özesm 1987; Kılınç & Dere 1988; Yıldız & Özkıran 1991; Dere & Sıvacı 1994; Kılınç occur there in low abundance. 1998; Kılınç & Sıvacı 2001; Akbulut 2003), Konya Judging by the apparent morphological simi- Closed River Basin (Skuja 1937; Cirik et al. 1991; Temel larity of specimens described in older reports, 1992, 1996; Yıldız et al. 1994; Demirsoy 1996; Mita- some previously reported diatoms may actually mura et al. 1997; Akköz & Obalı 1998; Elmacı & Obalı be taxa unknown from Turkey until now but which 1998; Morkoyunlu et al. 1996), Küçük Menderes (Aysel we found commonly in the material we studied et al. 1992, 1998a, b, 2001, 2002), Sakarya (Obalı 1984; (Caloneis fontinalis, C. lancettula, Cymbella affi ni- Yıldız 1987a; Obalı et al. 1989; Yıldız & Özkıran 1994; formis, C. subleptoceros, Pinnularia viridiformis). Atıcı & Yıldız 1996; Aykulu et al. 1999; Atıcı 2002; On the other hand, several of the diatoms most Yardımcı & Temel 2000), Susurluk (Skuja 1937; Dere et al. 2002; Karacaören et al. 2004). commonly reported from Turkey seem to be truly widespread, though they may also be accompanied GENERAL DISTRIBUTION AND ECOLOGY. Wide- or replaced by taxa of more limited distribution. spread, tychoplanktonic, mesosaprobous species Some of the 245 taxa we found in the material from (Denys 1991; Krammer & Lange-Bertalot 1991b; Türkmen Mt. need further taxonomical studies and Hofmann 1994; Van Dam et al. 1994), classifi ed as will be published separately. a circumneutral or alkaliphilous diatom (Håkans- son 1993; Van Dam et al. 1994). 1. Achnanthidium minutissimum (Kützing) Czarnecki Figs 2, 3, 123 & 124 2. Amphora ovalis (Kützing) Kützing Achnanthes minutissima Kützing Figs 4 & 5 Ref. Hustedt 1930 (p. 198, Fig. 274); Patrick & Re- Ref. Levkov 2009 (p. 96, Figs 1: 1–5; 2: 1–6); Hofmann imer 1966 (p. 253, Fig. 16: 9–10); Krammer & Lange- et al. 2011(p. 98, Fig. 90: 1–5); Krammer & Lange- Bertalot 1991b (p. 56, Fig. 32: 1–24); Hofmann et al. Bertalot 1986 (p. 344, Fig. 149: 1, 2). 2011 (p. 83, Fig. 23: 15–21). DIMENSIONS. Valves 47.4–64.4 μm long and DIMENSIONS. Valves 6–24 μm in length, 2.5– 9.4–15.7 μm wide, 9–13 striae in 10 μm. 4.0 μm wide, 24–30 striae in 10 μm. DISTRIBUTION IN TURKMEN MT. Akındere, DISTRIBUTION IN TURKMEN MT. Türkmenbaba, Güllüdere. Darıpınar, İdrıs, Söğüt-Lütfi ye. DISTRIBUTION IN TURKEY. Lakes and rivers in DISTRIBUTION IN TURKEY. Lakes and rivers most of the river basins (Fig. 1). in Eastern Anatolia (Fırat, Aras, Çoruh), Black Sea (East Black Sea River Basin), Inner Anatolia RIVER BASINS. Akarçay (Gönülol & Obalı 1986; Şen (Akarçay, Kızılırmak, Konya Closed River Basin, et al. 1994), Antalya (Skuja 1937; Vardar & Güner 1972; Sakarya) and Egean (Küçük Menderes and Susur- Conk & Cirik 1991; Morkoyunlu & Ertan 1995a, b; luk) river basins (Fig. 1). Ertan et al. 1996; Morkoyunlu et al. 1996; Savaş & Cirik 1997; Yüce et al. 1997; Ertan & Morkoyunlu 1998), RIVER BASINS. Akarçay (Gönülol & Obalı 1986; Şen Aras (Altuner 1988; Akbulut & Yıldız 2002; Gürbüz et al. 1994), Aras (Altuner 1988; Akbulut & Yıldız 2002; et al. 2002a, b; Kıvrak & Gürbüz 2005), Asi (Şen et al. Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Büyük 1997), Çoruh (Altuner 1984; Altuner & Aykulu 1987; Menderes (Skuja 1937; Güner 1966), Çoruh (Altuner Atıcı & Obalı 1999), Dicle (Şen 1988a; Şen et al. 1999; 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; Şen Baykal et al. 2004), East Black Sea River Basin (Şahin et al. 1999; Baykal et al. 2004), East Black Sea River & Gönülol 1997; Kolaylı et al. 1998; Kara & Şahin 2001; Basin (Şahin 1992, 1994, 1998a, b, 2000, 2001, 2002, Şahin 1992, 1994, 1998a, b, 2000, 2001, 2002, 2003), 2003; Şahin & Gönülol 1997; Kolaylı et al. 1998; Kara Fırat (Altuner & Gürbüz 1988, 1989, 1990a, b, 1991, & Şahin 2001), Fırat (Altuner & Gürbüz 1988, 1989, 1994, 1996; Şen & Çetin 1988; Altuner & Pabucçu 1993, 380 POLISH BOTANICAL JOURNAL 57(2). 2012

1994; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz Aras, Çoruh), Eagen (Gediz, Susurluk, Küçük 2000; Gürbüz & Altuner 2000), Gediz (Cirik 1994; Menderes) and Inner Anatolia (Konya Closed İşbakan et al. 1998; Gezerler et al. 1999), Kızılırmak River Basin, Kızılırmak, Sakarya) river basins (Kılınç & Dere 1988; Gönülol & Çomak 1990, 1992; (Fig. 1). Gönülol 1993; Yıldız & Özkıran 1991; Gönülol & Ar- slan 1992; Hasbenli & Yıldız 1993; Dere & Sıvacı 1994; RIVER BASINS. Aras (Altuner 1988; Akbulut & Yıldız Kılınç 1998; Kılınç & Sıvacı 2001), Konya Closed River 2002; Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Basin (Skuja 1937; Yıldız 1984a, b, 1985b; 1986a, b; Çoruh (Altuner 1984; Altuner & Aykulu 1987), Dicle Cirik et al. 1991; Temel 1992, 1996; Yıldız et al. 1994; (Şen 1988a; Şen et al. 1999; Baykal et al. 2004), East Demirsoy 1996; Mitamura et al. 1997; Akköz & Obalı Black Sea River Basin (Şahin 1992, 1994, 1998a, b, 1998; Elmacı & Obalı 1998; Morkoyunlu et al. 1999), 2000, 2001, 2002, 2003; Şahin & Gönülol 1997; Kolaylı Küçük Menderes (Skuja 1937; Kolaylı & Güner 1988; et al. 1998; Kara & Şahin 2001), Fırat (Altuner Sukatar & Güner 1988; Aysel et al. 1998a, b, 2001, 2002, & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, 1996; 2005; Gezerler et al. 1992, 1995; Balık & Gezerler-Şipal Altuner & Pabucçu 1993, 1994; Şen et al. 1995; Çetin 1995; Erener et al. 1995), Marmara (Albay & Aykulu & Şen 1998, 2004; Gürbüz 2000; Gürbüz & Altuner 1994a, b; Temel 1994, 1996, 1999, 2001, 2002, 2003), 2000), Gediz (Gezerler et al. 1996), Kızılırmak (Kılınç North Aegean (Güner 1970; Gezerler 1986; Gezerler- & Dere 1988; Yıldız & Özkıran 1991; Gönülol & Ar- Şipal et al. 1996), Sakarya (Gönülol & Aykulu 1984; slan 1992; Dere & Sıvacı 1994; Gönülol & Obalı 1998; Obalı 1984; Ünal 1984; Gönülol 1985a, b, 1987; Yıldız Kılınç 1998; Gönülol & Çomak 1990, 1992; Gönülol 1987a, b; Dere 1989; Obalı et al. 1989; Atıcı & Yıldız 1993; Kılınç & Sıvacı 2001), Konya Closed River Basin 1996; Aykulu et al. 1999; Temel 2001; Atıcı 2002), (Skuja 1937; Yıldız 1984a, b, 1985a, b, 1986a, b, 1987b; Seyhan (Kandemir-Çevik et al. 1994), Susurluk (Skuja Cirik et al. 1991; Temel 1992, Yıldız et al. 1994; Mita- 1937; Gezerler et al. 1994; Dere et al. 2002; Karacaoğlu mura et al. 1997; Demirsoy 1996; Akköz & Obalı 1998; et al. 2004), West Black Sea River Basin (Aysel et al. Elmacı & Obalı 1998; Morkoyunlu et al. 1996), Küçük 1992; Öztürk 1994), Yeşilırmak (Yazıcı & Gönülol 1994; Menderes (Aysel et al. 1992, 1998a, b, 2001, 2002), Altuner & Pabucçu 1996; Kır & Polat 1996; Gönülol Sakarya (Gönülol & Aykulu 1984; Obalı 1984; Ünal & Obalı 1998). 1984; Gönülol 1985a, b, 1987; Yıldız 1987a; Dere 1989; Obalı et al. 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız GENERAL DISTRIBUTION AND ECOLOGY. 1996; Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı Tycho planktonic, mesosaprobous and α-meso- 2002), Seyhan (Kandemir-Çevik et al. 1994), Susurluk eutraphentic species (Lange-Bertalot 1979; (Skuja 1937; Dere et al. 2002; Karacaören et al. 2004), Denys 1991), classifi ed as an alkaliphilous diatom Yeşilırmak (Altuner & Pabuçcu 1996). (Håkansson 1993; Van Dam et al. 1994). GENERAL DISTRIBUTION AND ECOLOGY. Wide- spread, tychoplanktonic, mesosaprobous and eu- 3. Amphora pediculus (Kützing) Grunow traphentic species (Denys 1991; Hofmann 1994; Figs 6–8 & 125–127 Van Dam et al. 1994; Levkov 2009), classifi ed Amphora ovalis var. pediculus Kützing as an alkaliphilous (Håkansson 1993; Van Dam Ref. Hustedt 1930 (p. 343, Fig. 629); Patrick & Re- et al. 1994). imer 1975 (p. 253, Fig. 16: 9–10); Krammer & Lange- Bertalot 1986 (p. 346, Fig. 150: 8–13); Levkov 2009 4. Caloneis fontinalis (Grunow) Lange-Bertalot (p. 101, Figs. 55: 31–34; 78: 40–47); Hofmann et al. & Reichardt Figs 9 & 10 2011 (p. 98, Fig. 91: 29–33). Ref. Hofmann et al. 2011 (p. 117, Fig. 67: 15–19). DIMENSIONS. Valves 7–15 μm long and 2–4 μm wide, 14–24 striae in 10 μm. DIMENSIONS. Valves 12.5–16.0 μm long and 4.0–4.7 μm wide, 24–25 striae in 10 μm. DISTRIBUTION IN TURKMEN MT. Dümbüldek, Darıpınar, İdris. DISTRIBUTION IN TURKMEN MT. Dümbüldek, İdris. DISTRIBUTION IN TURKEY. Lakes and riv- ers in Black Sea (East Black Sea River Basin, DISTRIBUTION IN TURKEY. New for fl ora of Kızılırmak, Yeşilırmak), Eastern Anatolia (Fırat, Turkey. C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 381

Figs 2–21. 2 & 3 – Achnanthidium minutissimum (Kützing) Czarnecki, 4 & 5 – Amphora ovalis (Kützing) Kützing, 6–8 – A. pe- diculus (Kützing) Grunow, 9 & 10 – Caloneis fontinalis (Grunow) Lange-Bertalot & Reichardt, 11–14 – C. lancettula (Schulz) Lange-Bertalot & Witkowski, 15 – Cocconeis pediculus Ehrenberg, 16 & 17 – C. placentula var. euglypta (Ehrenberg) Grunow, 18 – C. placentula var. lineata (Ehrenberg) Van Heurck, 19 – Craticula cuspidata (Kützing) D. G. Mann, 20 & 21 – Cyclotella ocellata Pantocsek. Scale bar = 10 μm. 382 POLISH BOTANICAL JOURNAL 57(2). 2012

GENERAL DISTRIBUTION AND ECOLOGY. Ac- & Gönülol 1997; Kolaylı et al. 1998; Kara & Şahin cording to Hofmann et al. (2011) Caloneis fon- 2001), Fırat (Altuner & Gürbüz 1988, 1989, 1990a, 1991; tinalis is widespread species occurring locally in Altuner & Pabucçu 1993, 1994; Şen et al. 1995; Çetin large number in calcium rich, mesotrophic and & Şen 1998, 2004; Gürbüz 2000; Gürbüz & Altuner 2000), Kızılırmak (Yıldız & Özkıran 1991; Gönülol eutrophic waters of low saprobity; most common & Çomak 1990, 1992; Gönülol 1993; Dere & Sıvacı in springs and running waters. 1994; Gönülol & Obalı 1998), Konya Closed River Basin (Skuja 1937; Yıldız 1984a, b, 1985a, b, 1986a, b, 1987b; 5. Caloneis lancettula (Schulz) Lange-Bertalot Cirik et al. 1991; Temel 1992, 1996; Yıldız et al. 1994; & Witkowski Figs 11–14 Demirsoy 1996; Morkoyunlu et al. 1996; Mitamura et al. 1997; Akköz & Obalı 1998; Elmacı & Obalı 1998), Ref. Hofmann et al. 2011 (p. 117, Fig. 67: 1–9). Sakarya (Obalı 1984; Yıldız 1987a; Obalı et al. 1989; DIMENSIONS. Valves 14–24 μm long and 4.0– Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu 5.3 μm wide, 22–30 striae in 10 μm. et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), Seyhan (Kandemir-Çevik et al. 1994), Susurluk (Skuja 1937; DISTRIBUTION IN TURKMEN MT. Dümbüldek. Dere et al. 2002; Karacaören et al. 2004), Yeşilırmak (Yazıcı & Gönülol 1994; Kır & Polat 1996). DISTRIBUTION IN TURKEY. New for fl ora of Turkey. GENERAL DISTRIBUTION AND ECOLOGY. Ty- choplanktonic, mesosaprobous and eutraphentic GENERAL DISTRIBUTION AND ECOLOGY. Caloneis lancettula is a common diatom in species (Denys 1991; Hofmann 1994; Van Dam α-mesotrophic up to eutrophic waters; good in- et al. 1994), classifi ed as an alkaliphilous diatom dicator of elevated trophic level (Hofmann et al. (Håkansson 1993; Van Dam et al. 1994). 2011). 7. Cocconeis placentula var. euglypta (Ehrenberg) 6. Cocconeis pediculus Ehrenberg Grunow Figs 16 & 17 Figs 15 & 128–130 Cocconeis placentula var. euglypta (Ehrenberg) Cleve Ref. Hustedt 1930 (p. 188, Fig. 259); Patrick & Reimer Ref. Hustedt 1930 (p. 190, Fig. 261); Patrick & Reimer 1966 (p. 240, Fig. 15: 3–4); Krammer & Lange-Bertalot 1966 (p. 241, Fig. 15: 8); Krammer & Lange-Bertalot 1991b (p. 88, Fig. 57: 1–4); Hofmann et al. 2011 (p. 132, 1991b (p. 87, Figs 53: 1–14; 54: 12). Fig. 19: 17–19). DIMENSIONS. Valves 12–40 μm long and DIMENSIONS. Valves 15–24 μm long and 7–25 μm wide, 16–26 striae in 10 μm. 11–20 μm wide, 14–20 striae in 10 μm. DISTRIBUTION IN TURKMEN MT. İdris. DISTRIBUTION IN TURKMEN MT. Türkmen- baba. DISTRIBUTION IN TURKEY. Lakes and rivers in most of the river basins (Fig. 1). DISTRIBUTION IN TURKEY. Lakes and rivers in Inner Anatolia (Konya Closed River Basin) RIVER BASINS. Akarçay (Gönülol & Obalı 1986; (Fig. 1). Şen et al. 1994), Antalya (Conk & Cirik 1991; Morkoy- unlu & Ertan 1995a, b; Morkoyunlu et al. 1996; Ertan RIVER BASINS. Antalya (Conk & Cirik 1991; Morko- et al. 1996; Yüce et al. 1997; Savaş & Cirik 1997; yunlu & Ertan 1995a, b; Ertan et al. 1996; Morkoy- Ertan & Morkoyunlu 1998), Aras (Altuner 1988; Ak- unlu et al. 1996; Yüce et al. 1997; Savaş & Cirik 1997; bulut & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak Ertan & Morkoyunlu 1998), Aras (Altuner 1988; Ak- & Gürbüz 2005), Çoruh (Altuner 1984; Altuner bulut & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; & Gürbüz 2005), Asi (Şen et al. 1997), Büyük Menderes Baykal et al. 2004), East Black Sea River Basin (Şahin (Skuja 1937; Güner 1966), Çoruh (Altuner 1984; Altuner 1992, 1994, 1998a, b, 2000, 2001, 2002, 2003; Şahin & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; & Gönülol 1997; Kolaylı et al. 1998; Kara & Şahin Baykal et al. 2004), East Black Sea River Basin (Şahin 2001), Fırat (Altuner & Gürbüz 1988, 1989, 1990a, b, 1992, 1994, 1998a, b, 2000, 2001, 2002, 2003; Şahin 1991, 1994, 1996; Altuner & Pabucçu 1993, 1994; Şen C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 383 et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; 1993; Gönülol & Obalı 1998), Konya Closed River Gürbüz & Altuner 2000), Gediz (Cirik 1994; Gezerler Basin (Yıldız 1984a, b, 1985a, b, 1986a, b, 1987b), et al. 1996, 1999), Kızılırmak (Gönülol & Çomak 1990, Küçük Menderes (Aysel et al. 1992, 1998a, b, 2001, 1992; Yıldız & Özkıran 1991; Gönülol & Arslan 1992; 2002), Susurluk (Skuja 1937; Gezerler 1992; Dere et al. Gönülol 1993; Dere & Sıvacı 1994; Gönülol & Obalı 2002; Karacaören et al. 2004). 1998), Konya Closed River Basin (Yıldız 1984a, b, 1985a, b, 1986a, b, 1987b; Cirik et al. 1991; Demirsoy GENERAL DISTRIBUTION AND ECOLOGY. Ty- 1996; Akköz & Obalı 1998), Küçük Menderes (Aysel choplanktonic, mesosaprobous and eutraphentic et al. 1992, 1998a, b, 2001, 2002), Marmara (Albay species (Denys 1991; Van Dam et al. 1994), clas- & Aykulu 1994a, b; Temel 1994, 2001, 1996, 1999, sifi ed as an alkaliphilous diatom (Håkansson 1993; 2002, 2003), Sakarya (Obalı 1984; Yıldız 1987a; Obalı Van Dam et al. 1994). et al. 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu et al. 1999; Atıcı 2002; Yardımcı & Temel 2000), 9. Craticula cuspidata (Kützing) D. G. Mann Susurluk (Skuja 1937; Dere et al. 2002; Karacaören Figs 19 & 133–135 et al. 2004), West Black Sea River Basin (Öztürk 1994; Cirik 1997), Yeşilırmak (Altuner & Pabuçcu 1996). Navicula cuspidata Kützing

GENERAL DISTRIBUTION & ECOLOGY. Tycho- Ref. Hustedt 1930 (p. 268, Fig. 433); Patrick & Reimer planktonic, mesosaprobous and eutraphentic spe- 1966 (p. 464, Fig. 43: 9–10); Krammer & Lange-Ber- talot 1986 (p. 126, Fig. 43: 1–8); Lange-Bertalot 2001 cies (Denys 1991; Van Dam et al. 1994), classifi ed (p. 111, Figs 82: 1, 2; 83: 1, 2); Hofmann et al. 2011 as an alkaliphilous diatom (Håkansson 1993; Van (p. 139, Fig. 44: 1, 2). Dam et al. 1994). DIMENSIONS. Valves 85–100 μm long and 8. Cocconeis placentula var. lineata (Ehrenberg) 20–22 μm wide, 14–17 striae in 10 μm. Van Heurck Figs 18, 131 & 132 DISTRIBUTION IN TURKMEN MT. Dümbüldek, Cocconeis placentula var. lineata (Ehrenberg) Cleve, Darıpınar, İdris. C. placentula var. lineata (Ehrenberg) Van Heurck DISTRIBUTION IN TURKEY. Lakes and rivers in Ref. Hustedt 1930 (p. 190, Fig. 262); Patrick & Reimer most of the river basins (Fig. 1). 1966 (p. 241, Fig. 15: 8); Krammer & Lange-Bertalot 1991b (p. 87, Fig. 52: 1–13). RIVER BASINS. Antalya (Conk & Cirik 1991; Morko- yunlu & Ertan 1995a, b; Morkoyunlu et al. 1996; Ertan DIMENSIONS. Valves 21.0–29.3 μm long and et al. 1996; Yüce et al. 1997; Savaş & Cirik 1997; 12.0–15.3 μm wide, 19–21 striae in 10 μm. Ertan & Morkoyunlu 1998), Aras (Altuner 1988; Ak- DISTRIBUTION IN TURKMEN MT. Türkmen- bulut & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak baba. & Gürbüz 2005), Asi (Şen et al. 1997), Büyük Men- deres (Skuja 1937; Güner 1966), Çoruh (Altuner DISTRIBUTION IN TURKEY. Lakes and rivers in 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; Eastern Anatolia (Fırat, Aras, Çoruh) and Egean Şen et al. 1999; Baykal et al. 2004), East Black Sea (Gediz, Küçük Menderes, Susurluk) river basins River Basin (Şahin 2002), Fırat (Şen & Çetin 1988; (Fig. 1). Altuner & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, 1996; Altuner & Pabucçu 1993, 1994; Şen et al. 1995; RIVER BASINS. Antalya (Skuja 1937; Vardar & Güner Çetin & Şen 1998, 2004; Gürbüz 2000; Gürbüz & Al- 1972), Aras (Gürbüz et al. 2002a, b; Kıvrak & Gürbüz tuner 2000), Gediz (Güner 1970; Cirik 1994; Gezerler 2005), Çoruh (Altuner 1984; Altuner & Aykulu 1987), et al. 1996, 1999), Kızılırmak (Özesm 1987; Gönülol Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. 2004), & Çomak 1990, 1992; Yıldız & Özkıran 1991; Gönülol East Black Sea River Basin (Şahin 2002), Fırat (Altuner & Arslan 1992; Gönülol 1993; Dere & Sıvacı 1994; & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, 1996; Gönülol & Obalı 1998; Akbulut 2003), Konya Closed Altuner & Pabucçu 1993, 1994; Şen et al. 1995; Çetin River Basin (Skuja 1937; Yıldız 1984a, b, 1985a, b, & Şen 1998, 2004; Gürbüz 2000; Gürbüz & Altuner 1986a, b, 1987b; Cirik et al. 1991; Temel 1992, 1996; 2000), Gediz (Gezerler et al. 1996), Kızılırmak (Gönülol Yıldız et al. 1994; Demirsoy 1996; Akköz & Obalı & Çomak 1990, 1992; Elmacı & Obalı 1992; Gönülol 1998; Morkoyunlu et al. 1999; Mitamura et al. 1997; 384 POLISH BOTANICAL JOURNAL 57(2). 2012

Elmacı & Obalı 1998), Küçük Menderes (Aysel et al. Gürbüz & Altuner 2000), Gediz (Gezerler et al. 1996), 1992, 1998a, b, 2001, 2002; Balık & Gezerler 1995), Konya Closed River Basin (Skuja 1937; Yıldız 1984a, b, Marmara (Albay & Aykulu 1994a, b; Temel 1994, 1996, 1985a, b, 1986a, b, 1987b; Cirik et al. 1991; Temel 1992, 1999, 2002, 2003), Sakarya (Gönülol & Aykulu 1984; 1996; Yıldız et al. 1994; Demirsoy 1996; Akköz & Obalı Obalı 1984; Ünal 1984; Gönülol 1985a, b, 1987; Yıldız 1998), Küçük Menderes (Aysel et al. 1992, 1998a, b, 1987a; Dere 1989; Obalı et al. 1989; Yıldız & Özkıran 2001, 2002; Balık & Gezerler 1995), Marmara (Albay 1994; Atıcı & Yıldız 1996; Aykulu et al. 1999; Yardımcı & Aykulu 1994a, b; Temel 1994, 1996, 1999, 2002, & Temel 2000; Atıcı 2002), Seyhan (Kandemir-Çevik 2003), Sakarya (Gönülol & Aykulu 1984; Obalı 1984; et al. 1994), Susurluk (Skuja 1937; Gezerler 1992; Dere Ünal 1984; Gönülol 1985a, b, 1987; Yıldız 1987a; Dere et al. 2002; Karacaören et al. 2004). 1989; Obalı et al. 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu et al. 1999), Seyhan (Kandemir- GENERAL DISTRIBUTION & ECOLOGY. Wide- Çevik et al. 1994), Susurluk (Skuja 1937; Dere et al. spread, tychoplanktonic, mesosaprobous and eut- 2002; Karacaören et al. 2004), West Black Sea River raphentic species (Lange-Bertalot 1979; Krammer Basin (Öztürk 1994), Kızılırmak (Özesm 1987; Kılınç & Lange-Bertalot 1986; Denys 1991; Hofmann & Dere 1988; Gönülol & Çomak 1990, 1992; Yıldız 1994; Van Dam et al. 1994; Lange-Bertalot 2001), & Özkıran 1991; Gönülol & Arslan 1992; Gönülol 1993; classifi ed as an alkaliphilous diatom (Håkansson Dere & Sıvacı 1994; Gönülol & Obalı 1998; Kılınç 1993; Van Dam et al. 1994). 1998; Kılınç & Sıvacı 2001; İşbakan et al. 2002; Ak- bulut 2003), Yeşilırmak (Yazıcı & Gönülol 1994; Altuner & Pabuçcu 1996; Kır & Polat 1996). 10. Cyclotella ocellata Pantocsek Figs 20, 21 & 142 GENERAL DISTRIBUTION AND ECOLOGY. Eu- planktonic, oligosaprobous and meso-eutraphentic Ref. Hustedt 1930 (p. 100, Fig. 67); Krammer & Lange- Bertalot 1991a (p. 51, Figs 50: 1–11, 13, 14; 51: 1–5); species (Denys 1991; Van Dam et al. 1994), clas- Håkansson 2002 (p. 85, Figs 309–318). sifi ed as an alkaliphilous diatom (Håkansson 1993; Van Dam et al. 1994). DIMENSIONS. Valves 9–23 μm in diameter and 20–22 striae in 10 μm. 11. Cymatopleura elliptica (Brébisson) W. Smith DISTRIBUTION IN TURKMEN MT. Dümbüldek, Fig. 23 Darıpınar, Söğüt-Lütfi ye, Türkmenbaba, Uluköy. Ref. Hustedt 1930 (p. 426, Fig. 825); Krammer & Lange- Bertalot 1988 (p. 170, Figs 119: 1–4; 120: 1–6; 122: 3); DISTRIBUTION IN TURKEY. Lakes and rivers in Hofmann et al. 2011 (p. 143, Fig. 123: 1–4). Eastern Anatolia (Fırat, Çoruh and Aras river ba- sins), Egean (Küçük Menderes River Basin), Medi- DIMENSIONS. Valves 65–89 μm long and μm terrnean (Antalya and Asi river basins) and Inner 42–52 μm wide, 3–4 fi bulae in 10 μm. Anatolia (Konya Closed River Basin) (Fig. 1). DISTRIBUTION IN TURKMEN MT. Söğüt-Lütfi ye, RIVER BASINS. Akarçay (Aykulu & Obalı 1981; Güllüdere-Türkmenbaba. Gönülol & Obalı 1986; Şen et al. 1994), Antalya (Conk DISTRIBUTION IN TURKEY. Lakes and rivers in & Cirik 1991; Morkoyunlu & Ertan 1995a, b; Ertan all river basins (Fig. 1). et al. 1996; Morkoyunlu et al. 1996; Savaş & Cirik 1997; Yüce et al. 1997; Mitamura et al. 1997; Elmacı RIVER BASINS. Akarçay (Aykulu & Obalı 1981), An- & Obalı 1998; Ertan & Morkoyunlu 1998; Morkoy- talya (Conk & Cirik 1991; Morkoyunlu & Ertan 1995a, unlu et al. 1999), Aras (Altuner 1988; Akbulut & Yıldız b; Ertan et al. 1996; Morkoyunlu et al. 1996; Savaş 2002; Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), & Cirik 1997; Yüce et al. 1997; Ertan & Morkoyunlu Asi (Şen et al. 1997), Çoruh (Altuner 1984; Altuner 1998), Aras (Altuner 1988; Akbulut & Yıldız 2002; & Aykulu 1987; Atıcı 2002; Yardımcı & Temel 2000), Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Çoruh Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. 2004), (Altuner 1984; Altuner & Aykulu 1987), Dicle (Şen East Black Sea River Basin (Şahin 2002), Fırat (Altuner 1988a; Şen et al. 1999; Baykal et al. 2004), East Black & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, 1996; Sea River Basin (Şahin 2002), Fırat (Altuner & Pabucçu Şen & Çetin 1988; Altuner & Pabucçu 1993, 1994; Şen 1993, 1994; Şen et al. 1995; Çetin & Şen 1998, 2004; et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Gürbüz 2000; Gürbüz & Altuner 2000), Gediz (Cirik C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 385

Figs 22–30. 22 – Cymatopleura solea (Brébisson) W. Smith, 23 – C. elliptica (Brébisson) W. Smith, 24, 25 – Cymbella affi niformis Krammer, 26 – C. cymbiformis Agardh, 27 – C. helvetica Kützing, 28 – C. neocistula Krammer, 29 & 30 – C. subleptoceros Krammer. Scale bar = 10 μm. 386 POLISH BOTANICAL JOURNAL 57(2). 2012

1994; Gezerler et al. 1999), Kızılırmak (Özesm 1987; (Altuner & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, Kılınç & Dere 1988; Yıldız & Özkıran 1991; Gönülol 1996; Altuner & Pabucçu 1993, 1994; Şen & Çetin 1988; & Arslan 1992; Dere & Sıvacı 1994; Kılınç 1998; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Kılınç & Sıvacı 2001; Akbulut 2003), Konya Closed Gürbüz & Altuner 2000), Gediz (Cirik 1994; Gezerler River Basin (Skuja 1937; Yıldız 1984a, b, 1985a, b, et al. 1996, 1999), Kızılırmak (Özesm 1987; Kılınç 1986a, b, 1987b; Cirik et al. 1991; Temel 1992, 1996; & Dere 1988; Gönülol & Çomak 1990, 1992; Yıldız Yıldız et al. 1994; Demirsoy 1996; Akköz & Obalı 1998; & Özkıran 1991; Gönülol & Arslan 1992; Gönülol Morkoyunlu et al. 1999; Mitamura et al. 1997; Elmacı 1993; Dere & Sıvacı 1994; Gönülol & Obalı 1998; & Obalı 1998), Küçük Menderes (Aysel et al. 1992, Kılınç 1998; Kılınç & Sıvacı 2001; Akbulut 2003), 1998a, b, 2001, 2002), Marmara (Temel 1996, 1999, Konya Closed River Basin (Skuja 1937; Cirik et al. 2002), Sakarya (Gönülol 1985a, 1987; Yıldız 1987a; 1991; Temel 1992, 1996; Yıldız 1984a, b, 1985a, b, Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu 1986a, b, 1987b; Yıldız et al. 1994; Demirsoy 1996; et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), Su- Mitamura et al. 1997; Akköz & Obalı 1998; Elmacı surluk (Skuja 1937; Gezerler 1992; Dere et al. 2002; & Obalı 1998; Morkoyunlu et al. 1999), Küçük Men- Karacaören et al. 2004), West Black Sea River Basin deres (Aysel et al. 1992, 1998a, b, 2001, 2002; Balık (Öztürk 1994), Yeşilırmak (Yazıcı & Gönülol 1994; Al- & Gezerler 1995), Marmara (Albay & Aykulu 1994a, b; tuner & Pabuçcu 1996; Kır & Polat 1996). Temel 1994, 1996, 1999, 2002, 2003), Sakarya (Gönülol & Aykulu 1984; Obalı 1984; Ünal 1984; Gönülol 1985a, GENERAL DISTRIBUTION AND ECOLOGY. Wide- b, 1987; Yıldız 1987a; Dere 1989; Obalı et al. 1989; spread, euplanktonic, mesosaprobous and eutra- Kandemir-Çevik et al. 1994; Yıldız & Özkıran 1994; phentic species (Krammer & Lange-Bertalot 1988; Atıcı & Yıldız 1996; Aykulu et al. 1999; Atıcı 2002; Denys 1991; Hofmann 1994; Van Dam et al. 1994), Yardımcı & Temel 2000), Susurluk (Brunnthaler 1903, classifi ed as an alkaliphilous diatom (Håkansson Skuja 1937; Gezerler 1992; Dere et al. 2002; Karacaören 1993; Van Dam et al. 1994). et al. 2004), West Black Sea River Basin (Öztürk 1994), Yeşilırmak (Yazıcı & Gönülol 1994; Altuner & Pabuçcu 1996; Kır & Polat 1996). 12. Cymatopleura solea (Brébisson) W. Smith Fig. 22 GENERAL DISTRIBUTION AND ECOLOGY. Wide- spread, euplanktonic, mesosaprobous and eutra- Ref. Hustedt 1930 (p. 425, Fig. 823a); Krammer phentic species (Krammer & Lange-Bertalot 1988; & Lange-Bertalot 1988 (p. 168, Figs 117: 1–5; 118: 1, 3); Hofmann et al. 2011 (p. 144, Figs 124–125). Lange-Bertalot 1979; Denys 1991; Hofmann 1994; Van Dam et al. 1994), classifi ed as an alkaliphilous DIMENSIONS. Valves 65–131 μm long and (Håkansson 1993; Van Dam et al. 1994). 13–23 μm wide, 7–8 fi bulae in 10 μm. 13. Cymbella affi niformis Krammer DISTRIBUTION IN TURKMEN MT. Söğüt-Lütfi ye, Figs 24, 25 & 136–141 Güllüdere. Ref. Krammer 2002 (p. 45, Fig. 25; 1–8). DISTRIBUTION IN TURKEY. Lakes and rivers in most of the river basins (Fig. 1). DIMENSIONS. Valves 23.0–38.7 μm long and 6–10 μm wide, 10–12 striae in 10 μm. RIVER BASINS. Akarçay (Aykulu & Obalı 1981; Gönülol & Obalı 1986; Şen et al. 1994), Antalya (Conk DISTRIBUTION IN TURKMEN MT. Darıpınar, & Cirik 1991; Ertan et al. 1996; Yüce et al. 1997; Dümbüldek, Çobanlar, Türkmenbaba. Savaş & Cirik 1997; Morkoyunlu & Ertan 1995a, b; Morkoyunlu et al. 1996; Ertan & Morkoyunlu 1998), DISTRIBUTION IN TURKEY. New for fl ora of Aras (Altuner 1988; Akbulut & Yıldız 2002; Gürbüz Turkey. et al. 2002a, b; Kıvrak & Gürbüz 2005), Asi (Şen GENERAL DISTRIBUTION AND ECOLOGY. Cym- et al. 1997), Çoruh (Altuner 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. bella affi niformis is probably widespread, thuogh 2004), East Black Sea River Basin (Şahin 1992, 1994, not distinguished from commonly reported taxa. 1998a, b, 2000, 2001, 2002, 2003; Şahin & Gönülol The species ecology and distribution remain very 1997; Kolaylı et al. 1998; Kara & Şahin 2001), Fırat poorly known. It has been described in 2002 by C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 387

Krammer from material collected from pond (Ger- Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Demirsoy many). Later, very similar diatom – Cymbella yakii 1996; Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı Jüttner & Van de Vijver (2010) has been described 2002), Seyhan (Kandemir-Çevik et al. 1994), Susurluk from Asia (Nepal). (Skuja 1937; Dere et al. 2002; Karacaoğlu et al. 2004), West Black Sea River Basin (Öztürk 1994), Yeşilırmak (Yazıcı & Gönülol 1994; Altuner & Pabucçu 1996; Kır 14. Cymbella cymbiformis Agardh Fig. 26 & Polat 1996; Gönülol & Obalı 1998). Ref. Hustedt 1930 (p. 362, Fig. 672); Patrick & Reimer GENERAL DISTRIBUTION AND ECOLOGY. Wide- 1975 (p. 54, Fig. 10: 3–4); Krammer & Lange-Bertalot spread, euplanktonic, oligosaprobous, oligo-meso- 1986 (p. 317, Fig. 129: 2–6); Krammer 2002 (p. 76, Fig. 58; 59; 61; 62: 1–3; 63; 65); Hofmann et al. 2011 traphentic species (Denys 1991; Van Dam et al. (p. 148, Fig. 80: 1–11). 1994; Krammer 2002), classifi ed as a circumneutal or alkaliphilous (Håkansson 1993; Van Dam et al. DIMENSIONS. Valves 44–83 μm long and 1994). 13–16 μm wide, 9–11 striae in 10 μm.

DISTRIBUTION IN TURKMEN MT. Türkmenbaba, 15. Cymbella helvetica Kützing Fig. 27 Darıpınar, İdrıs, Söğüt-Lütfi ye. Ref. Hustedt 1930 (p. 364, Fig. 678); Krammer & Lange-Bertalot 1986 (p. 324, Figs 132: 2–4; 133: DISTRIBUTION IN TURKEY. Lakes and rivers in 1–8); Krammer 2002 (p. 154, Figs 183: 1–3; 184: 1–4; most of the river basins (Fig. 1). 185: 1–6; 187: 1–5); Hofmann et al. 2011 (p. 151, Fig. 81: 3–4). RIVER BASINS. Antalya (Conk & Cirik 1991; Mor- koyunlu & Ertan 1995a, b; Ertan et al. 1996; Morko- DIMENSIONS. Valves 40–120 μm long and yunlu et al. 1996; Savaş & Cirik 1997; Yüce et al. 10–21 μm wide, maximal length/breadth ratio 1997; Ertan & Morkoyunlu 1998), Aras (Altuner 1988; about 6.6 and 10–11 striae in 10 μm. Akbulut & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Asi (Şen et al. 1997), Büyük Menderes DISTRIBUTION IN TURKMEN MT. Güllüdere. (Güner 1966; Skuja 1937), Çoruh (Altuner 1984; Altuner & Aykulu 1987), Dicle (Sen 1988b; Şen & Aksakal DISTRIBUTION IN TURKEY. Lakes and rivers 1988; Şen et al. 1999; Baykal et al. 2004), East Black in Eastern Anatolia (Fırat, Aras, Çoruh), Egean Sea River Basin (Şahin 1992, 1993, 1994, 1998a, b, (Küçük Menderes, Gediz, Susurluk), Black Sea 2000, 2001, 2002, 2003; Şahin & Gönülol 1997; Ko- (East Black Sea River Basin, Yeşilırmak), Inner laylı et al. 1998; Kara & Şahin 2001), Fırat (Altuner Anatolia (Konya Closed River Basin, Sakarya, & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, 1996; Kızılırmak) river basins (Fig. 1). Altuner & Pabucçu 1993, 1994; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Gürbüz & Altuner RIVER BASINS. Akarçay (Gönülol & Obalı 1986; Şen 2000); Gediz (Güner 1970; Cirik 1994; Gezerler et al. et al. 1994), Aras (Altuner 1988; Akbulut & Yıldız 2002; 1996, 1999), Kızılırmak (Kılınç & Dere 1988; Gönülol Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Asi (Şen & Çomak 1990, 1992; Yıldız & Özkıran 1991; Gönülol et al. 1997), Çoruh (Altuner 1984; Altuner & Aykulu & Arslan 1992; Gönülol 1993; Dere & Sıvacı 1994; 1987), Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. Kılınç 1998; Kılınç & Sıvacı 2001; İşbakan et al. 2002), 2004), East Black Sea River Basin (Şahin 1992, 1994, Konya Closed River Basin (Skuja 1937; Yıldız 1984a, b, 1998a, b, 2000, 2001, 2002, 2003; Şahin & Gönülol 1985a, b, 1986a, b, 1987b Cirik et al. 1991; Temel 1992, 1997; Kolaylı et al. 1998; Kara & Şahin 2001), Fırat 1996; Yıldız et al. 1994; Demirsoy 1996; Morkoyunlu (Altuner & Pabucçu 1993, 1994; Şen et al. 1995; Gürbüz et al. 1996; Mitamura et al. 1997; Akköz & Obalı 1998; 2000; Gürbüz & Altuner 2000; Çetin & Şen 1998, Elmacı & Obalı 1998), Küçük Menderes (Skuja 1937; 2004), Gediz (Gezerler et al. 1996), Kızılırmak (Yıldız Cirik & Cirik 1989; Aysel et al. 1992, 1998a, b, 2001, & Özkıran 1991; Dere & Sıvacı 1994), Konya Closed 2002; Balık & Gezerler 1995), Marmara (Albay & Ay- River Basin (Skuja 1937; Cirik et al. 1991; Temel 1992, kulu 1994a, b; Yardımcı & Temel 2000; Temel 1994, 1996; Yıldız et al. 1994; Demirsoy 1996; Mitamura 1996, 1999, 2002, 2003), Sakarya (Aykulu et al. 1983; et al. 1997; Akköz & Obalı 1998; Elmacı & Obalı 1998; Gönülol & Aykulu 1984; Obalı 1984; Ünal 1984; Gö- Morkoyunlu et al. 1999), Küçük Menderes (Aysel et al. nülol 1985b; Yıldız 1987a; Dere 1989; Obalı et al. 1989; 1992, 1998a, b, 2001, 2002), Marmara (Albay & Aykulu 388 POLISH BOTANICAL JOURNAL 57(2). 2012

1994a, b; Temel 1994, 1996, 1999, 2002, 2003), Sakarya 2000), Gediz (Cirik 1994; Temel 1994, 1996, 1999, (Gönülol 1985a, 1987; Yıldız 1987a; Yıldız & Özkıran 2002, 2003; Gezerler et al. 1996, 1999); Kızılırmak 1994; Atıcı & Yıldız 1996; Aykulu et al. 1999; Yardımcı (Kılınç & Dere 1988; Gönülol & Çomak 1990, 1992; & Temel 2000; Atıcı 2002), Seyhan (Kandemir-Çevik Yıldız & Özkıran 1991; Gönülol 1993; Dere & Sıvacı et al. 1994). 1994; Kılınç 1998; Kılınç & Sıvacı 2001), Konya Closed River Basin (Skuja 1937; Cirik et al. 1991; Temel 1992, GENERAL DISTRIBUTION AND ECOLOGY. Eu- 1996; Yıldız et al. 1994; Demirsoy 1996; Morkoyunlu planktonic, oligosaprobous, oligo-mesotraphentic et al. 1996; Mitamura et al. 1997; Akköz & Obalı 1998; or mesotraphentic species (Denys 1991; Hofmann Elmacı & Obalı 1998), Küçük Menderes (Skuja 1937; 1994; Van Dam et al. 1994; Krammer 2002), clas- Aysel et al. 1992, 1998a, b, 2001, 2002; Balık & Gez- sifi ed as an alkaliphilous diatom (Håkansson 1993; erler 1995), Marmara (Temel 1996, 1999, 2002), Sakarya Van Dam et al. 1994). (Obalı 1984; Yıldız 1987a; Obalı et al. 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), Susurluk (Skuja 16. Cymbella neocistula Krammer Fig. 28 1937; Gezerler 1992; Dere et al. 2002; Karacaoğlu et al. 2004), Yeşilırmak (Gönülol & Obalı 1998). Cymbella cistula (Hemprich) Grunow, C. cistula (Ehren- berg) Kirchner GENERAL DISTRIBUTION AND ECOLOGY. Wide- Ref. Hustedt 1930 (p. 363, Fig. 676a); Patrick & Reimer spread, tychoplanktonic, mesosaprobous and eu- 1975 (p. 62, Fig. 11: 3–4); Krammer & Lange-Bertalot traphentic species (Denys 1991; Hofmann 1994; 1986 (p. 316, Figs 127: 8–11; 128: 1–6); Krammer 2002 Van Dam et al. 1994; Krammer 2002), classifi ed (p. 94, Figs 85: 1–4; 86: 1–7; 87: 1–9; 88: 1–8); Hof- as an alkaliphilous diatom (Håkansson 1993; Van mann et al. 2011 (p. 155, Fig. 79: 11–15). Dam et al. 1994). DIMENSIONS. Valves 32–82 μm long and 12.0– 15.7 μm wide, maximal length/breadth ratio about 17. Cymbella subleptoceros Krammer 6.1 and 9–11 striae in 10 μm. Figs 29 & 30 Ref. Krammer 2002 (p. 133, Figs 154: 2–17; 155: 1–7). DISTRIBUTION IN TURKMEN MT. Gölcük, Darıpınar. DIMENSIONS. Valves 14.0–40.7 μm long, 6–11 μm wide and 9–13 striae in 10 μm. DISTRIBUTION IN TURKEY. Lakes and rivers in Mediterranean (Antalya River Basin), Black DISTRIBUTION IN TURKMEN MT. Darıpınar. Sea (East Black Sea River Basin and Kızılırmak), DISTRIBUTION IN TURKEY. New for fl ora of Egean (Gediz, Küçük Menderes), Inner Anatolia Turkey. (Konya Closed River Basin, Sakarya), Eastern Anatolia (Fırat) river basins (Fig. 1). GENERAL DISTRIBUTION AND ECOLOGY. Tycho- planktonic, oligo- or mesosaprobous and oligotra- RIVER BASINS. Akarçay (Gönülol & Obalı 1986; Şen et al. 1994), Antalya (Conk & Cirik 1991; Morkoy- phentic species (Denys 1991; Hofmann 1994; Van unlu & Ertan 1995a, b; Ertan et al. 1996; Morkoyunlu Dam et al. 1994), classifi ed as alkaliphilous (Van et al. 1996; Savaş & Cirik 1997; Yüce et al. 1997; Dam et al. 1994). Ertan & Morkoyunlu 1998), Aras (Altuner 1988; Ak- bulut & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak 18. Cymbopleura amphicephala (Naegeli) & Gürbüz 2005), Büyük Menderes (Skuja 1937; Güner Krammer Figs 31 & 32 1966), Çoruh (Altuner 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. 2004), SYNONYMS. Cymbella amphicephala Naegeli, C. amphi- East Black Sea River Basin (Şahin 1992, 1994, 1998a, cephala Naegeli ex Kützing b, 2000, 2001, 2002, 2003; Şahin & Gönülol 1997; Ref. Hustedt 1930 (p. 355, Fig. 651); Patrick & Reimer Kolaylı et al. 1998; Kara & Şahin 2001), Fırat (Altuner 1975 (p. 33, Fig. 4: 11); Krammer & Lange-Bertalot & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, 1996; 1986 (p. 335, Fig. 142: 3–14, 21); Krammer 2003 (p. 70, Altuner & Pabucçu 1993, 1994; Şen et al. 1995; Çetin Figs 91: 1–18; 93: 2–8); Hofmann et al. 2011 (p. 160, & Şen 1998, 2004; Gürbüz 2000; Gürbüz & Altuner Fig. 83: 11–15). C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 389

Figs 31–54. 31 & 32 – Cymbopleura amphicephala (Naegeli) Krammer, 33–35 – Denticula tenuis Kützing, 36 & 37 – Diatoma mesodon (Ehrenberg) Kützing, 38–40 – D. tenuis Agardh, 41 – D. vulgaris Bory, 42–44 – Epithemia sorex Kützing, 45 – Fragilaria capucina Desmaziéres, 46–48 – F. vaucheriae (Kützing) Petersen, 49 & 50 – Gomphonema olivaceum (Hornemann) Brébisson, 51, 52 – G. parvulum (Kützing) Kützing, 53, 54 – Hantzschia amphioxys (Ehrenberg) Grunow. Scale bar = 10 μm. 390 POLISH BOTANICAL JOURNAL 57(2). 2012

DIMENSIONS. Valves 23–36 μm long and DISTRIBUTION IN TURKMEN MT. Dümbüldek, 7–10 μm wide, maximal length/breadth ratio about Seydi. 4.4 and 11–15 striae in 10 μm. DISTRIBUTION IN TURKEY. Lakes and rivers in DISTRIBUTION IN TURKMEN MT. Dümbüldek, Egean (Gediz, Küçük Menderes) and Black Sea Darıpınar, Güllüdere, Türkmenbaba, Akındere. (Yeşilırmak, Kızılırmak) river basins (Fig. 1).

DISTRIBUTION IN TURKEY. Lakes and rivers RIVER BASINS. Akarçay (Gönülol & Obalı 1986; in Mediterranean (Antalya River Basin), Eastern Şen et al. 1994), Fırat (Altuner & Gürbüz 1988, 1989, Anatolia (Fırat, Aras, Çoruh river basins), Inner 1990a, 1991), Gediz (Gezerler et al. 1996), Kızılırmak (Özesm 1987; Gönülol & Çomak 1990, 1992; Gönülol Anatolia (Konya Closed River Basin, Sakarya, 1993; Akbulut 2003), Küçük Menderes (Aysel et al. Kızılırmak river basins) (Fig. 1). 1992, 1998a, b, 2001, 2002), Sakarya (Obalı 1984; Obalı RIVER BASINS. Antalya (Conk & Cirik 1991; et al. 1989), Yeşilırmak (Gönülol & Obalı, 1998). Morkoyunlu & Ertan 1995a, b; Ertan et al. 1996; GENERAL DISTRIBUTION AND ECOLOGY. Wide- Morkoyunlu et al. 1996; Savaş & Cirik 1997; Yüce spread, tychoplanktonic, oligo- or mesosaprobous et al. 1997; Ertan & Morkoyunlu 1998), Aras (Altuner 1988; Akbulut & Yıldız 2002; Gürbüz et al. 2002a, b; and mesotraphentic species (Krammer & Lange- Kıvrak & Gürbüz 2005), Çoruh (Altuner 1984; Altuner Bertalot 1988; Denys 1991; Hofmann 1994; Van & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; Dam et al. 1994), classifi ed as a circumneutral Baykal et al. 2004), East Black Sea River Basin (Şahin (Håkansson 1993) or alkaliphilous diatom (Van 1992, 1994, 1998a, b, 2000, 2001, 2002, 2003; Şahin Dam et al. 1994). & Gönülol 1997; Kolaylı et al. 1998; Kara & Şahin 2001), Fırat (Altuner & Gürbüz 1988, 1989, 1990a, b, 20. Diatoma mesodon (Ehrenberg) Kützing 1991, 1994, 1996; Şen & Çetin 1988; Altuner & Pabucçu Figs 36 & 37 1993, 1994; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Gürbüz & Altuner 2000), Gediz (Gez- Diatoma hiemale var. mesodon (Ehrenberg) Grunow erler et al. 1996), Kızılırmak (Yıldız & Özkıran 1991; Ref. Hustedt 1930 (p. 129, Fig. 116); Krammer & Lange- Dere & Sıvacı 1994), Konya Closed River Basin Bertalot 1991a (p. 100, Figs 98: 7; 99: 1–12); Hofmann (Yıldız 1984a, b,1985a, b,1986a, b,1987b; Cirik et al. et al. 2011 (p. 173, Fig. 2: 1–5). 1991; Demirsoy 1996; Akköz & Obalı 1998), Küçük Menderes (Aysel et al. 1992, 1998a, b, 2001, 2002), DIMENSIONS. Valves 13–19 μm long and 6.5– Sakarya (Gönülol & Aykulu 1984; Obalı 1984; Ünal 9.0 μm wide, 3–4 transapical bars and 22–35 striae 1984; Gönülol 1985a, b, 1987; Yıldız 1987a; Dere 1989; in 10 μm. Obalı et al. 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız DISTRIBUTION IN TURKMEN MT. Uluköy, İnli, 1996; Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), Seyhan (Kandemir-Çevik et al. 1994). Söğüt-Lütfi ye, Türkmenbaba. DISTRIBUTION IN TURKEY. Egean (Küçük Men- GENERAL DISTRIBUTION AND ECOLOGY. Wide- spread, tychoplanktonic, oligo- or oligo-meso- deres, Gediz, Susurluk) river basins (Fig. 1). saprobous species (Denys 1991; Hofmann 1994; RIVER BASINS. Aras (Altuner 1988; Akbulut & Yıldız Van Dam et al. 1994; Krammer 2003), classifi ed 2002; Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), as a circumneutral diatom (Håkansson 1993; Van Çoruh (Altuner 1984; Altuner & Aykulu 1987; Altuner Dam et al. 1994). & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, 1996), East Black Sea River Basin (Şahin 2002), Fırat (Altuner & Pabucçu 1993, 1994; Gürbüz 2000; Gürbüz & Altuner 19. Denticula tenuis Kützing Figs 33–35 & 143 2000), Gediz (Gezerler et al. 1996), Küçük Menderes Ref. Hustedt 1930 (p. 381, Fig. 723); Krammer & Lange- (Aysel et al. 1992, 1998a, b, 2001, 2002), Susurluk (Skuja Bertalot 1988 (p. 139, Fig. 95: 4–25); Hofmann et al. 1937; Dere et al. 2002, Karacaoğlu et al. 2004). 2011 (p. 169, Fig. 117: 32–37). GENERAL DISTRIBUTION AND ECOLOGY. DIMENSIONS. Valves 11.5–18.0 μm long and Widespread, oligo- or oligo-mesosaprobous and 3.0–5.3 μm wide, 5–7 fi bulae in 10 μm. oligo- or oligo-mesotraphentic species (Krammer C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 391

& Lange-Bertalot 1991a; Hofmann 1994; Van Hofmann 1994; Van Dam et al. 1994), classifi ed as Dam et al. 1994), classifi ed as a circumneutral an alkaliphilous diatom (Van Dam et al. 1994). (Van Dam et al. 1994) or an alkaliphilous diatom (Håkansson 1993). 22. Diatoma vulgaris Bory Figs 41 & 144–147 Ref. Hustedt 1930 (p. 127, Fig. 103); Krammer & Lange- 21. Diatoma tenuis Agardh Figs 38–40 Bertalot 1991a (p. 95, Fig. 93: 1–12; 94: 1–13; 97: 3–5); Diatoma elongatum var. tenuis Agardh, D. tenue var. Hofmann et al. 2011 (p. 175, Fig. 3: 20–25). elongatum Lyngbye DIMENSIONS. Valves 26–40 μm long and 5.0– Ref. Hustedt 1930 (p. 127, Fig. 111); Patrick & Reimer 13.3 μm wide, 6–10 transapical bars and over 40 1966 (p. 109, Fig. 2: 6); Krammer & Lange-Bertalot striae in 10 μm. 1991a (p. 97, Fig. 96: 1–9). DISTRIBUTION IN TURKMEN MT. Dümbüldek, DIMENSIONS. Valves 24–36 μm long and Söğüt-Lütfi ye. 3.0–4.7 μm wide, 8–10 transapical bars in 10 μm. DISTRIBUTION IN TURKEY. Lakes and rivers in DISTRIBUTION IN TURKMEN MT. Dümbüldek, Egean (Küçük Menderes, Susurluk, Gediz), Med- Darıpınar, Uluköy. iterranean (Antalya), Inner Anatolian (Sakarya, Konya Closed River Basin) and Black Sea (East DISTRIBUTION IN TURKEY. Lakes and riv- Black Sea River Basin, West Black Sea River Basin, ers in Mediterranean (Antalya), Eastern Anato- Kızılırmak, Yeşilırmak) river basins (Fig. 1). lia (Fırat), Inner Anatolia (Konya Closed River Basin, Sakarya), Egean (Küçük Menderes) river RIVER BASINS. Akarçay (Gönülol & Obalı 1986; basins (Fig. 1). Şen et al. 1994), Antalya (Conk & Cirik 1991; Morkoy- unlu & Ertan 1995a, b; Ertan et al. 1996; Morkoyunlu RIVER BASINS. Akarçay (Gönülol & Obalı 1986; et al. 1996; Savaş & Cirik 1997; Yüce et al. 1997; Ertan Şen et al. 1994), Antalya (Conk & Cirik 1991; Morkoy- & Morkoyunlu 1998), Aras (Altuner 1988; Akbulut unlu & Ertan 1995a, b; Ertan et al. 1996; Morkoyunlu & Yıldız 2002), Asi (Şen et al. 1997), Büyük Menderes et al. 1996; Savaş & Cirik 1997; Yüce et al. 1997; Ertan (Cirik et al. 1989), Dicle (Şen 1988a; Şen et al. 1999; & Morkoyunlu 1998), Aras (Altuner 1988; Akbulut Baykal et al. 2004), East Black Sea River Basin (Şahin & Yıldız 2002), Asi (Şen et al. 1997), Büyük Mend- 1992, 1994, 1998a, b, 2000, 2001, 2003; Şahin & Gönülol eres (Skuja 1937; Güner 1966), Dicle (Şen 1988a; Şen 1997; Kolaylı et al. 1998; Kara & Şahin 2001), Fırat et al. 1999; Baykal et al. 2004), Fırat (Altuner & Gürbüz (Altuner & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, 1988, 1989, 1990a, b, 1991, 1994, 1996; Şen et al. 1995; 1996; Şen et al. 1995; Çetin & Şen 1998, 2004), Gediz Çetin & Şen 1998, 2004), Gediz (Gezerler et al. 1996), (Gezerler et al. 1996), Kızılırmak (Gönülol & Çomak Kızılırmak (Kılınç & Dere 1988; Yıldız & Özkıran 1991; 1990, 1992; Yıldız & Özkıran 1991; Gönülol & Arslan Dere & Sıvacı 1994; Kılınç 1998; Kılınç & Sıvacı 2001), 1992; Gönülol 1993; Dere & Sıvacı 1994; Gönülol Konya Closed River Basin (Skuja 1937; Cirik et al. 1991; & Obalı 1998; İşbakan et al. 2002), Konya Closed River Temel 1992, 1996; Yıldız et al. 1994; Demirsoy 1996; Basin (Skuja 1937; Yıldız 1984a, b, 1985a, b, 1986a, b, Morkoyunlu et al. 1996; Mitamura et al. 1997; Akköz 1987b; Cirik et al. 1991; Temel 1992, 1996; Yıldız et al. & Obalı 1998; Elmacı & Obalı 1998), Küçük Menderes 1994; Demirsoy 1996; Morkoyunlu et al. 1996; Mita- (Skuja 1937; Aysel et al. 1992, 1998a, b, 2001, 2002; Balık mura et al. 1997; Akköz & Obalı 1998; Elmacı & Obalı & Gezerler 1995), Marmara (Albay & Aykulu 1994a, b; 1998), Küçük Menderes (Aysel et al. 1992, 1998a, b, Temel 1994, 1996, 1999, 2002, 2003), Sakarya (Obalı 2001, 2002; Balık & Gezerler 1995), Marmara (Albay 1984; Gönülol 1985a, 1987; Yıldız 1987a; Obalı et al. & Aykulu 1994a, b; Temel 1994, 1996, 1999, 2002, 2003), 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu Sakarya (Aykulu et al. 1983; Gönülol & Aykulu 1984; et al. 1999; Yardımcı & Temel 2000; Atıcı 2002). Obalı 1984; Gönülol 1985a, b, 1987; Yıldız 1987a; Obalı et al. 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; GENERAL DISTRIBUTION AND ECOLOGY. Wide- Demirsoy 1996; Aykulu et al. 1999; Yardımcı & Temel spread, euplanktonic, mesosaprobous and meso- 2000; Atıcı 2002), Seyhan (Kandemir-Çevik et al. 1994), or eutraphentic species (Lange-Bertalot 1979; Susurluk (Skuja 1937; Güner et al. 1987; Gezerler 1992; Denys 1991; Krammer & Lange-Bertalot 1991a; Dere et al. 2002; Karacaören et al. 2004), West Black Sea 392 POLISH BOTANICAL JOURNAL 57(2). 2012

River Basin (Öztürk 1994), Yeşilırmak (Yazıcı & Gönülol GENERAL DISTRIBUTION AND ECOLOGY. Wide- 1994; Altuner & Pabuçcu 1996; Kır & Polat 1996). spread, tychoplanktonic, mesosaprobous and meso- or eutraphentic species (Krammer & Lange- GENERAL DISTRIBUTION AND ECOLOGY. Ty- choplanktonic, mesosaprobous and meso- or eutra- Bertalot 1988; Denys 1991; Hofmann 1994; Van phentic species (Denys 1991; Hofmann 1994; Van Dam et al. 1994), classifi ed as an alkalibiontic Dam et al. 1994), classifi ed as an alkaliphilous- diatom (Håkansson 1993; Van Dam et al. 1994). alkalibiontic (Håkansson 1993) or an alkalibiontic diatom (Van Dam et al. 1994). 24. Fragilaria capucina Desmaziéres Figs 45, 148 & 149 23. Epithemia sorex Kützing Figs 42–44 Ref. Hustedt 1930 (p. 138, Fig. 126); Patrick & Reimer 1966 (p.118, Fig. 3: 5); Krammer & Lange-Bertalot Ref. Hustedt 1930 (p. 388, Fig. 736); Patrick & Reimer 1991a (p. 121, Fig. 108: 1–8); Hofmann et al. 2011 1975 (p. 188, Fig. 27: 4); Krammer & Lange-Bertalot (p. 259, Fig. 9: 8–12). 1988 (p. 154, Fig. 106: 1–13); Hofmann et al. 2011 (p. 206, Fig. 121: 1–7) DIMENSIONS. Valves 22.0–65.3 μm long and 3.5–5.0 μm wide, 14–17 striae in 10 μm. DIMENSIONS. Valves 23.5–47.3 μm long and 6.5–10.0 μm wide, 5–8 fi bulae in 10 μm. DISTRIBUTION IN TURKMEN MT. Darıpınar, Dümbüldek. DISTRIBUTION IN TURKMEN MT. Dümbüldek, Güllüdere, Seydi. DISTRIBUTION IN TURKEY. Lakes and rivers in Egean (Küçük Menderes, Gediz, Susurluk) river DISTRIBUTION IN TURKEY. Lakes and rivers basins (Fig. 1). in Inner Anatolia (Akarçay, Kızılırmak, Sakarya, Konya Closed River Basin), Eastern Anatolia RIVER BASINS. Aras (Altuner 1988; Akbulut & Yıldız (Fırat, Aras, Çoruh), Egean (Gediz, Küçük Men- 2002), Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. deres Susurluk) river basins (Fig. 1). 2004), Fırat (Şen et al. 1995; Çetin & Şen 1998, 2004), Gediz (Cirik 1994; Gezerler et al. 1999), Kızılırmak RIVER BASINS. Akarçay (Aykulu & Obalı 1981; (Özesm 1987; Gönülol & Çomak 1990, 1992; Gönülol Gönülol & Obalı 1986; Şen et al. 1994), Aras (Altuner 1993; Gönülol & Obalı 1998; Akbulut 2003), Küçük 1988; Akbulut & Yıldız 2002; Gürbüz et al. 2002a, b; Menderes (Cirik & Cirik 1989; Aysel et al. 1992, 1998a, Kıvrak & Gürbüz 2005), Çoruh (Altuner 1984; Altuner b, 2001, 2002; Balık & Gezerler 1995), Marmara (Albay & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; & Aykulu 1994a, b; Temel 1994, 2000, 2003), Susurluk Baykal et al. 2004), East Black Sea River Basin (Şahin (Brunnthaler 1903, Skuja 1937; Gezerler 1992; Dere 2002), Fırat (Altuner & Gürbüz 1988, 1989, 1990a, b, et al. 2002; Karacaoğlu et al. 2004), West Black Sea 1991, 1994, 1996; Altuner & Pabucçu 1993, 1994; Şen River Basin (Aysel et al. 1992). et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Gürbüz & Altuner 2000), Gediz (Cirik 1994; Temel GENERAL DISTRIBUTION AND ECOLOGY. Ty- 1994, 1996, 1999, 2002, 2003; Gezerler et al. 1996, choplanktonic, mesosaprobous and mesotraphen- 1999), Kızılırmak (Özesm 1987; Kılınç & Dere 1988; tic species (Denys 1991; Hofmann 1994; Van Gönülol & Çomak 1990, 1992; Yıldız & Özkıran 1991; Dam et al. 1994), classifi ed as a circumneutral Gönülol 1993; Dere & Sıvacı 1994; Gönülol & Obalı (Van Dam et al. 1994) or an alkaliphilous diatom 1998; Kılınç 1998; Kılınç & Sıvacı 2001; Akbulut 2003), (Håkansson 1993). Konya Closed River Basin (Skuja 1937; Cirik et al. 1991; Temel 1992, 1996; Yıldız et al. 1994; Demirsoy 1996; Mitamura et al. 1997; Akköz & Obalı 1998; Elmacı 25. Fragilaria vaucheriae (Kützing) Petersen & Obalı 1998; Morkoyunlu et al. 1999), Küçük Mend- Figs 46–48 eres (Aysel et al. 1992, 1998a, b, 2001, 2002), Sakarya Fragilaria vaucheriae (Grunow) Patrick, F. capucina (Gönülol 1985a, 1987; Yıldız 1987a; Yıldız & Özkıran Desmaziéres var. vaucheriae (Kützing) Lange-Ber- 1994; Atıcı & Yıldız 1996; Aykulu et al. 1999; Yardımcı talot & Temel 2000; Atıcı 2002), Susurluk (Skuja 1937; Güner et al. 1987; Dere et al. 2002; Karacaören et al. 2004). Ref. Hustedt 1930 (p. 139, Fig. 130); Patrick C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 393

& Reimer 1966 (p. 120, Fig. 3: 14, 15); Krammer DISTRIBUTION IN TURKMEN MT. Türkmenbaba, & Lange-Bertalot 1991a (p. 124, Fig. 108: 10–15); İdris. Hofmann et al. 2011 (p. 277, Fig. 9: 1–7). DISTRIBUTION IN TURKEY. Lakes and rivers in DIMENSIONS. Valves 13.0–36.7 μm long and most of the river basins (Fig. 1). 3–5 μm wide, 9–17 striae in 10 μm. RIVER BASINS. Akarçay (Aykulu & Obalı 1981; Obalı DISTRIBUTION IN TURKMEN MT. Söğüt-Lütfi ye, 1982; Gönülol & Obalı 1986; Şen et al. 1994), Antalya Güllüdere-Türkmenbaba, Lütfi ye-Güllüdere. (Conk & Cirik 1991; Morkoyunlu & Ertan 1995a, b; Ertan et al. 1996; Morkoyunlu et al. 1996; Savaş & Cirik DISTRIBUTION IN TURKEY. Lakes and rivers in 1997; Yüce et al. 1997; Ertan & Morkoyunlu 1998), Aras Eastern Anatolia (Fırat, Aras, Çoruh) and Black (Altuner 1988; Akbulut & Yıldız 2002; Gürbüz et al. Sea (Kızılırmak) river basins (Fig. 1). 2002a, b; Kıvrak & Gürbüz 2005), Asi (Şen et al. 1997), Büyük Menderes (Cirik et al. 1989), Çoruh (Altuner RIVER BASINS. Akarçay (Gönülol & Obalı 1986; Şen et al. 1994), Aras (Altuner 1988; Akbulut & Yıldız 2002; 1984; Altuner & Aykulu 1987), Dicle (Sen 1988b; Şen Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Çoruh & Aksakal 1988; Şen et al. 1999; Baykal et al. 2004), (Altuner 1984; Altuner & Aykulu 1987), Dicle (Şen East Black Sea River Basin (Şahin 1992, 1994, 1998a, b, 1988a; Şen et al. 1999; Baykal et al. 2004), East Black 2000, 2001, 2002, 2003; Şahin & Gönülol 1997; Kolaylı Sea River Basin (Şahin 2002), Fırat (Altuner & Pabucçu et al. 1998; Kara & Şahin 2001), Fırat (Altuner & Gürbüz 1988, 1989, 1990a, b, 1991, 1993, 1994, 1996; Şen et al. 1988, 1989, 1990a, b, 1991, 1994, 1996; Altuner & Pa- 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Gürbüz bucçu 1993, 1994; Şen et al. 1995; Çetin & Şen 1998, & Altuner 2000), Gediz (Güner 1970; Cirik 1994; Gez- 2004; Gürbüz 2000; Gürbüz & Altuner 2000), Gediz erler et al. 1999), Kızılırmak (Kılınç & Dere 1988; (Cirik 1981, 1994; Temel 1994, 1996, 1999, 2002, 2003; Gönülol & Çomak 1990, 1992; Yıldız & Özkıran 1991; Gezerler et al. 1996, 1999), Kızılırmak (Kılınç & Dere Gönülol 1993; Dere & Sıvacı 1994; Gönülol & Obalı 1988; Kılınç 1998; Gönülol & Çomak 1990, 1992; Yıldız 1998; Kılınç 1998; Kılınç & Sıvacı 2001; İşbakan et al. & Özkıran 1991; Gönülol & Arslan 1992; Gönülol 1993; 2002), Konya Closed River Basin (Yıldız 1985a, 1986a, Dere & Sıvacı 1994; Kılınç & Sıvacı 2001; İşbakan et al. b, 1987b; Cirik et al. 1991; Demirsoy 1996; Akköz 2002), Konya Closed River Basin (Skuja 1937; Yıldız & Obalı 1998), Sakarya (Obalı 1984; Gönülol 1985a, 1984a, b, 1985a, b, 1986a, b, 1987b; Cirik et al. 1991; 1987; Yıldız 1987a; Obalı et al. 1989; Yıldız & Özkıran Temel 1992, 1996; Yıldız et al. 1994; Demirsoy 1996; 1994; Atıcı & Yıldız 1996; Aykulu et al. 1999; Yardımcı Akköz & Obalı 1998; Morkoyunlu et al. 1999; Mitamura & Temel 2000; Atıcı 2002), Susurluk (Skuja 1937; Dere et al. 1997; Elmacı & Obalı 1998), Küçük Menderes et al. 2002; Karacaören et al. 2004). (Skuja 1937; Aysel et al. 1992, 1998a, b, 2001, 2002; Erener et al. 1995), Marmara (Albay & Aykulu 1994a, GENERAL DISTRIBUTION AND ECOLOGY. Ty- b; Yardımcı & Temel 2000; Temel 1994, 1996, 1999, choplanktonic, mesosaprobous and eutraphentic 2002, 2003), Sakarya (Aykulu et al. 1983; Gönülol species (Lange-Bertalot 1979; Denys 1991; Hof- & Aykulu 1984; Obalı 1984; Ünal 1984; Gönülol 1985a, mann 1994; Van Dam et al. 1994), classifi ed as b, 1987; Yıldız 1987a; Dere 1989; Obalı et al. 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Demirsoy an alkaliphilous diatom (Håkansson 1993; Van 1996; Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı Dam et al. 1994). 2002), Seyhan (Kandemir-Çevik et al. 1994), Susurluk (Skuja 1937; Dere et al. 2002; Karacaoğlu et al. 2004), 26. Gomphonema olivaceum (Hornemann) Bré- Yeşilırmak (Yazıcı & Gönülol 1994; Altuner & Pabucçu bisson Figs 49 & 50 1996; Kır & Polat 1996; Gönülol & Obalı 1998). Gomphonema olivaceum (Lyngbye) Kützing GENERAL DISTRIBUTION AND ECOLOGY. Wide- Ref. Hustedt 1930 (p. 378, Fig. 719); Patrick & Re- spread, tychoplanktonic, mesosaprobous and meso- imer 1975 (p. 139, Fig. 8: 13–14); Krammer & Lange- eutraphentic species (Krammer & Lange-Bertalot Bertalot 1986 (p. 374, Fig. 165: 1–8); Hofmann et al. 1986; Denys 1991; Hofmann 1994; Van Dam et al. 2011 (p. 310, Fig. 95: 1–6). 1994), classifi ed as an alkaliphilous-alkalibiontic DIMENSIONS. Valves 15–38 μm long and 5.5– (Håkansson 1993) or an alkalibiontic diatom (Van 8.7 μm wide, 8–12 striae in 10 μm. Dam et al. 1994). 394 POLISH BOTANICAL JOURNAL 57(2). 2012

27. Gomphonema parvulum (Kützing) Kützing GENERAL DISTRIBUTION AND ECOLOGY. Wide- Figs 51, 52 & 152–157 spread, tychoplanktonic, polysaprobous and eu- traphentic diatom (Krammer & Lange-Bertalot Gomphonema parvulum (Kützing) Grunow 1986; Denys 1991; Hofmann 1994; Van Dam Ref. Hustedt 1930 (p. 372, Fig. 713a); Patrick & Re- et al. 1994), classifi ed as a circumneutral diatom imer 1975 (p. 122, Fig. 17: 7–12); Krammer & Lange- (Håkansson 1993; Van Dam et al. 1994). Bertalot 1986 (p. 358, Fig. 154: 1–25); Hofmann et al. 2011 (p. 312, Fig. 99: 1–10). 28. Hantzschia amphioxys (Ehrenberg) Grunow DIMENSIONS. Valves 9.5–33.0 μm long and Figs 53 & 54 5–8 μm wide, 9–16 striae in 10 μm. Ref. Hustedt 1930 (p. 394, Fig. 747); Krammer & Lange- Bertalot 1988 (p. 128, Fig. 88: 1–7); Hofmann et al. DISTRIBUTION IN TURKMEN MT. Türkmenbaba, 2011 (p. 333, Fig. 102: 1–5). Uluköy, Darıpınar, Çifteler, Çobanlar. DIMENSIONS. Valves 22–61 μm long and DISTRIBUTION IN TURKEY. Lakes and rivers 5–9 μm wide, 23–25 striae in 10 μm. in Eastern Anatolia (Fırat, Aras, Çoruh), Medi- terraean (Antalya, Asi), Black Sea (East Black DISTRIBUTION IN TURKMEN MT. Darıpınar, Sea River Basin, Yeşilırmak, Kızılırmak), Egean Çobanlar, Akındere. (Gediz, Küçük Menderes) and Inner Anatolia DISTRIBUTION IN TURKEY. Lakes and rivers in (Sakarya, Konya Closed River Basin, Kızılırmak) most of the river basins (Fig. 1). river basins (Fig. 1). RIVER BASINS. Akarçay (Aykulu & Obalı 1981; RIVER BASINS. Akarçay (Obalı 1982; Gönülol Obalı 1982; Gönülol & Obalı 1986; Şen et al. 1994), & Obalı 1986; Şen et al. 1994), Antalya (Conk & Cirik Antalya (Conk & Cirik 1991; Morkoyunlu & Ertan 1991; Morkoyunlu & Ertan 1995a, b; Ertan et al. 1996; 1995a, b; Ertan et al. 1996; Morkoyunlu et al. 1996; Morkoyunlu et al. 1996; Savaş & Cirik 1997; Yüce Savaş & Cirik 1997; Yüce et al. 1997; Ertan & Morko- et al. 1997; Ertan & Morkoyunlu 1998), Aras (Altuner yunlu 1998), Aras (Altuner 1988; Akbulut & Yıldız 1988; Akbulut & Yıldız 2002; Gürbüz et al. 2002a, b; 2002; Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Kıvrak & Gürbüz 2005), Asi (Şen et al. 1997), Çoruh Asi (Şen et al. 1997), Çoruh (Altuner 1984; Altuner (Altuner 1984; Altuner & Aykulu 1987), East Black Sea & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; River Basin (Şahin 1992, 1994, 1998a, b, 2000, 2001, Baykal et al. 2004), East Black Sea River Basin (Şahin 2002, 2003; Şahin & Gönülol 1997; Kolaylı et al. 1998; 1992, 1994, 1998a, b, 2000, 2001, 2002, 2003; Şahin Kara & Şahin 2001), Fırat (Altuner & Gürbüz 1988, & Gönülol 1997; Kolaylı et al. 1998; Kara & Şahin 1989, 1990a, b, 1991, 1994, 1996; Altuner & Pabucçu 2001), Fırat (Altuner & Gürbüz 1988, 1989, 1990a, b, 1993, 1994; Gürbüz 2000; Gürbüz & Altuner 2000), 1991, 1994, 1996; Şen & Çetin 1988; Altuner & Pabucçu Gediz (Gezerler et al. 1996), Kızılırmak (Özesm 1987; 1993, 1994; Şen et al. 1995; Çetin & Şen 1998, 2004; Kılınç & Dere 1988; Gönülol & Obalı 1998; Kılınç Gürbüz 2000; Gürbüz & Altuner 2000), Gediz (Gezerler 1998; Gönülol & Çomak 1990, 1992; Yıldız & Özkıran et al. 1996), Kızılırmak (Özesm 1987; Kılınç & Dere 1991; Gönülol & Arslan 1992; Gönülol 1993; Dere 1988; Gönülol & Çomak 1990, 1992; Yıldız & Özkıran & Sıvacı 1994; Kılınç & Sıvacı 2001; Akbulut 2003), 1991; Gönülol 1993; Dere & Sıvacı 1994; Kılınç 1998; Konya Closed River Basin (Skuja 1937; Temel 1992, Kılınç & Sıvacı 2001; Akbulut 2003), Konya Closed 1996; Yıldız et al. 1994; Morkoyunlu et al. 1999; River Basin (Skuja 1937; Yıldız 1984a, b, 1985a, b, Mitamura et al. 1997; Elmacı & Obalı 1998), Küçük 1986a, b, 1987b; Cirik et al. 1991; Temel 1992, 1996; Menderes (Aysel et al. 1992, 1998a, b, 2001, 2002; Yıldız et al. 1994; Demirsoy 1996; Mitamura et al. Balık & Gezerler 1995), Sakarya (Aykulu et al. 1983; 1997; Akköz & Obalı 1998; Elmacı & Obalı 1998; Gönülol & Aykulu 1984; Gönülol 1985b; Obalı 1984; Morkoyunlu et al. 1999), Küçük Menderes (Aysel Yıldız 1987a; Obalı et al. 1989; Yıldız & Özkıran 1994; et al. 1992, 1998a, b, 2001, 2002), Marmara (Albay Atıcı & Yıldız 1996; Demirsoy 1996; Aykulu et al. & Aykulu 1994a, b; Temel 1994, 1996, 1999, 2002, 1999; Yardımcı & Temel 2000; Atıcı 2002), Yeşilırmak 2003), Sakarya (Gönülol & Aykulu 1984; Obalı 1984; (Yazıcı & Gönülol 1994; Altuner & Pabuçcu 1996; Kır Ünal 1984; Gönülol 1985a, b, 1987; Yıldız 1987a; Dere & Polat 1996). 1989; Obalı et al. 1989; Yıldız & Özkıran 1994; Atıcı C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 395

Figs 55–72. 55 – Melosira varians Agardh, 56 & 57 – Meridion circulare (Gréville) Agardh, 58 & 59 – Navicula cincta (Ehrenberg) Ralfs, 60–62 – N. cryptocephala Kützing, 63 & 64 – N. radiosa Kützing, 65 & 66 – N. rhynchocephala Kützing, 67 & 68 – N. tripunctata (O.Müller) Bory, 69 & 70 – N. trivialis Lange-Bertalot, 71 & 72 – N. veneta Kützing. Scale bar = 10 μm. 396 POLISH BOTANICAL JOURNAL 57(2). 2012

& Yıldız 1996; Aykulu et al. 1999; Yardımcı & Temel 2003), Sakarya (Gönülol & Aykulu 1984; Obalı 1984; 2000; Atıcı 2002), Susurluk (Skuja 1937; Gezerler 1992; Gönülol 1985a, b, 1987; Yıldız 1987a; Obalı et al. 1989; Dere et al. 2002; Karacaören et al. 2004), Yeşilırmak Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu (Gönülol & Obalı 1998). et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), Seyhan (Kandemir-Çevik et al. 1994), Susurluk (Skuja 1937; GENERAL DISTRIBUTION AND ECOLOGY. Wide- Gezerler 1992; Dere et al. 2002; Karacaören et al. 2004), spread, tychoplanktonic and mesosaprobous spe- West Black Sea River Basin (Öztürk 1994), Yeşilırmak cies (Krammer & Lange-Bertalot 1988; Denys (Altuner & Pabuçcu 1996). 1991; Van Dam et al. 1994), classifi ed as a cir- cumneutral diatom (Håkansson 1993; Van Dam GENERAL DISTRIBUTION AND ECOLOGY. Ty- et al. 1994). choplanktonic, mesosaprobous and eutraphentic species (Lange-Bertalot 1979; Denys 1991; Van Dam et al. 1994), classifi ed as an alkaliphilous 29. Melosira varians Agardh Fig. 55 diatom (Håkansson 1993; Van Dam et al. 1994). Ref. Hustedt 1930 (p. 85, Fig. 41); Krammer & Lange- Bertalot 1991a (p. 7, Fig 4: 1–8); Hofmann et al. 2011 30. Meridion circulare (Gréville) Agardh (p. 357, Fig. 1: 6–9). Figs 56, 57, 150 & 151 DIMENSIONS. Valves 7.0–23.3 μm in diam- Ref. Hustedt 1930, (p. 130, Fig. 118); Patrick & Reimer eter. 1966 (p. 113, Fig. 2: 15); Krammer & Lange-Bertalot 1991a (p. 101, Fig. 101: 1–3); Hofmann et al. 2011 DISTRIBUTION IN TURKMEN MT. Darıpınar. (p. 359, Fig. 1: 1–14). DISTRIBUTION IN TURKEY. Lakes and rivers DIMENSIONS. Valves 27–62 μm long and in Mediterranean (Antalya), Eastern Anatolia 6–8 μm wide, 4–5 costae in 10 μm. (Fırat, Aras, Çoruh), Egean (Küçük Menderes, Gediz), Inner Anatolia (Konya Closed River Basin, DISTRIBUTION IN TURKMEN MT. Dümbüldek, Kızılırmak, Sakarya) river basins (Fig. 1). Güllüdere, Güllüdere-Lütfi ye, Gölcük.

RIVER BASINS. Akarçay (Aykulu & Obalı 1981), An- DISTRIBUTION IN TURKEY. Lakes and rivers in talya (Conk & Cirik 1991; Morkoyunlu & Ertan 1995a, b; Mediterranean (Antalya), Black Sea (East Black Ertan et al. 1996; Morkoyunlu et al. 1996; Savaş & Cirik Sea River Basin, West Black Sea River Basin), 1997; Yüce et al. 1997; Ertan & Morkoyunlu 1998), Eastern Anatolia (Fırat, Aras, Çoruh) river basins Aras (Altuner 1988; Akbulut & Yıldız 2002; Gürbüz (Fig. 1). et al. 2002a, b; Kıvrak & Gürbüz 2005), Çoruh (Altuner 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; Şen RIVER BASINS. Antalya (Conk & Cirik 1991; Mor- et al. 1999; Baykal et al. 2004), East Black Sea River koyunlu & Ertan 1995a, b; Morkoyunlu et al. 1996; Basin (Şahin 1992, 1994, 1998a, b, 2000, 2001, 2002, Ertan et al. 1996; Yüce et al. 1997; Savaş & Cirik 2003; Şahin & Gönülol 1997; Kolaylı et al. 1998; Kara 1997; Ertan & Morkoyunlu 1998), Aras (Altuner & Şahin 2001), Fırat (Altuner & Gürbüz 1988, 1989, 1988; Akbulut & Yıldız 2002; Gürbüz et al. 2002a, b; 1990a, b, 1991, 1994, 1996; Altuner & Pabucçu 1993, Kıvrak & Gürbüz 2005), Çoruh (Altuner 1984; Altuner 1994; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; 2000; Gürbüz & Altuner 2000), Gediz (Cirik 1994; Gez- Baykal et al. 2004), East Black Sea River Basin (Şahin erler et al. 1996, 1999), Kızılırmak (Gönülol & Çomak 1992, 1994, 1998a, b, 2000, 2001, 2002, 2003; Şahin 1990, 1992; Yıldız & Özkıran 1991; Gönülol 1993; & Gönülol 1997; Kolaylı et al. 1998; Kara & Şahin Dere & Sıvacı 1994; Gönülol & Obalı 1998; İşbakan 2001), Fırat (Altuner & Gürbüz 1988, 1989, 1990a, b, et al. 2002), Konya Closed River Basin (Skuja 1937; 1991, 1994, 1996; Altuner & Pabucçu 1993, 1994; Şen Cirik et al. 1991; Temel 1992, 1996; Yıldız et al. 1994; et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Demirsoy 1996; Morkoyunlu et al. 1996; Mitamura Gürbüz & Altuner 2000), Kızılırmak (Yıldız & Özkıran et al. 1997; Akköz & Obalı 1998; Elmacı & Obalı 1991; Dere & Sıvacı 1994), Konya Closed River Basin 1998), Küçük Menderes (Aysel et al. 1992, 1998a, b, (Skuja 1937; Yıldız 1985a, 1986a, b, 1987b; Temel 2001, 2002; Balık & Gezerler 1995), Marmara (Albay 1992, 1996; Yıldız et al. 1994; Morkoyunlu et al. 1999; & Aykulu 1994a, b; Temel 1994, 1996, 1999, 2002, Mitamura et al. 1997; Elmacı & Obalı 1998), Mar- C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 397 mara (Albay & Aykulu 1994a, b; Temel 1994, 1996, 32. Navicula cryptocephala Kützing 1999, 2002, 2003), Sakarya (Gönülol 1985a, 1987; Figs 60–62 & 158 Yıldız & Özkıran 1994), Seyhan (Kandemir-Çevik et al. 1994), Susurluk (Skuja 1937; Dere et al. 2002; Ref. Hustedt 1930 (p. 295, Fig. 496); Patrick & Reimer Karacaören et al. 2004), West Black Sea River Basin 1966 (p. 503, Fig. 48: 3); Krammer & Lange-Bertalot (Öztürk 1994). 1986 (p. 102, Fig. 30: 8–14); Lange-Bertalot 2001 (p. 27, Figs 17: 1–10, 18: 9–20); Hofmann et al. 2011 (p. 377, GENERAL DISTRIBUTION AND ECOLOGY. Wide- Fig. 31: 6–10). spread, tychoplanktonic, mesosaprobous and eut- DIMENSIONS. Valves 22.0–41.3 μm long and raphentic diatom (Lange-Bertalot 1979; Krammer 5–7 μm wide, 13–18 striae in 10 μm. & Lange-Bertalot 1991a; Denys 1991; Van Dam et al. 1994), classifi ed as an alkaliphilous diatom DISTRIBUTION IN TURKMEN MT. Darıpınar, (Håkansson 1993; Van Dam et al. 1994). Söğüt, Güllüdere, Türkmenbaba Çobanlar, Ul- uköy.

31. Navicula cincta (Ehrenberg) Ralfs DISTRIBUTION IN TURKEY. Lakes and rivers in Figs 58, 59 most of the river basins (Fig. 1).

Navicula cincta (Ehrenberg) Kützing, N. cincta (Ehren- RIVER BASINS. Akarçay (Aykulu & Obalı 1981; berg) Ralfs in Pritchard Obalı 1982), Antalya (Conk & Cirik 1991; Morkoy- Ref. Hustedt 1930, (p. 298, Fig. 510); Patrick & Reimer unlu & Ertan 1995a, b; Ertan et al. 1996; Morkoyunlu 1966 (p. 516, Fig. 49: 8); Krammer & Lange-Bertalot et al. 1996; Savaş & Cirik 1997; Yüce et al. 1997; Ertan 1986 (p. 98, Fig. 28: 8–15); Lange-Bertalot 2001 (p. 26, & Morkoyunlu 1998), Aras (Altuner 1988; Akbulut Fig. 41: 1–29); Hofmann et al. 2011 (p. 376, Fig. 32: & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 31–36). 2005), Asi (Şen et al. 1997), Büyük Menderes (Skuja 1937; Güner 1966, Cirik et al. 1989), Çoruh (Altuner DIMENSIONS. Valves 16.0–33.3 μm long and 1984), Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. 5–7 μm wide, 8–13 striae in 10 μm. 2004), East Black Sea River Basin (Şahin 1992, 1994, 1998a, b, 2000, 2001, 2002, 2003; Şahin & Gönülol DISTRIBUTION IN TURKMEN MT. Türkmenbaba, 1997; Kolaylı et al. 1998; Kara & Şahin 2001), Fırat Dümbüldek, Darıpınar, Güllüdere. (Altuner & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, 1996; Şen & Çetin 1988; Altuner & Pabucçu 1993, 1994; DISTRIBUTION IN TURKEY. Lakes and rivers in Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Egean (Gediz, Susurluk, Küçük Menderes) river Gürbüz & Altuner 2000), Gediz (Cirik 1994; Gezerler basins (Fig. 1). et al. 1996, 1999), Kızılırmak (Özesm 1987; Kılınç & Dere 1988; Gönülol & Çomak 1990, 1992; Yıldız RIVER BASINS. Asi (Şen et al. 1997), Dicle (Şen & Özkıran 1991; Gönülol & Arslan 1992; Gönülol 1993; 1988a; Şen et al. 1999; Baykal et al. 2004), Fırat (Şen Dere & Sıvacı 1994; Gönülol & Obalı 1998; Kılınç et al. 1995; Çetin & Şen 1998, 2004), Gediz (Gezerler 1998; Kılınç & Sıvacı 2001; İşbakan et al. 2002; Akbulut et al. 1996), Kızılırmak (Özesm 1987; Elmacı & Obalı 2003), Konya Closed River Basin (Skuja 1937; Yıldız 1992; Akbulut 2003), Küçük Menderes (Aysel et al. 1984a, b, 1985a, b, 1986a, b, 1987b; Cirik et al. 1991; 1992, 1998a, b, 2001, 2002), Sakarya (Obalı 1984; Temel 1992, 1996; Yıldız et al. 1994; Demirsoy 1996; Obalı et al. 1989), Susurluk (Skuja 1937; Dere et al. Morkoyunlu et al. 1996; Mitamura et al. 1997; Akköz 2002; Karacaören et al. 2004), Yeşilırmak (Altuner & Obalı 1998; Elmacı & Obalı 1998), Küçük Menderes & Pabuçcu 1996). (Aysel et al. 1992, 1998a, b, 2001, 2002), Marmara GENERAL DISTRIBUTION AND ECOLOGY. Wide- (Albay & Aykulu 1994a, b; Temel 1994, 1996, 1999, spread, tychoplanktonic, mesosaprobous and eu- 2002, 2003), Sakarya (Aykulu et al. 1983; Obalı 1984; Gönülol & Aykulu 1984; Ünal 1984; Gönülol 1985a, traphentic diatom (Lange-Bertalot 1979, 2001; b, 1987; Yıldız 1987a; Dere 1989; Obalı et al. 1989; Krammer & Lange-Bertalot 1986; Denys 1991; Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Demirsoy Hofmann 1994; Van Dam et al. 1994), classifi ed 1996; Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı as an alkaliphilous diatom (Håkansson 1993; Van 2002), Seyhan (Kandemir-Çevik et al. 1994), Susurluk Dam et al. 1994). (Skuja 1937; Dere et al. 2002; Karacaören et al. 2004), 398 POLISH BOTANICAL JOURNAL 57(2). 2012

Van Closed River Basin (Gessner 1957; Öztığ 1957), Konya Closed River Basin (Skuja 1937; Yıldız 1984a, b, West Black Sea River Basin (Aysel et al. 1992; Öztürk 1985a, b, 1986a, b, 1987b; Cirik et al. 1991; Yıldız et al. 1994; Cirik 1997), Yeşilırmak (Yazıcı & Gönülol 1994; 1994; Temel 1992, 1996; Demirsoy 1996; Mitamura Altuner & Pabuçcu 1996; Kır & Polat 1996). et al. 1997; Akköz & Obalı 1998; Elmacı & Obalı 1998; Morkoyunlu et al. 1999), Küçük Menderes (Aysel et al. GENERAL DISTRIBUTION AND ECOLOGY. Wide- 1992, 1998a, b, 2001, 2002; Balık & Gezerler 1995), spread, tychoplanktonic, mesosaprobous and eutra- Marmara (Albay & Aykulu 1994a, b; Temel 1994, 1996, phentic species (Krammer & Lange-Bertalot 1986; 1999, 2002, 2003), Sakarya (Gönülol & Aykulu 1984; Denys 1991; Hofmann 1994; Van Dam et al. 1994) Obalı 1984; Ünal 1984; Gönülol 1985a, b, 1987; Yıldız classifi ed as a circumneutral (Van Dam et al. 1994) 1987a; Dere 1989; Obalı et al. 1989; Yıldız & Özkıran or an alkaliphilous diatom (Håkansson 1993). 1994; Atıcı & Yıldız 1996; Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), Seyhan (Kandemir-Çevik 33. Navicula radiosa Kützing Figs 63 & 64 et al. 1994), Susurluk (Skuja 1937; Dere et al. 2002; Karacaören et al. 2004), West Black Sea River Basin Ref. Hustedt 1930 (p. 299, Fig. 513); Patrick & Reimer (Cirik 1997), Yeşilırmak (Yazıcı & Gönülol 1994; Kır 1966 (p. 509, Fig. 48: 15); Krammer & Lange-Bertalot & Polat 1996). 1986 (p. 99, Fig. 29: 1–4); Lange-Bertalot 2001 (p. 59, Fig. 8: 1–7); Hofmann et al. 2011 (p. 394, Fig. 35: GENERAL DISTRIBUTION AND ECOLOGY. Wide- 1–5). spread, tychoplanktonic, mesosaprobous and meso-eutraphentic species (Krammer & Lange- DIMENSIONS. Valves 54–85 μm long and 7.0– Bertalot 1986; Denys 1991; Hofmann 1994; Van 13.3 μm wide, 7–13 striae in 10 μm. Dam et al. 1994), classifi ed as circumneutral (Van DISTRIBUTION IN TURKMEN MT. Dümbüldek, Dam et al. 1994) or an alkaliphilous diatom (Hå- İdris, Söğüt-Lütfi ye, Lütfi ye-Güllüdere. kansson 1993).

DISTRIBUTION IN TURKEY. Lakes and rivers 34. Navicula rhynchocephala Kützing in Mediterranean (Antalya, Seyhan, Asi), Egean Figs 65 & 66 (Susurluk, Büyük Menderes, Küçük Menderes), Eastern Anatolia (Fırat, Aras, Çoruh), Black Sea Ref. Hustedt 1930 (p. 296, Fig. 501); Patrick & Reimer 1966 (p. 505, Fig. 48: 6); Krammer & Lange-Bertalot (Kızılırmak, East Black Sea River Basin, West 1986 (p. 101, Figs 30: 5–8; 31: 1, 2); Lange-Bertalot Black Sea River Basin) river basins (Fig. 1). 2001 (p. 64, Fig. 9: 6–10); Hofmann et al. 2011 (p. 396, RIVER BASINS. Akarçay (Obalı 1982; Gönülol Fig. 39: 12–17). & Obalı 1986; Şen et al. 1994), Antalya (Conk & Cirik DIMENSIONS. Valves 50–64 μm long and 8.0– 1991; Morkoyunlu & Ertan 1995a, b; Ertan et al. 1996; Morkoyunlu et al. 1996; Savaş & Cirik 1997; Yüce et al. 11.3 μm wide, 7–11 striae in 10 μm. 1997; Ertan & Morkoyunlu 1998), Aras (Altuner 1988; DISTRIBUTION IN TURKMEN MT. Çobanlar. Şen et al. 1997; Akbulut & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Büyük Menderes DISTRIBUTION IN TURKEY. Lakes and rivers in (Skuja 1937; Güner 1966), Çoruh (Altuner 1984; Altuner Mediterraean (Antalya), Eastern Anatolia (Fırat, & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; Aras Çoruh), Egean (Gediz, Susurluk, Küçük Baykal et al. 2004), East Black Sea River Basin (Şahin Menderes), Inner Anatolia (Konya Closed River 1992, 1994, 1998a, b, 2000, 2001, 2002, 2003; Şahin Basin), Black Sea (East Black Sea River Basin) & Gönülol 1997; Kolaylı et al. 1998; Kara & Şahin river basins (Fig. 1). 2001), Fırat (Altuner & Pabucçu 1993, 1994; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Gürbüz RIVER BASINS. Antalya (Conk & Cirik 1991; & Altuner 2000), Gediz (Cirik 1994; Temel 1994, Morkoyunlu & Ertan 1995a, b; Morkoyunlu et al. 1996; 1996, 1999, 2002, 2003; Gezerler et al. 1996, 1999), Savaş & Cirik 1997; Yüce et al. 1997; Ertan et al. 1996; Kızılırmak (Kılınç & Dere 1988; Gönülol & Çomak Ertan & Morkoyunlu 1998), Aras (Altuner 1988; Ak- 1990, 1992; Yıldız & Özkıran 1991; Gönülol & Ar- bulut & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak slan 1992; Gönülol 1993; Dere & Sıvacı 1994; Gönülol & Gürbüz 2005), Asi (Şen et al. 1997), Çoruh (Altuner & Obalı 1998; Kılınç 1998; Kılınç & Sıvacı 2001), 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; Şen C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 399 et al. 1999; Baykal et al. 2004), East Black Sea River Ertan & Morkoyunlu 1998), Aras (Altuner 1988; Ak- Basin (Şahin 1992, 1994, 1998a, b, 2000, 2001, 2002, bulut & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak 2003; Şahin & Gönülol 1997; Kolaylı et al. 1998; Kara & Gürbüz 2005), Asi (Şen et al. 1997), Çoruh (Altuner & Şahin 2001), Fırat (Altuner & Gürbüz 1988, 1989, 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; Şen 1990a, b, 1991, 1994, 1996; Altuner & Pabucçu 1993, et al. 1999; Baykal et al. 2004), East Black Sea River 1994; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz Basin (Şahin 1992, 1994, 1998a, b, 2000, 2001, 2002, 2000; Gürbüz & Altuner 2000), Gediz (Gezerler et al. 2003; Şahin & Gönülol 1997; Kolaylı et al. 1998; Kara 1996), Kızılırmak (Gönülol & Çomak 1990, 1992; & Şahin 2001), Fırat (Altuner & Gürbüz 1988, 1989, Gönülol 1993; Gönülol & Obalı 1998; İşbakan et al. 1990a, 1991; Altuner & Pabucçu 1993, 1994; Şen et al. 2002), Konya Closed River Basin (Skuja 1937; Temel 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Gürbüz 1992, 1996; Yıldız et al. 1994; Mitamura et al. 1997; & Altuner 2000), Gediz (Cirik 1994; Temel 1994, 1996, Elmacı & Obalı 1998; Morkoyunlu et al. 1999), Küçük 1999, 2002, 2003; Gezerler et al. 1996, 1999), Kızılırmak Menderes (Aysel et al. 1992, 1998a, b, 2001, 2002), (Kılınç & Dere 1988; Yıldız & Özkıran 1991; Gönülol Sakarya (Gönülol 1985a, 1987), Susurluk (Skuja 1937; & Arslan 1992; Dere & Sıvacı 1994; Kılınç 1998; Kılınç Dere et al. 2002; Karacaören et al. 2004). & Sıvacı 2001), Konya Closed River Basin (Skuja 1937; Yıldız 1984a, b, 1985a, b, 1986a, b, 1987b; Cirik et al. GENERAL DISTRIBUTION AND ECOLOGY. Wide- 1991; Temel 1992, 1996; Yıldız et al. 1994; Demirsoy spread, tychoplanktonic and mesosaprobous spe- 1996; Morkoyunlu et al. 1996; Mitamura et al. 1997; cies (Krammer & Lange-Bertalot 1986; Denys Akköz & Obalı 1998; Elmacı & Obalı 1998), Küçük 1991; Hofmann 1994; Van Dam et al. 1994). Na- Menderes (Aysel et al. 1992, 1998a, b, 2001, 2002; Balık vicula rhynchocephala is a circumneutral (Håkans- & Gezerler 1995), Marmara (Albay & Aykulu 1994a, b; son 1993) or an alkalihilous diatom (Van Dam Temel 1994, 1996, 1999, 2002, 2003), Sakarya (Gönülol et al. 1994). & Aykulu 1984; Obalı 1984; Yıldız 1987a; Gönülol 1985b; Obalı et al. 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu et al. 1999; Yardımcı & Temel 35. Navicula tripunctata (O. Müller) Bory 2000; Atıcı 2002), Seyhan (Kandemir-Çevik et al. 1994). Figs 67, 68, 160 & 161 Susurluk (Skuja 1937; Dere et al. 2002; Karacaören Navicula gracilis Ehrenberg et al. 2004), West Black Sea River Basin (Öztürk 1994), Yeşilırmak (Yazıcı & Gönülol 1994; Altuner & Pabuçcu Ref. Hustedt 1930 (p. 299, Fig. 514); Patrick & Reimer 1996; Kır & Polat 1996). 1966 (p. 513, Fig. 49: 3); Krammer & Lange-Bertalot 1986 (p. 95, Fig. 27: 1–3); Lange-Bertalot 2001 (p. 73, GENERAL DISTRIBUTION AND ECOLOGY. Wide- Fig. 1: 1–8); Hofmann et al. 2011 (p. 403, Fig. 35: spread, tychoplanktonic, mesosaprobous and eu- 11–16). traphentic species (Krammer & Lange-Bertalot DIMENSIONS. Valves 32–80 μm long and 1986; Denys 1991; Hofmann 1994; Van Dam 6–11 μm wide, 9–12 striae in 10 μm. et al. 1994), classifi ed as an alkaliphilous diatom (Håkansson 1993; Van Dam et al. 1994). DISTRIBUTION IN TURKMEN MT. Söğüt-Lütfi ye, Lütfi ye-Güllüdere, Güllüdere, Güllüdere-Türk- 36. Navicula trivialis Lange-Bertalot menbaba, İdris, Akındere. Figs 69 & 70

DISTRIBUTION IN TURKEY. Lakes and rivers Ref. Krammer & Lange-Bertalot 1986 (p. 110, Fig. 35: in Mediterranean (Antalya, Seyhan), Black Sea 1–4); Lange-Bertalot 2001 (p. 73, Fig. 29: 1–6); Hof- (East Black Sea River Basin, West Black Sea River mann et al. 2011 (p. 403, Fig. 33: 11–15). Basin, Kızılırmak), Egean (Gediz, Küçük Mender- DIMENSIONS. Valves 27.0–43.3 μm long and ess), Inner Anatolia (Konya Closed River Basin, 7.0–10.7 μm wide, 10–14 striae in 10 μm. Kızılırmak, Sakarya), Eastern Anatolia (Fırat, Aras, Çoruh) river basins (Fig. 1). DISTRIBUTION IN TURKMEN MT. Dümbüldek, Darıpınar, Seydi. RIVER BASINS. Antalya (Conk & Cirik 1991; Morko- yunlu & Ertan 1995a, b; Ertan et al. 1996; Morkoyunlu DISTRIBUTION IN TURKEY. Lakes and rivers in et al. 1996; Savaş & Cirik 1997; Yüce et al. 1997; Eastern Anatolia (Fırat River Basin) (Fig. 1). 400 POLISH BOTANICAL JOURNAL 57(2). 2012

RIVER BASINS. Dicle (Şen 1988a; Şen et al. 1999; Akköz & Obalı 1998; Elmacı & Obalı 1998), Küçük Baykal et al. 2004), Fırat (Şen & Çetin 1988; Şen et al. Menderes (Cirik & Cirik 1989), Sakarya (Gönülol 1995; Çetin & Şen 1998, 2004). & Aykulu 1984; Obalı 1984; Ünal 1984; Gönülol 1985a, b, 1987; Yıldız 1987a; Dere 1989; Obalı et al. 1989; GENERAL DISTRIBUTION AND ECOLOGY. Wide- Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu spread, tychoplanktonic, mesosaprobous and eu- et al. 1999; Atıcı 2002; Yardımcı & Temel 2000), Su- traphentic species (Krammer & Lange-Bertalot surluk (Skuja 1937; Dere et al. 2002; Karacaören et al. 1986; Denys 1991; Hofmann 1994; Van Dam et al. 2004), West Black Sea River Basin (Öztürk 1994). 1994), classifi ed as an alkaliphilous diatom (Van GENERAL DISTRIBUTION AND ECOLOGY. Wide- Dam et al. 1994). spread, tychoplanktonic, mesopoly- or polys- aprobous and eutraphentic species (Krammer 37. Navicula veneta Kützing Figs 71, 72 & 159 & Lange-Bertalot 1986; Denys 1991; Hofmann Navicula cryptocephala var. veneta (Kützing) Grunow 1994; Van Dam et al. 1994), classifi ed as an alka- Ref. Hustedt 1930 (p. 295, Fig. 497a); Patrick & Reimer liphilous diatom (Van Dam et al. 1994). 1966 (p. 504, Fig. 48: 5); Krammer & Lange-Bertalot 1986 (p. 104, Fig. 32: 1–4); Lange-Bertalot 2001 (p. 74, 38. Nitzschia acicularis (Kützing) W. Smith Fig. 14: 23–30); Hofmann et al. 2011 (p. 406, Fig. 9: Figs 73 & 74 8–12). Nitzschia acicularis W. Smith DIMENSIONS. Valves 18.0–26.7 μm long and Ref. Hustedt 1930 (p. 423, Fig. 821); Krammer & Lange- 4.0–6.7 μm wide, 13–17 striae in 10 μm. Bertalot 1988 (p. 123, Fig. 85: 1–4). ISTRIBUTION IN URKMEN T D T M . Dümbüldek, DIMENSIONS. Valves 36–130 μm long and Darıpınar, İnli. 2–5 μm wide, 17–19 fi bulae in 10 μm. DISTRIBUTION IN TURKEY. Lakes and rivers DISTRIBUTION IN TURKMEN MT. İdris, Çoban- in Inner Anatolia (Akarçay, Konya Closed River lar, Seydi, Lütfi ye-Güllüdere. Basin, Sakarya, Kızılırmak), Egean (Gediz, Su- surluk, Büyük Menderes, Küçük Menderes), DISTRIBUTION IN TURKEY. Lakes and rivers in Eastern Anatolia (Fırat, Aras, Çoruh), Black Sea Mediterranean (Antalya), Eastern Anatolia (Fırat, (East Black Sea River Basin, West Black Sea River Aras), Inner Anatolia (Konya Closed River Basin, Basin) river basins (Fig. 1). Sakarya), Egean (Gediz, Küçük Menderes) river basins (Fig. 1). RIVER BASINS. Akarçay (Obalı 1982; Gönülol & Obalı 1986; Şen et al. 1994), Aras (Kıvrak & Gürbüz RIVER BASINS. Akarçay (Aykulu & Obalı 1981; 2005), Büyük Menderes (Skuja 1937; Güner 1966), Gönülol & Obalı 1986; Şen et al. 1994), Antalya (Conk Çoruh (Altuner 1984; Altuner & Aykulu 1987; Gürbüz & Cirik 1991; Morkoyunlu & Ertan 1995a, b; Ertan et al. 2002a, b), Dicle (Şen 1988a; Şen et al. 1999; et al. 1996; Morkoyunlu et al. 1996; Savaş & Cirik Baykal et al. 2004), East Black Sea River Basin (Şahin 1997; Yüce et al. 1997; Ertan & Morkoyunlu 1998), 1992, 1994, 1998a, b, 2000, 2001, 2002, 2003; Şahin Aras (Altuner 1988; Akbulut & Yıldız 2002; Gürbüz & Gönülol 1997; Kolaylı et al. 1998; Kara & Şahin et al. 2002a, b; Kıvrak & Gürbüz 2005), Büyük Men- 2001), Fırat (Altuner & Gürbüz 1990b, 1994, 1996; deres (Cirik et al. 1989), Çoruh (Altuner 1984; Altuner Altuner & Pabucçu 1993, 1994; Şen et al. 1995; Çetin & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; & Şen 1998, 2004; Gürbüz 2000; Gürbüz & Altuner Baykal et al. 2004), East Black Sea River Basin (Şahin 2000), Gediz (Güner 1970; Cirik 1994; Gezerler et al. 2002), Fırat (Altuner & Gürbüz 1988, 1989, 1990a, 1999), Kızılırmak (Gönülol & Çomak 1990, 1992; Yıldız b, 1991, 1994, 1996; Altuner & Pabucçu 1993, 1994; & Özkıran 1991; Gönülol & Arslan 1992; Gönülol 1993; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz Dere & Sıvacı 1994; Gönülol & Obalı 1998; İşbakan 2000; Gürbüz & Altuner 2000), Gediz (Cirik 1994; et al. 2002), Konya Closed River Basin (Skuja 1937; Temel 1994, 1996, 1999, 2002, 2003; Gezerler et al. Yıldız 1984a, b, 1985a, b, 1986a, b, 1987b; Cirik et al. 1996, 1999), Kızılırmak (Kılınç & Dere 1988; Gönülol 1991; Temel 1992, 1996; Yıldız et al. 1994; Demirsoy & Çomak 1990, 1992; Yıldız & Özkıran 1991; Gönülol 1996; Morkoyunlu et al. 1996; Mitamura et al. 1997; & Arslan 1992; Gönülol 1993; Dere & Sıvacı 1994; C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 401

Figs 73–91. 73 & 74 – Nitzschia acicularis (Kützing) W. Smith, 75–77 – N. amphibia Grunow, 78–80 – N. dissipata (Kützing) Grunow, 81–83 – N. fonticola Grunow, 84–86 – N. frustulum (Kützing) Grunow, 87–89 – N. linearis (Agardh) W. Smith, 90 & 91 – N. palea (Kützing) W. Smith. Scale bar = 10 μm.

Gönülol & Obalı 1998; Kılınç 1998; Kılınç & Sıvacı Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), 2001; İşbakan et al. 2002), Konya Closed River Basin Susurluk (Skuja 1937; Güner et al. 1987; Gezerler 1992; (Skuja 1937; Yıldız 1985a, 1986a, b, 1987b; Temel Dere et al. 2002; Karacaören et al. 2004), West Black 1992,1996; Yıldız et al. 1994; Mitamura et al. 1997; Sea River Basin (Öztürk 1994; Cirik 1997), Yeşilırmak Elmacı & Obalı 1998; Morkoyunlu et al. 1999), Küçük (Altuner & Pabuçcu 1996). Menderes (Aysel et al. 1992, 1998a, b, 2001, 2002; Balık & Gezerler 1995), Marmara (Albay & Aykulu 1994a, b; GENERAL DISTRIBUTION AND ECOLOGY. Wide- Temel 1994, 1996, 1999, 2002, 2003), Sakarya (Obalı spread, tychoplanktonic, mesosaprobous and eu- 1984; Ünal 1984; Yıldız 1987a; Dere 1989; Obalı et al. traphentic species (Krammer & Lange-Bertalot 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; 1986; Denys 1991; Hofmann 1994; Van Dam et al. 402 POLISH BOTANICAL JOURNAL 57(2). 2012

1994), classifi ed as an alkaliphilous diatom (Van DIMENSIONS. Valves 16–45 μm long and 3.5– Dam et al. 1994). 7.0 μm wide, 7–11 fi bulae in 10 μm.

DISTRIBUTION IN TURKMEN MT. Dümbüldek, 39. Nitzschia amphibia Grunow Figs 75–77 Söğüt-Lütfi ye, Çobanlar, İdris, Akındere. Ref. Hustedt 1930 (p. 414, Fig. 793); Krammer & Lange- Bertalot 1988 (p. 108, Fig. 78: 13–21); Hofmann et al. DISTRIBUTION IN TURKEY. Lakes and rivers 2011 (p. 433, Fig. 117: 9–15). in Eastern Anatolia (Fırat, Aras, Çoruh), Egean (Gediz, Küçük Menderes), Black Sea (Kızılırmak), DIMENSIONS. Valves 18–40 μm long and Inner Anatolia (Sakarya) river basins (Fig. 1). 4–5 μm wide, 7–9 fi bulae and 14–18 striae in 10 μm. RIVER BASINS. Akarçay (Obalı 1982; Gönülol & Obalı 1986; Şen et al. 1994), Aras (Altuner 1988; DISTRIBUTION IN TURKMEN MT. Darıpınar, Akbulut & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak Güllüdere-Türkmenbaba, Çobanlar, İdris, İnli, & Gürbüz 2005), Asi (Şen et al. 1997), Büyük Mend- Akındere. eres (Skuja 1937; Güner 1966), Çoruh (Altuner 1984; Altuner & Aykulu 1987), East Black Sea River Basin DISTRIBUTION IN TURKEY. Lakes and rivers in (Şahin 2002), Fırat (Altuner & Gürbüz 1988, 1989, Eastern Anatolia (Fırat), Black Sea (Kızılırmak, 1990a, b, 1991, 1994, 1996; Altuner & Pabucçu 1993, East Black Sea River Basin) river basins (Fig. 1). 1994; Gürbüz 2000; Gürbüz & Altuner 2000), Gediz (Cirik 1994; Temel 1994, 1996, 1999, 2002, 2003; Gez- RIVER BASINS. Akarçay (Gönülol & Obalı 1986; erler et al. 1996, 1999), Kızılırmak (Gönülol & Çomak Şen et al. 1994), Aras (Gürbüz et al. 2002a, b; Kıvrak 1990, 1992; Yıldız & Özkıran 1991; Gönülol & Arslan & Gürbüz 2005), Çoruh (Altuner 1984; Altuner 1992; Gönülol 1993; Dere & Sıvacı 1994; Gönülol & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; & Obalı 1998), Konya Closed River Basin (Yıldız Baykal et al. 2004), East Black Sea River Basin (Şahin 1985a; 1986a; b; 1987b), Küçük Menderes (Cirik 2002), Fırat (Altuner & Gürbüz 1988, 1989, 1990a, b, & Cirik 1989; Aysel et al. 1992, 1998a, b, 2001, 2002), 1991, 1994, 1996; Altuner & Pabucçu 1993, 1994; Şen Sakarya (Gönülol & Aykulu 1984; Gönülol 1985b; et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Yıldız 1987a, b; Yıldız & Özkıran 1994; Atıcı & Yıldız Gürbüz & Altuner 2000), Kızılırmak (Gönülol & Çomak 1996; Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı 1990, 1992; Yıldız & Özkıran 1991; Gönülol & Arslan 2002), Susurluk (Skuja 1937; Dere et al. 2002; Kara- 1992; Gönülol 1993; Dere & Sıvacı 1994; Gönülol caören et al. 2004), Yeşilırmak (Yazıcı & Gönülol 1994; & Obalı 1998), Konya Closed River Basin (Cirik et al. Kır & Polat 1996). 1991; Demirsoy 1996; Akköz & Obalı 1998), Sakarya (Obalı 1984; Obalı et al. 1989; Yıldız & Özkıran 1994), GENERAL DISTRIBUTION AND ECOLOGY. Susurluk (Skuja 1937; Dere et al. 2002; Karacaören Widespread, tychoplanktonic, mesosaprobous et al. 2004), Yeşilırmak (Yazıcı & Gönülol 1994; Altuner and meso-eutraphentic or eutraphentic species & Pabuçcu 1996; Kır & Polat 1996). (Lange-Bertalot 1979; Krammer & Lange-Bertalot GENERAL DISTRIBUTION AND ECOLOGY. 1986; Denys 1991; Hofmann 1994; Van Dam et al. Widespread, tychoplanktonic, mesosaprobous 1994). Nitzschia dissipata is an alkaliphilous dia- and eutraphentic species (Lange-Bertalot 1979; tom (Håkansson 1993; Van Dam et al. 1994). Krammer & Lange-Bertalot 1986; Denys 1991; Hofmann 1994; Van Dam et al. 1994), classifi ed 41. Nitzschia fonticola Grunow Figs 81–83 as alkaliphilous diatom (Håkansson 1993; Van Ref. Hustedt 1930 (p. 415, Fig. 800); Krammer & Lange- Dam et al. 1994). Bertalot 1988 (p. 103, Fig. 75: 1–22); Hofmann et al. 2011 (p. 259, Fig. 9: 8–12). 40. Nitzschia dissipata (Kützing) Grunow Figs 78–80 DIMENSIONS. Valves 11–49 μm long and 3–5 μm wide, 10–16 fi bulae in 10 μm. Ref. Hustedt 1930 (p. 412, Fig. 789); Krammer & Lange- Bertalot 1988 (p. 19, Fig. 11: 1–7); Hofmann et al. 2011 DISTRIBUTION IN TURKMEN MT. Türkmenbaba, (p. 441, Fig. 109: 8–18). Dümbüldek, Darıpınar, Akındere. C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 403

DISTRIBUTION IN TURKEY. Lakes and riv- 1994), Konya Closed River Basin (Skuja 1937; Yıldız ers in Inner Anatolia (Sakarya, Konya Closed 1984a, b, 1985a, b, 1986a, b, 1987b; Temel 1992, 1996; River Basin), Eastern Anatolia (Fırat), Black Sea Yıldız et al. 1994; Mitamura et al. 1997; Elmacı & Obalı (Kızılırmak) river basins (Fig. 1). 1998; Morkoyunlu et al. 1999), Sakarya (Aykulu et al. 1983; Gönülol & Aykulu 1984; Gönülol 1985b; RIVER BASINS. Akarçay (Gönülol & Obalı 1986; Demirsoy 1996), Van Closed River Basin (Gessner Şen et al. 1994), Antalya (Skuja 1937; Vardar & Güner 1957; Öztığ 1957), Yeşilırmak (Yazıcı & Gönülol 1994; 1972), Aras (Altuner 1988; Akbulut & Yıldız 2002; Kır & Polat 1996). Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Çoruh (Altuner 1984; Altuner & Aykulu 1987), East GENERAL DISTRIBUTION AND ECOLOGY. Wide- Black Sea River Basin (Şahin 2002), Fırat (Altuner spread, tychoplanktonic, mesosaprobous and eu- & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, 1996; traphentic species (Krammer & Lange-Bertalot Altuner & Pabucçu 1993, 1994; Gürbüz 2000; Gürbüz 1986; Denys 1991; Hofmann 1994; Van Dam & Altuner 2000), Kızılırmak (Gönülol & Çomak 1990, et al. 1994), classifi ed as an alkaliphilous diatom 1992; Elmacı & Obalı 1992; Gönülol 1993; Gönülol (Håkansson 1993; Van Dam et al. 1994). & Obalı 1998), Konya Closed River Basin (Skuja 1937; Temel 1992, 1996; Yıldız et al. 1994; Mitamura et al. 1997; Elmacı & Obalı 1998; Morkoyunlu et al. 43. Nitzschia linearis (Agardh) W. Smith 1999), Sakarya (Gönülol & Aykulu 1984; Obalı 1984; Figs 87–89 Gönülol 1985b; Obalı et al. 1989; Yıldız & Özkıran Nitzschia linearis W. Smith 1994), Susurluk (Skuja 1937; Dere et al. 2002; Ka- racaören et al. 2004), Van Closed River Basin (Öztığ Ref. Hustedt 1930 (p. 409, Fig. 784); Krammer & Lange- 1957; Gessner 1957). Bertalot 1988 (p. 69, Fig. 55: 1–4); Hofmann et al. 2011 (p. 452, Fig. 106: 1–3). GENERAL DISTRIBUTION AND ECOLOGY. Ty- choplanktonic, mesosaprobous and eutraphentic DIMENSIONS. Valves 56–132 μm long and species (Lange-Bertalot 1979; Denys 1991; Hof- 3.0–6.7 μm wide, 9–15 fi bulae in 10 μm. mann 1994; Van Dam et al. 1994), classifi ed as DISTRIBUTION IN TURKMEN MT. Dümbüldek, an alkaliphilous diatom (Håkansson 1993; Van Darıpınar, Güllüdere-Türkmenbaba, Çobanlar, Dam et al. 1994). Akındere, Çifteler.

42. Nitzschia frustulum (Kützing) Grunow DISTRIBUTION IN TURKEY. Lakes and rivers in Figs 84–86 & 164 Mediterranean (Antalya), Inner Anatolia (Konya Closed River Basin, Kızılırmak, Sakarya), Eastern Ref. Hustedt 1930 (p. 414, Fig. 795); Krammer & Lange- Anatolia (Fırat), Egean (Gediz, Küçük Menderes) Bertalot 1988 (p. 94, Fig. 68: 1–8); Hofmann et al. 2011 (p. 445, Fig. 112: 28–40). river basins (Fig. 1). RIVER BASINS. Antalya (Conk & Cirik 1991; DIMENSIONS. Valves 9.0–28.7 μm long and Morkoyunlu & Ertan 1995a, b; Ertan et al. 1996; 3.0–4.3 μm wide, 9–12 fi bulae and 20–26 striae Morkoyunlu et al. 1996; Savaş & Cirik 1997; Yüce in 10 μm. et al. 1997; Ertan & Morkoyunlu 1998), Asi (Şen et al. 1997), Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. DISTRIBUTION IN TURKMEN MT. Dümbüldek, 2004), Fırat (Şen & Çetin 1988; Altuner & Gürbüz Darıpınar, Güllüdere-Türkmenbaba, İdris, İnli. 1990b, 1994, 1996; Şen et al. 1995; Çetin & Şen DISTRIBUTION IN TURKEY. Lakes and rivers 1998, 2004), Gediz (Gezerler et al. 1996), Kızılırmak in Inner Anatolia (Konya Closed River Basin, (Kılınç & Dere 1988; Gönülol & Çomak 1990, Kızılırmak, Sakarya) river basins (Fig. 1). 1992; Yıldız & Özkıran 1991; Gönülol 1993; Dere & Sıvacı 1994; Gönülol & Obalı 1998; Kılınç 1998; RIVER BASINS. Akarçay (Obalı 1982), Aras (Al- Kılınç & Sıvacı 2001), Konya Closed River Basin tuner 1988; Akbulut & Yıldız 2002), Fırat (Altuner (Skuja 1937; Yıldız 1984a, b, 1985a, b, 1986a, b, & Gürbüz 1988, 1989, 1990a, 1991), Kızılırmak (Yıldız 1987b; Cirik et al. 1991; Temel 1992, 1996; Yıldız & Özkıran 1991; Elmacı & Obalı 1992; Dere & Sıvacı et al. 1994; Mitamura et al. 1997; Demirsoy 1996; 404 POLISH BOTANICAL JOURNAL 57(2). 2012

Morkoyunlu et al. 1996; Akköz & Obalı 1998; Elmacı Gönülol & Arslan 1992; Gönülol 1993; Dere & Sıvacı & Obalı 1998), Küçük Menderes (Aysel et al. 1992, 1994; Gönülol & Obalı 1998; Kılınç 1998; Kılınç 1998a, b, 2001, 2002), Marmara (Albay & Aykulu & Sıvacı 2001; Akbulut 2003), Konya Closed River 1994a, b; Temel 1994, 1996, 1999, 2002, 2003), Basin (Skuja 1937; Yıldız 1984a, b, 1985a, b, 1986a, b, Sakarya (Aykulu et al. 1983; Yıldız 1987a; Yıldız 1987b; Cirik et al. 1991; Temel 1992, 1996; Yıldız et al. & Özkıran 1994; Atıcı & Yıldız 1996; Demirsoy 1994; Demirsoy 1996; Mitamura et al. 1997; Akköz 1996; Aykulu et al. 1999; Yardımcı & Temel 2000; & Obalı 1998; Elmacı & Obalı 1998; Morkoyunlu et al. Atıcı 2002), Susurluk (Skuja 1937; Dere et al. 2002; 1999), Küçük Menderes (Aysel et al. 1992, 1998a, b, Karacaören et al. 2004), Yeşilırmak (Yazıcı & Gönülol 2001, 2002; Balık & Gezerler 1995), Marmara (Albay 1994; Kır & Polat 1996). & Aykulu 1994a, b; Temel 1994, 1996, 1999, 2002, 2003), Sakarya (Aykulu et al. 1983; Gönülol & Aykulu GENERAL DISTRIBUTION AND ECOLOGY. Wide- 1984; Obalı 1984; Ünal 1984; Gönülol 1985a, b, 1987; spread, tychoplanktonic, mesosaprobous and meso- Yıldız 1987a; Dere 1989; Obalı et al. 1989; Yıldız eutraphentic or eutraphentic species (Krammer & Özkıran 1994; Atıcı & Yıldız 1996; Demirsoy 1996; & Lange-Bertalot 1986; Denys 1991; Hofmann Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), 1994; Van Dam et al. 1994), classifi ed as an alka- Susurluk (Skuja 1937; Dere et al. 2002; Karacaören liphilous diatom (Van Dam et al. 1994). et al. 2004), Yeşilırmak (Yazıcı & Gönülol 1994; Altuner & Pabuçcu 1996; Kır & Polat 1996).

44. Nitzschia palea (Kützing) W. Smith GENERAL DISTRIBUTION AND ECOLOGY. Wide- Figs 90, 91, 162 & 163 spread, tychoplanktonic, polysaprobous and eu- Ref. Hustedt 1930 (p. 416, Fig. 801); Krammer & Lange- traphentic species (Krammer & Lange-Bertalot Bertalot 1988 (p. 85, Figs 59: 1–24; 60: 1–6); Hofmann 1986; Denys 1991; Hofmann 1994; Van Dam et al. 2011 (p. 454, Fig. 111: 1–20). et al. 1994), classifi ed as a circumneutral diatom (Håkansson 1993; Van Dam et al. 1994). DIMENSIONS. Valves 20.0–55.7 μm long and 2.0–5.7 μm wide, 10–16 fi bulae in 10 μm. 45. Pinnularia borealis Ehrenberg Figs 92–94 DISTRIBUTION IN TURKMEN MT. Dümbüldek, İnli. Ref. Hustedt 1930 (p. 326, Fig. 597); Patrick & Reimer 1966 (p. 618, Fig. 58: 13); Krammer & Lange-Bertalot DISTRIBUTION IN TURKEY. Lakes and rivers 1986 (p. 405, Fig. 177: 1–12); Hofmann et al. 2011 in Inner Anatolia (Akarçay, Konya Closed River (p. 481, Fig. 75: 27–33). Basin, Kızılırmak, Sakarya), Eastern Anatolia DIMENSIONS. Valves 29.0–40.7 μm long and (Fırat, Aras), Black Sea (East Black Sea River 7–10 μm wide, 4–6 striae in 10 μm. Basin, Kızılırmak), Egean (Küçük Menderes) river basins (Fig. 1). DISTRIBUTION IN TURKMEN MT. Türkmenbaba, Çobanlar, İnli, Uluköy. RIVER BASINS. Akarçay (Aykulu & Obalı 1981; Obalı 1982; Gönülol & Obalı 1986; Şen et al. 1994), DISTRIBUTION IN TURKEY. Lakes and rivers Aras (Altuner 1988; Akbulut & Yıldız 2002; Gürbüz in Eastern Anatolia (Fırat, Aras, Çoruh), Black et al. 2002a, b; Kıvrak & Gürbüz 2005), Asi (Şen Sea (East Black Sea River Basin) river basins et al. 1997), Çoruh (Altuner 1984; Altuner & Aykulu (Fig. 1). 1987), Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. 2004), East Black Sea River Basin (Şahin 1992, 1994, RIVER BASINS. Aras (Altuner 1988; Akbulut 1998a, b, 2000, 2001, 2002, 2003; Şahin & Gönülol & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 1997; Kolaylı et al. 1998; Kara & Şahin 2001), Fırat 2005), Çoruh (Altuner 1984; Altuner & Aykulu 1987), (Altuner & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, East Black Sea River Basin (Şahin 1992, 1994, 1996; Şen & Çetin 1988; Altuner & Pabucçu 1993, 1998a, b, 2000, 2001, 2002, 2003; Şahin & Gönülol 1994; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 1997; Kolaylı et al. 1998; Kara & Şahin 2001), Fırat 2000; Gürbüz & Altuner 2000), Gediz (Gezerler et al. (Altuner & Gürbüz 1988, 1989, 1990a, 1991; Altuner 1996), Kızılırmak (Özesm 1987; Kılınç & Dere 1988; & Pabucçu 1993, 1994; Gürbüz 2000; Gürbüz & Altuner Gönülol & Çomak 1990, 1992; Yıldız & Özkıran 1991; 2000), Kızılırmak (Özesm 1987; Yıldız & Özkıran 1991; C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 405

Figs 92–109. 92–94 – Pinnularia borealis Ehrenberg, 95 – P. microstauron (Ehrenberg) Cleve, 96 – P. viridiformis Krammer, 97–99 – Planothidium lanceolatum (Brébisson ex Kützing) Lange-Bertalot, 100–102 – Pseudostaurosira brevistriata Grunow, 103 & 104 – Rhoicosphaenia abbreviata (C. Agardh) Lange-Bertalot, 105 & 106 – Sellaphora pupula (Kützing) Mereschkowsky, 107–109 – Staurosirella pinnata (Ehrenberg) D. M. Wiliams & Round. Scale bar = 10 μm.

Dere & Sıvacı 1994; Akbulut 2003), Sakarya (Yıldız GENERAL DISTRIBUTION AND ECOLOGY. The & Özkıran 1994), Susurluk (Skuja 1937; Dere et al. species is an acidophilous diatom (Håkansson 2002; Karacaören et al. 2004). 1993). 406 POLISH BOTANICAL JOURNAL 57(2). 2012

46. Pinnularia microstauron (Ehrenberg) Cleve GENERAL DISTRIBUTION AND ECOLOGY. Wide- Fig. 95 spread, tychoplanktonic and mesosaprobous spe- cies (Krammer & Lange-Bertalot 1986; Denys Ref. Hustedt 1930 (p. 320, Fig. 582); Patrick and Reimer 1966 (p.128, Fig. 14: 4); Krammer & Lange-Bertalot 1991; Van Dam et al. 1994), classifi ed as a cir- 1986 (p. 425, Fig. 191: 1–6; 192: 1–16); Hofmann et al. cumneutral diatom (Van Dam et al. 1994). 2011 (p. 485, Fig. 71: 9–14). 47. Pinnularia viridiformis Krammer Fig. 96 DIMENSIONS. Valves 29–50 μm long and 8–11 μm wide, 11–13 striae in 10 μm. Ref. Krammer 2000 (p. 167, Figs 160–162); Hofmann et al. 2011 (p. 493, Fig. 72: 3–4). DISTRIBUTION IN TURKMEN MT. Türkmenbaba, DIMENSIONS. Valves 58–82 μm long and 12.0– Çobanlar, İnli. 17.3 μm wide, 7–11 striae in 10 μm. DISTRIBUTION IN TURKEY. Lakes and rivers DISTRIBUTION IN TURKMEN MTS. Türkmen- in Eastern Anatolia (Fırat, Aras, Çoruh), Egean baba, Çobanlar, İnli. (Gediz, Büyük Menderes, Küçük Menderes), Black Sea (Yeşilırmak, East Black Sea River Basin, West DISTRIBUTION IN TURKEY. New for fl ora of Black Sea River Basin) river basins (Fig. 1). Turkey.

RIVER BASINS. Akarçay (Gönülol & Obalı 1986; GENERAL DISTRIBUTION AND ECOLOGY. Ty- Şen et al. 1994), Antalya (Skuja 1937; Vardar & Güner choplanktonic (Denys 1991) and mesosaprobic 1972), Asi (Şen et al. 1997), Aras (Altuner 1988; Ak- (Hofmann 1994), classifi ed as a circumneutral bulut & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak diatom (Håkansson 1993). & Gürbüz 2005), Büyük Menderes (Skuja 1937; Güner 1966), Çoruh (Altuner 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. 2004), 48. Planothidium lanceolatum (Brébisson ex East Black Sea River Basin (Şahin 2002), Fırat (Al- Kützing) Round & Bukhtiyarova Figs 97–99 tuner & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, Achnanthes lanceolata Brébisson, A. lanceolata Bré- 1996; Şen & Çetin 1988; Altuner & Pabucçu 1993, bisson ex Kützing, A. lanceolata (Brébisson) Grunow 1994; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz Ref. Hustedt 1930 (p. 207, Fig. 306a); Patrick & Re- 2000; Gürbüz & Altuner 2000), Gediz (Güner 1970; imer 1966 (p. 269, Fig. 18: 1–10); Krammer & Lange- Temel 1994, 1996, 1999, 2002, 2003; Gezerler et al. Bertalot 1991b (p. 75, Fig. 41: 11–13); Hofmann et al. 1996), Kızılırmak (Özesm 1987; Yıldız & Özkıran 2011 (p. 510, Fig. 24: 41–47). 1991; Gönülol & Arslan 1992; Dere & Sıvacı 1994; Akbulut 2003), Konya Closed River Basin (Skuja DIMENSIONS. Valves 6.0–39.3 μm long and 1937; Yıldız 1984a, b, 1985a, b, 1986a, b, 1987b; Cirik 4.0–9.3 μm wide, 9–18 striae in 10 μm. et al. 1991; Temel 1992, 1996; Yıldız et al. 1994; Demirsoy 1996; Morkoyunlu et al. 1996; Mitamura DISTRIBUTION IN TURKMEN MT. Darıpınar, et al. 1997; Akköz & Obalı 1998; Elmacı & Obalı Türkmenbaba, Çobanlar, İdris, Uluköy. 1998), Küçük Menderes (Aysel et al. 1992, 1998a, b, 2001, 2002; Balık & Gezerler 1995), Marmara (Albay DISTRIBUTION IN TURKEY. Lakes and rivers in & Aykulu 1994a, b; Temel 1994, 1996, 1999, 2002, Mediterranean (Antalya, Seyhan), Inner Anantolia 2003), Sakarya (Aykulu et al. 1983; Gönülol & Aykulu (Akarçay, Kızıırmak, Konya Closed River Basin, 1984; Obalı 1984; Ünal 1984; Gönülol 1985a, b, 1987; Sakarya), Eastern Anatolia (Fırat, Aras, Çoruh), Yıldız 1987a; Dere 1989; Obalı et al. 1989; Yıldız Egean (Gediz, Susurluk, Büyük, Küçük Mend- & Özkıran 1994; Atıcı & Yıldız 1996; Demirsoy 1996; eres), Black Sea (East Black Sea River Basin) river Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı basins (Fig. 1). 2002), Susurluk (Skuja 1937; Güner et al. 1987; Gez- erler 1992; Dere et al. 2002; Karacaören et al. 2004), RIVER BASINS. Akarçay (Obalı 1982), Antalya (Conk West Black Sea River Basin (Öztürk 1994), Yeşilırmak & Cirik 1991; Morkoyunlu & Ertan 1995a, b; Ertan et al. (Yazıcı & Gönülol 1994; Altuner & Pabuçcu 1996; 1996; Morkoyunlu et al. 1996; Savaş & Cirik 1997; Kır & Polat 1996). Yüce et al. 1997; Ertan & Morkoyunlu 1998), Aras C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 407

(Altuner 1988; Akbulut & Yıldız 2002; Gürbüz et al. Temel 1992, 1996; Yıldız et al. 1994; Mitamura et al. 2002a, b; Kıvrak & Gürbüz 2005), Büyük Menderes 1997; Elmacı & Obalı 1998; Morkoyunlu et al. 1999), (Skuja 1937; Güner 1966), Çoruh (Altuner 1984; Altuner Sakarya (Gönülol 1985a, 1987). & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. 2004), East Black Sea River Basin (Şahin GENERAL DISTRIBUTION AND ECOLOGY. Wide- 1992, 1994, 1998a, b, 2000, 2001, 2002, 2003; Şahin spread, tychoplanktonic, mesosaprobous and me- & Gönülol 1997; Kolaylı et al. 1998; Kara & Şahin sotraphentic species (Krammer & Lange-Bertalot 2001), Fırat (Altuner & Gürbüz 1988, 1989, 1990a, b, 1986; Denys 1991; Hofmann 1994; Van Dam 1991, 1994, 1996; Altuner & Pabucçu 1993, 1994; Şen et al. 1994), classifi ed as an alkaliphilous diatom et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; (Håkansson 1993; Van Dam et al. 1994). Gürbüz & Altuner 2000), Gediz (Gezerler et al. 1996), Kızılırmak (Kılınç & Dere 1988; Yıldız & Özkıran 1991; Gönülol & Arslan 1992; Dere & Sıvacı 1994; 50. Rhoicosphenia abbreviata (C. Agardh) Lange- Kılınç 1998; Kılınç & Sıvacı 2001), Konya Closed River Bertalot Figs 103, 104 & 165 Basin (Cirik et al. 1991; Demirsoy 1996; Akköz & Obalı Rhoicosphenia curvata (Kützing) Grunow, R. curvata 1998), Küçük Menderes (Aysel et al. 1992, 1998a, b, (Kützing) Grunow ex Rabenhorst 2001, 2002), Marmara (Albay & Aykulu 1994a, b; Temel Ref. Hustedt 1930 (p. 207, Fig. 306a); Patrick & Reimer 1994, 1996, 1999, 2002, 2003), Sakarya (Obalı 1984; 1966 (p. 282, Fig. 20: 1–5); Krammer & Lange-Bertalot Ünal 1984; Yıldız 1987a; Dere 1989; Obalı et al. 1989; 1986 (p. 381, Fig. 91: 20–28); Hofmann et al. 2011 Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu (p. 527, Fig. 18: 42–48). et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), Seyhan (Kandemir-Çevik et al. 1994), Susurluk (Skuja 1937; DIMENSIONS. Valves 15.0–32.7 μm long and Dere et al. 2002; Karacaören et al. 2004), West Black 7.0–10.7 μm wide, 11–18 striae in 10 μm. Sea River Basin (Öztürk 1994; Cirik 1997), Yeşilırmak (Altuner & Pabuçcu 1996). DISTRIBUTION IN TURKMEN MT. Çobanlar.

GENERAL DISTRIBUTION AND ECOLOGY. Ty- DISTRIBUTION IN TURKEY. Lakes and rivers choplanktonic, mesosaprobous and eutraphentic in Inner Anatolia (Akarçay, Kızılırmak, Konya species (Denys 1991; Hofmann 1994; Van Dam Closed River Basin, Sakarya), Mediterranean (An- et al. 1994), classifi ed as an alkaliphilous diatom talya, Asi, Seyhan), Eastern Anatolia (Fırat, Aras, (Håkansson 1993; Van Dam et al. 1994). Çoruh), Egean (Gediz, Büyük Menderes, Küçük Menderes), Black Sea (East Black Sea River Basin, 49. Pseudostaurosira brevistriata Grunow Kızılırmak, Yeşilırmak) river basins (Fig. 1).

Figs 100–102 RIVER BASINS. Akarçay (Aykulu & Obalı 1981; Ref. Hustedt 1930 (p. 145, Fig. 151); Patrick & Reimer Gönülol & Obalı 1986; Şen et al. 1994), Antalya (Conk 1966 (p.128, Fig. 4: 15); Krammer & Lange-Bertalot & Cirik 1991; Morkoyunlu & Ertan 1995a, b; Ertan et al. 1991a (p. 162, Fig. 130: 9–16); Hofmann et al. 2011 1996; Morkoyunlu et al. 1996; Savaş & Cirik 1997; (p. 258, Fig. 9: 25–29). Yüce et al. 1997; Ertan & Morkoyunlu 1998), Aras (Altuner 1988; Akbulut & Yıldız 2002; Gürbüz et al. DIMENSIONS. Valves 9.0–22.3 μm long and 2002a, b; Kıvrak & Gürbüz 2005), Asi (Şen et al. 1997), 3–5 μm wide, 13–15 striae in 10 μm. Büyük Menderes (Skuja 1937; Güner 1966), Çoruh (Al- tuner 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; DISTRIBUTION IN TURKMEN MT. Çobanlar, Şen & Aksakal 1988; Şen et al. 1999; Baykal et al. Darıpınar. 2004), East Black Sea River Basin (Şahin 1992, 1994, DISTRIBUTION IN TURKEY. Lakes and rivers 1998a, b, 2000, 2001, 2002, 2003; Şahin & Gönülol in Inner Anatolia (Konya Closed River Basin) 1997; Kolaylı et al. 1998; Kara & Şahin 2001), Fırat (Fig. 1). (Altuner & Gürbüz 1988, 1989, 1990a, 1991; Altuner & Pabucçu 1993, 1994; Şen et al. 1995; Çetin & Şen RIVER BASINS. Dicle (Şen 1988a; Şen et al. 1999; 1998, 2004; Gürbüz 2000; Gürbüz & Altuner 2000), Baykal et al. 2004), Fırat (Şen et al. 1995; Çetin & Şen Gediz (Güner 1969; Güner 1970; Cirik & Cirik 1989; 1998, 2004), Konya Closed River Basin (Skuja 1937; Gezerler et al. 1996), Kızılırmak (Özesm 1987; Gönülol 408 POLISH BOTANICAL JOURNAL 57(2). 2012

& Çomak 1990, 1992; Yıldız & Özkıran 1991; Elmacı & Pabucçu 1993, 1994; Şen et al. 1995; Çetin & Şen & Obalı 1992; Gönülol & Arslan 1992; Gönülol 1993; 1998, 2004; Gürbüz 2000; Gürbüz & Altuner 2000), Dere & Sıvacı 1994; Gönülol & Obalı 1998; İşbakan Gediz (Cirik 1994; Temel 1994, 1996, 1999, 2002, 2003; et al. 2002; Akbulut 2003), Konya Closed River Basin Gezerler et al. 1996, 1999), Kızılırmak (Özesm 1987; (Skuja 1937; Yıldız 1984a, b, 1985a, b, 1986a, b, Gönülol & Çomak 1990, 1992; Yıldız & Özkıran 1991; 1987b; Cirik et al. 1991; Temel 1992, 1996; Yıldız et al. Elmacı & Obalı 1992; Gönülol 1993; Dere & Sıvacı 1994; Mitamura et al. 1997; Demirsoy 1996; Akköz 1994; Akbulut 2003), Konya Closed River Basin (Skuja & Obalı 1998; Elmacı & Obalı 1998; Morkoyunlu et al. 1937; Yıldız 1985a, 1986a, b, 1987b; Cirik et al. 1991; 1996), Küçük Menderes (Aysel et al. 1992, 1998a, b, Temel 1992, 1996; Yıldız et al. 1994; Demirsoy 1996; 2001, 2002; Balık & Gezerler 1995), Marmara (Albay Morkoyunlu et al. 1996; Mitamura et al. 1997; Akköz & Aykulu 1994a, b; Temel 1994, 1996, 1999, 2002, & Obalı 1998; Elmacı & Obalı 1998), Küçük Mend- 2003), Sakarya (Gönülol & Aykulu 1984; Obalı 1984; eres (Skuja 1937; Aysel et al. 1992, 1998a, b, 2001, Gönülol 1985a, b, 1987; Yıldız 1987a; Obalı et al. 1989; 2002; Erener et al. 1995), Sakarya (Obalı 1984; Ünal Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu 1984; Gönülol 1985a, 1987; Yıldız 1987a; Obalı et al. et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), Seyhan 1988; Dere 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız (Kandemir-Çevik et al. 1994), Susurluk (Skuja 1937; 1996; Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı Gezerler 1992; Dere et al. 2002; Karacaören et al. 2004), 2002), Susurluk (Skuja 1937; Gezerler 1992; Dere et al. West Black Sea River Basin (Öztürk 1994), Yeşilırmak 2002; Karacaoğlu et al. 2004), Yeşilırmak (Gönülol (Yazıcı & Gönülol 1994; Altuner & Pabuçcu 1996; Kır & Obalı 1998). & Polat 1996). GENERAL DISTRIBUTION AND ECOLOGY. Ty- GENERAL DISTRIBUTION AND ECOLOGY. Wide- choplanktonic, mesosaprobous and eutraphen- spread, tychoplanktonic, mesosaprobous and eu- tic species (Denys 1991; Hofmann 1994; Van traphentic species (Krammer & Lange-Bertalot Dam et al. 1994), classifi ed as an alkaliphilous 1986; Denys 1991; Hofmann 1994; Van Dam (Håkansson 1993) or alcalibiontic diatom (Van et al. 1994), classifi ed as an alkaliphilous diatom Dam et al. 1994). (Håkansson 1993; Van Dam et al. 1994). 52. Sellaphora pupula (Kützing) Mereschkow sky 51. Rhopalodia gibba (Ehrenberg) Müller Figs 105, 106, 166 & 167 Fig. 117 Navicula pupula Kützing Ref. Hustedt 1930 (p. 390, Fig. 740); Patrick & Reimer Ref. Hustedt 1930 (p. 281, Fig. 467a); Patrick & Reimer 1975 (p. 189, Fig. 28: 1); Krammer & Lange-Bertalot 1966 (p. 495, Fig. 47: 7); Krammer & Lange-Bertalot 1988 (p. 159, Fig. 59: 1, 2, 4, 6, 7–13); Hofmann et al. 1986 (p. 190, Fig. 68: 1–12); Hofmann et al. 2011 2011 (p. 529, Fig. 122: 4–8). (p. 536, Fig. 41: 1–14). DIMENSIONS. Valves 48–100 μm long and DIMENSIONS. Valves 13–31 μm long and 10.0–12.7 μm wide, 13–15 fi bulae in 10 μm. 4.0–8.7 μm wide, 18–24 striae in 10 μm. DISTRIBUTION IN TURKMEN MT. Çobanlar, DISTRIBUTION IN TURKMEN MT. Dümbüldek, Darıpınar. Darıpınar, Söğüt, Güllüdere-Türkmenbaba, Türk- DISTRIBUTION IN TURKEY. Lakes and rivers menbaba, Çobanlar, İdris, Uluköy. in Eaean (Gediz), Eastern Anatolia (Fırat), Inner DISTRIBUTION IN TURKEY. Lakes and rivers in Anatolia (Konya Closed River Basin, Sakarya) Eastern Anatolia (Fırat, Aras, Çoruh), Black Sea river basins (Fig. 1). (East Black Sea River Basin, Kızılırmak), Egean RIVER BASINS. Akarçay (Gönülol & Obalı 1986; (Gediz, Büyük Menderes, Küçük Menderes), Inner Şen et al. 1994), Aras (Altuner 1988; Akbulut & Yıldız Anatolia (Konya Closed River Basin, Sakarya, 2002; Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Kızılırmak) river basins (Fig. 1). Çoruh (Altuner 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. 2004), East RIVER BASINS. Aras (Altuner 1988; Akbulut & Yıldız Black Sea River Basin (Şahin 2002), Fırat (Altuner 2002; Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 409

Figs 110–116. 110–112 – Surirella angusta Kützing, 113 & 114 – S. brebissonii Krammer & Lange-Bertalot, 115 – S. ovalis Brébisson, 116 – S. tenera Gregory. Scale bar = 10 μm. 410 POLISH BOTANICAL JOURNAL 57(2). 2012

Büyük Menderes (Skuja 1937; Güner 1966), Çoruh (Al- RIVER BASINS. Aras (Altuner 1988; Akbulut tuner 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; & Yıldız 2002; Gürbüz et al. 2002a, b; Kıvrak Şen et al. 1999; Baykal et al. 2004), East Black Sea & Gürbüz 2005), Asi (Şen et al. 1997), Çoruh (Altuner River Basin (Şahin 1992, 1994, 1998a, b, 2000, 2001, 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; 2002, 2003; Şahin & Gönülol 1997; Kolaylı et al. 1998; Şen et al. 1999; Baykal et al. 2004), East Black Sea Kara & Şahin 2001), Fırat (Altuner & Gürbüz 1988, River Basin (Şahin 1992, 1994, 1998a, b, 2000, 2001, 1989, 1990a, b, 1991, 1994, 1996; Şen & Çetin 1988; 2002, 2003; Şahin & Gönülol 1997; Kolaylı et al. Altuner & Pabucçu 1993, 1994; Şen et al. 1995; Çetin 1998; Kara & Şahin 2001), Fırat (Altuner & Gürbüz & Şen 1998, 2004; Gürbüz 2000; Gürbüz & Altuner 1990b, 1994, 1996; Altuner & Pabucçu 1993, 1994; 2000), Gediz (Cirik 1994; Temel 1994, 1996, 1999, 2002, Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2003; Gezerler et al. 1996, 1999), Kızılırmak (Kılınç 2000; Gürbüz & Altuner 2000), Gediz (Gezerler et al. & Dere 1988; Gönülol & Çomak 1990, 1992; Yıldız 1996), Kızılırmak (Özesm 1987; Yıldız & Özkıran & Özkıran 1991; Gönülol & Arslan 1992; Gönülol 1993; 1991; Gönülol & Arslan 1992; Dere & Sıvacı 1994; Dere & Sıvacı 1994; Gönülol & Obalı 1998; Kılınç Akbulut 2003), Konya Closed River Basin (Cirik et al. 1998; Kılınç & Sıvacı 2001), Konya Closed River Basin 1991; Demirsoy 1996; Akköz & Obalı 1998), Küçük (Yıldız 1984a, b, 1985a, b, 1986a, b, 1987b), Küçük Menderes (Aysel et al. 1992, 1998a, b, 2001, 2002), Menderes (Aysel et al. 1992, 1998a, b, 2001, 2002), Sakarya (Obalı 1984; Yıldız 1987a; Obalı et al. 1989; Marmara (Albay & Aykulu 1994a, b; Temel 1994, 1996, Atıcı & Yıldız 1996; Aykulu et al. 1999; Yardımcı 1999, 2002, 2003), Sakarya (Gönülol & Aykulu 1984; & Temel 2000; Atıcı 2002), West Black Sea River Obalı 1984; Ünal 1984; Gönülol 1985a, b, 1987; Yıldız Basin (Öztürk 1994). 1987a; Dere 1989; Obalı et al. 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu et al. 1999; Yardımcı GENERAL DISTRIBUTION AND ECOLOGY. Ty- & Temel 2000; Atıcı 2002), Seyhan (Kandemir-Çevik choplanktonic and mesosaprobous species (Denys et al. 1994), Susurluk (Skuja 1937; Dere et al. 2002; 1991; Hofmann 1994; Van Dam et al. 1994), clas- Karacaören et al. 2004), West Black Sea River Basin sifi ed as an alkaliphilous diatom (Håkansson 1993; (Cirik 1997), Yeşilırmak (Altuner & Pabuçcu 1996). Van Dam et al. 1994). GENERAL DISTRIBUTION AND ECOLOGY. Tycho- planktonic, mesosaprobous and meso-eutraphentic 54. Surirella angusta Kützing Figs 110–112 species (Denys 1991; Hofmann 1994; Van Dam Ref. Hustedt 1930 (p. 435, Fig. 844–845); Krammer et al. 1994), classifi ed as a circumneutral diatom & Lange-Bertalot 1988 (p. 187, Figs 133: 6–13; 134: 1, (Håkansson 1993; Van Dam et al. 1994). 6–10); Hofmann et al. 2011 (p. 554, Fig. 131: 1–5). DIMENSIONS. Valves 24.0–50.7 μm long and 53. Staurosirella pinnata (Ehrenberg) 7.0–10.7 μm wide, 6–7 fi bulae in 10 μm. D. M. Wiliams & Round Figs 107–109 DISTRIBUTION IN TURKMEN MT. Türkmenbaba, Ref. Hustedt 1930 (p. 142, Fig. 141); Patrick & Reimer Dümbüldk, Çobanlar. 1966 (p. 127, Fig. 4: 10); Krammer & Lange-Bertalot 1991a (p. 156, Fig. 133: 1–18, 32, 32a); Hofmann et al. DISTRIBUTION IN TURKEY. Lakes and rivers in 2011 (p. 272, Fig. 10: 30–35). Mediterranean (Antalya), Black Sea (East Black Sea River Basin), Eastern Anatolia (Fırat), Egean DIMENSIONS. Valves 4.5–20.0 μm long and (Küçük Menderes), Inner Anatolian (Konya Closed 3.5–6.0 μm wide, 9–14 striae in 10 μm. River Basin, Sakarya) river basins (Fig. 1).

DISTRIBUTION IN TURKMEN MT. Dümbüldek, RIVER BASINS. Antalya (Conk & Cirik 1991; Morko- Türkmenbaba, Çobanlar, İnli, Seydi. yunlu & Ertan 1995a, b; Ertan et al. 1996; Morkoyunlu et al. 1996; Savaş & Cirik 1997; Yüce et al. 1997; Ertan DISTRIBUTION IN TURKEY. Lakes and rivers & Morkoyunlu 1998), Aras (Gürbüz et al. 2002a, b; in Inner Anatolian (Konya Closed River Basin, Kıvrak & Gürbüz 2005), Büyük Menderes (Skuja 1937; Kızılırmak, Sakarya), Egean (Gediz, Susurluk, Güner 1966), Çoruh (Altuner 1984; Altuner & Aykulu Küçük Menderes), Eastern Anatolia (Fırat, Aras, 1987), East Black Sea River Basin (Şahin 1992, 1994, Çoruh) river basins (Fig. 1). 1998a, b, 2000, 2001, 2002, 2003; Şahin & Gönülol C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 411

Figs 117–122. 117 – Rhopalodia gibba (Ehrenberg) Müller, 118 & 119 – Tryblionella hungarica (Grunow) Frenguelli, 120–122 – Ulnaria ulna (Nitzsch) P. Compère. Scale bar = 10 μm.

1997; Kolaylı et al. 1998; Kara & Şahin 2001), Fırat Kızılırmak (Yıldız & Özkıran 1991; Gönülol & Ar- (Altuner & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, slan 1992; Dere & Sıvacı 1994), Konya Closed River 1996; Altuner & Pabucçu 1993, 1994; Gürbüz 2000; Basin (Skuja 1937; Yıldız 1984a, b, 1985a, b, 1986a, Gürbüz & Altuner 2000), Gediz (Gezerler et al. 1996), b, 1987b; Cirik et al. 1991; Temel 1992, 1996; Yıldız 412 POLISH BOTANICAL JOURNAL 57(2). 2012

Figs 123–130. 123 & 124 – Achnanthidium minutissimum (Kützing) Czarnecki, 125–127 – Amphora pediculus (Kützing) Grunow, 128–130 – Cocconeis pediculus Ehrenberg, external (123, 125, 128, 130) and internal (124, 126, 127, 129) view, all SEM. Scale bars: 123–125 = 5 μm, 126 = 3 μm, 127 = 1 μm, 128–130 = 10 μm. et al. 1994; Demirsoy 1996; Morkoyunlu et al. 1996; Sakarya (Gönülol & Aykulu 1984; Gönülol 1985a, Mitamura et al. 1997; Akköz & Obalı 1998; Elmacı b; 1987; Yıldız 1987a; Yıldız & Özkıran 1994; Atıcı & Obalı 1998), Küçük Menderes (Aysel et al. 1992, & Yıldız 1996; Aykulu et al. 1999; Yardımcı & Temel 1998a, b, 2001, 2002), Marmara (Albay & Aykulu 2000; Atıcı 2002), Susurluk (Skuja 1937; Dere et al. 1994a, b; Temel 1994, 1996, 1999, 2002, 2003), 2002; Karacaören et al. 2004). C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 413

Figs 131–141. 131 & 132 – Cocconeis placentula var. lineata (Ehrenberg) Van Heurck, 133–135 – Craticula cuspidata (Kützing) D. G. Mann, 136–141 – Cymbella cymbiformis Agardh, external (131, 136, 137, 141) and internal (132–135, 138–140) view; all SEM. Scale bars: 131, 132, 136, 138 = 10 μm; 133 = 30 μm; 134, 135, 137, 141 = 5 μm; 139 & 140 = 3 μm.

GENERAL DISTRIBUTION AND ECOLOGY. Ty- et al. 1994), classifi ed as an alkaliphilous diatom choplanktonic, mesosaprobous and eutraphentic (Van Dam et al. 1994). species (Denys 1991; Hofmann 1994; Van Dam 414 POLISH BOTANICAL JOURNAL 57(2). 2012

55. Surirella brebissonii Krammer species (Denys 1991; Hofmann 1994), classifi ed as & Lange-Bertalot Figs 113 & 114 an alkaliphilous diatom (Van Dam et al. 1994). Ref. Krammer & Lange-Bertalot 1988 (p. 179, Figs 123: 4–5; 126: 2–6; 127: 1–13); Hofmann et al. 2011 56. Surirella ovalis Brébisson Fig. 115 (p. 556, Fig. 130: 11–21). Ref. Hustedt 1930 (p. 441, Fig. 860–861); Krammer DIMENSIONS. Valves 21.5–43.0 μm long and & Lange-Bertalot 1988 (p. 178, Fig. 125: 1–7); Hofmann 7.0–10.5 μm wide, 6 fi bulae in 10 μm. et al. 2011 (p. 558, Fig. 129: 8–10).

DISTRIBUTION IN TURKMEN MT. Akındere, DIMENSIONS. Valves 31.0–84.7 μm long and Çobanlar, Güllüdere, İdris. 19.0–37.3 μm wide, 3–5 fi bulae and 14–16 striae in 10 μm. DISTRIBUTION IN TURKEY. Lakes and rivers in most of the river basins (Fig. 1). DISTRIBUTION IN TURKMEN MT. İdris.

RIVER BASINS. Akarçay (Aykulu & Obalı 1981; Obalı DISTRIBUTION IN TURKEY. Lakes and rivers in 1982), Antalya (Conk & Cirik 1991; Morkoyunlu & Ertan most of the river basins (Fig. 1). 1995a, b; Ertan et al. 1996; Morkoyunlu et al. 1996; Savaş & Cirik 1997; Yüce et al. 1997; Ertan & Morkoy- RIVER BASINS. Akarçay (Obalı 1982; Gönülol unlu 1998), Aras (Altuner 1988; Akbulut & Yıldız 2002; & Obalı 1986; Şen et al. 1994), Antalya (Conk Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Asi (Şen & Cirik 1991; Morkoyunlu & Ertan 1995a, b; Ertan et al. 1997), Çoruh (Altuner 1984; Altuner & Aykulu et al. 1996; Morkoyunlu et al. 1996; Yüce et al. 1997; 1987), Dicle (Şen 1988a; Şen et al. 1999; Baykal et al. Savaş & Cirik 1997; Ertan & Morkoyunlu 1998), Aras 2004), East Black Sea River Basin (Şahin 1992, 1994, (Altuner 1988; Akbulut & Yıldız 2002; Gürbüz et al. 1998a, b, 2000, 2001, 2002, 2003; Şahin & Gönülol 1997; 2002a, b; Kıvrak & Gürbüz 2005), Çoruh (Altuner Kolaylı et al. 1998; Kara & Şahin 2001), Fırat (Altuner 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; Şen & Gürbüz 1988, 1989, 1990a, b, 1991, 1994, 1996; Şen et al. 1999; Baykal et al. 2004), East Black Sea River & Çetin 1988; Altuner & Pabucçu 1993, 1994; Şen et al. Basin (Şahin 1992, 1994, 1998a, b, 2000, 2001, 2002, 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Gürbüz 2003; Şahin & Gönülol 1997; Kolaylı et al. 1998; Kara & Altuner 2000), Gediz (Güner 1970; Temel 1994, 1996, & Şahin 2001), Fırat (Altuner & Pabucçu 1993, 1994; 1999, 2002, 2003; Gezerler et al. 1996), Kızılırmak Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; (Kılınç & Dere 1988; Gönülol & Çomak 1990, 1992; Gürbüz & Altuner 2000), Gediz (Gezerler et al. 1996), Yıldız & Özkıran 1991; Elmacı & Obalı 1992; Gönülol Kızılırmak (Kılınç & Dere 1988; Yıldız & Özkıran 1991; & Arslan 1992; Gönülol 1993; Dere & Sıvacı 1994; Gönülol & Arslan 1992; Dere & Sıvacı 1994; Kılınç Gönülol & Obalı 1998; Kılınç 1998; Kılınç & Sıvacı 1998; Kılınç & Sıvacı 2001), Konya Closed River Basin 2001; İşbakan et al. 2002), Konya Closed River Basin (Skuja 1937; Cirik et al. 1991; Temel 1992, 1996; Yıldız (Skuja 1937; Yıldız 1984a, b, 1985a, b, 1986a, b, 1987b; et al. 1994; Demirsoy 1996; Morkoyunlu et al. 1996; Temel 1992,1996; Yıldız et al. 1994; Mitamura et al. Mitamura et al. 1997; Akköz & Obalı 1998; Elmacı 1997; Morkoyunlu et al. 1999; Elmacı & Obalı 1998), & Obalı 1998), Küçük Menderes (Aysel et al. 1992, Küçük Menderes (Cirik & Cirik 1989; Aysel et al. 1992, 1998a, b, 2001, 2002; Balık & Gezerler 1995), Marmara 1998a, b, 2001, 2002), Marmara (Albay & Aykulu (Albay & Aykulu 1994a, b; Temel 1994, 1996, 1999, 1994a, b; Temel 1994, 1996, 1999, 2002, 2003), Sakarya 2002, 2003), Sakarya (Gönülol & Aykulu 1984; Obalı (Aykulu et al. 1983; Gönülol & Aykulu 1984; Ünal 1984; 1984; Gönülol 1985a, b, 1987; Yıldız 1987a; Obalı et al. Gönülol 1985a, b, 1987; Yıldız 1987a; Dere 1989; Yıldız 1989; Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; & Özkıran 1994; Atıcı & Yıldız 1996; Demirsoy 1996; Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), Aykulu et al. 1999; Yardımcı & Temel 2000; Atıcı 2002), Susurluk (Skuja 1937; Dere et al. 2002; Karacaören Seyhan (Kandemir-Çevik et al. 1994), Susurluk (Skuja et al. 2004), West Black Sea River Basin (Öztürk 1994), 1937; Dere et al. 2002; Karacaören et al. 2004), West Yeşilırmak (Altuner & Pabuçcu 1996). Black Sea River Basin (Öztürk 1994), Yeşilırmak (Altuner GENERAL DISTRIBUTION AND ECOLOGY. Wide- & Pabuçcu 1996). spread, tychoplanktonic, mesosaprobous and eutra- GENERAL DISTRIBUTION AND ECOLOGY. Ty- phentic species (Krammer & Lange-Bertalot 1986; choplanktonic, mesosaprobous and eutraphentic Denys 1991; Van Dam et al. 1994). C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 415

57. Surirella tenera Gregory Fig. 116 et al. 1999), Kızılırmak (Yıldız & Özkıran 1991; Gönülol & Arslan 1992; Dere & Sıvacı 1994), Konya Closed River Ref. Hustedt 1930 (p. 401, Fig. 766); Krammer & Lange- Basin (Skuja 1937; Yıldız 1984a, b, 1985a, b, 1986a, b, Bertalot 1988 (p. 203, Figs 164: 1–4; 165: 1–3). 1987b; Temel 1992, 1996; Yıldız et al. 1994; Mitamura DIMENSIONS. Valves 31–100 μm long and et al. 1997; Elmacı & Obalı 1998; Morkoyunlu et al. 19.0–28.7 μm wide, 2–5 fi bulae. 1999), Marmara (Albay & Aykulu 1994a, b; Temel 1994, 1996, 1999, 2002, 2003; Yardımcı & Temel 2000), DISTRIBUTION IN TURKMEN MT. Çobanlar, Sakarya (Obalı 1984; Yıldız 1987a; Obalı et al. 1989; Darıpınar. Yıldız & Özkıran 1994; Atıcı & Yıldız 1996; Aykulu et al. 1999; Atıcı 2002; Yardımcı & Temel 2000), Susurluk DISTRIBUTION IN TURKEY. Lakes and rivers in (Skuja 1937; Dere et al. 2002; Karacaören et al. 2004), Black Sea (East Black Sea River Basin), Eastern West Black Sea River Basin (Öztürk 1994). Anatolia (Fırat) river basins (Fig. 1). GENERAL DISTRIBUTION & ECOLOGY. Wide- RIVER BASINS. Aras (Gürbüz et al. 2002a, b; spread, euplanktonic, mesosaprobous and eutra- Kıvrak & Gürbüz 2005), Çoruh (Altuner 1984; Altuner & Aykulu 1987), Dicle (Şen 1988a; Şen et al. 1999; phentic (Krammer & Lange-Bertalot 1986; Denys Baykal et al. 2004), East Black Sea River Basin (Şahin 1991; Hofmann 1994; Van Dam et al. 1994), clas- 1992, 1994, 1998a, b, 2000, 2001, 2002, 2003; Şahin sifi ed as an alkaliphilous diatom (Håkansson 1993; & Gönülol 1997; Kolaylı et al. 1998; Kara & Şahin Van Dam et al. 1994). 2001), Fırat (Altuner & Gürbüz 1988, 1989, 1990a, 1991; Şen & Çetin 1988; Altuner & Pabucçu 1993, 1994; 59. Ulnaria ulna (Nitzsch) P. Compère Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz 2000; Figs 120–122 Gürbüz & Altuner 2000), Konya Closed River Basin (Yıldız 1984a, b, 1985a, b, 1986a, b,1987b), Yeşilırmak Synedra ulna (Nitzsch) Ehrenberg, Fragilaria ulna (Altuner & Pabuçcu 1996). (Nitzsch) Lange-Bertalot Ref. Hustedt 1930 (p. 151, Figs 158,159); Patrick GENERAL DISTRIBUTION AND ECOLOGY. Wide- & Reimer 1966 (p. 148, Fig. 7: 1, 2); Krammer & Lange- spread, euplanktonic, mesosaprobous and eutra- Bertalot 1991a (p. 143, Fig. 122: 1–8); Hofmann et al. phentic species (Krammer & Lange-Bertalot 1986; 2011 (p. 276, Fig. 5: 6–11). Denys 1991; Van Dam et al. 1994), classifi ed as an alkaliphilous diatom (Håkansson 1993; Van DIMENSIONS. Valves 64–145 μm long and Dam et al. 1994). 2–9 μm wide, 9–15 striae in 10 μm. DISTRIBUTION IN TURKMEN MT. Darıpınar, 58. Tryblionella hungarica (Grunow) Frenguelli İdrıs, Lütfi ye-Güllüdere. Figs 118 & 119 DISTRIBUTION IN TURKEY. Lakes and rivers in Nitzschia hungarica Grunow most of the river basins (Fig. 1). Ref. Hustedt 1930 (p. 401, Fig. 766); Krammer & Lange- Bertalot 1988 (p. 42, Fig. 34: 1–3). RIVER BASINS. Akarçay (Aykulu & Obalı 1981; Obalı 1982; Gönülol & Obalı 1986; Şen et al. 1994), Antalya DIMENSIONS. Valves 69.0–90.7 μm long and (Conk & Cirik 1991; Morkoyunlu & Ertan 1995a, b; 6–8 μm wide, 9–11 fi bulae, and 15–19 striae in Ertan et al. 1996; Morkoyunlu et al. 1996, Savaş & Cirik 10 μm. 1997; Yüce et al. 1997; Ertan & Morkoyunlu 1998), Aras (Altuner 1988; Şen et al. 1997; Akbulut & Yıldız 2002; DISTRIBUTION IN TURKMEN MT. Dümbüldek. Gürbüz et al. 2002a, b; Kıvrak & Gürbüz 2005), Büyük DISTRIBUTION IN TURKEY. Lakes and rivers Menderes (Skuja 1937; Güner 1966; Cirik et al. 1989), in Inner Anatolia (Konya Closed River Basin, Çoruh (Altuner 1984; Altuner & Aykulu 1987), Dicle (Şen Sakarya), Egean (Gediz) and Marmara (Marmara) 1988a; Şen et al. 1999; Baykal et al. 2004), East Black Sea River Basin (Şahin 1992, 1994, 1998a, b, 2000, 2001, river basins (Fig. 1). 2002, 2003, Kolaylı et al. 1998; Şahin & Gönülol 1997; RIVER BASINS. Aras (Altuner 1988; Akbulut & Yıldız Kara & Şahin 2001), Fırat (Altuner & Gürbüz 1988, 1989, 2002), Asi (Şen et al. 1997), Gediz (Cirik 1994; Gezerler 1990a, b, 1991, 1994, 1996; Altuner & Pabucçu 1993, 416 POLISH BOTANICAL JOURNAL 57(2). 2012

Figs 142–157. 142 – Cyclotella ocellata Pantocsek, 143 – Denticula tenuis Kützing, 144 –147 – Diatoma vulgaris Bory, 148 & 149 – Fragilaria capucina Desmazieres s.l., 150 & 151 – Meridion circulare (Gréville) Agardh, 152–157 – Gomphonema parvulum (Kützing) Kützing, external (142, 144, 146, 147, 154–157) and internal (143, 145, 148–153) view; all SEM. Scale bars: 142 & 154 = 5 μm; 143 = 4 μm; 144–146, 150, 152 = 10 μm; 147, 151, 153 = 5 μm; 148, 149 = 15 μm; 155–157 = 2 μm. C. N. SOLAK & A. Z. WOJTAL: DIATOMS IN SPRINGS AND STREAMS OF TÜRKMEN MT. 417

Figs 158–167. 158 – Navicula cryptocephala Kützing, 159 – N. veneta Kützing, 160 & 161– N. tripunctata (O. Müller) Bory, 162, 163 – Nitzschia palea (Kützing) W. Smith, 164 – N. frustulum (Kützing) Grunow, 165 – Rhoicosphaenia abbreviata (C. Agardh) Lange-Bertalot, 166 & 167 – Sellaphora pupula (Kützing) Mereschkowsky, internal (158) and external (159–167) view; all SEM. Scale bars: 158, 159, 162, 165, 166 = 10 μm; 160 = 20 μm; 161 & 167 = 3 μm; 163 = 2 μm; 164 = 5 μm. 418 POLISH BOTANICAL JOURNAL 57(2). 2012

1994; Şen et al. 1995; Çetin & Şen 1998, 2004; Gürbüz Bol ve Yaygın Türlerin Dağılımı. In: XIV. Ulusal Su Ürün- 2000; Gürbüz & Altuner 2000), Gediz (Güner 1969; Cirik leri Sempozyumu, p. 28. Muğla Üniversitesi, Muğla. 1994; Cirik & Cirik 1989, 1990; Gezerler et al. 1999), ÁCS É., BORICS G., FEHÉR G., KISS K. T., RESKÓNE N. M., Kızılırmak (Özesm 1987; Kılınç & Dere 1988; Kılınç STENGER-KOVÁCS C. & VÁBRÍRÓ G. 2009. Implementa- 1998; Gönülol & Çomak 1990, 1992; Yıldız & Özkıran tion of the European Water Framework Directive to as- 1991; Gönülol & Arslan 1992; Gönülol 1993; Dere sessment the water quality of Hungarian running waters & Sıvacı 1994; Kılınç & Sıvacı 2001; Akbulut 2003), with diatoms. Diatomededelingen 33: 29–33. Konya Closed River Basin (Skuja 1937; Yıldız 1984a, AKBULUT A. 2003. Planktonic diatom (Bacillariophyceae) fl ora b, 1985a, b, 1986a, b, 1987b; Cirik et al. 1991; Temel of Sultan Sazlıgı Marshes (Kayseri). Turkish Journal of 1992, 1996, 1997a; Yıldız et al. 1994; Demirsoy 1996; Botany 27: 285–301. Morkoyunlu et al. 1996; Mitamura et al. 1997; Akköz AKBULUT A. 2010. The diatom composition of the Salt Lake & Obalı 1998; Elmacı & Obalı 1998), Küçük Menderes Basin and its relationship with salinity. Ekoloji 74: 150– (Skuja 1937; Kolaylı & Güner 1988; Cirik & Cirik 1989; 159. Aysel et al. 1992, 1998a, b, 2001, 2002; Balık & Gez- AKBULUT A. & YILDIZ K. 2002. The phytoplanktonic dia- erler 1995; Erener et al. 1995; Gezerler 1992; Gezerler toms of Çıldır Lake (Ardahan-Turkey). Turkish Journal et al. 1996), Marmara (Albay & Aykulu 1994a, b; Temel of Botany 26: 55–75. 1994, 1996, 1999, 2002, 2003; Yardımcı & Temel 2000), AKIN M. & AKIN G. 2007. Importance of water, water potentıal Sakarya (Aykulu et al. 1983; Gönülol & Aykulu 1984; in Turkey, water basins and water pollution. Ankara Obalı 1984; Ünal 1984; Gönülol 1985a, b, 1987; Yıldız Üniversitesi Dil ve Tarih-Coğrafya Fakültesi Dergisi 47(2): 1987a, b; Dere 1989; Obalı et al. 1989; Yıldız & Özkıran 105–118 (in Turkish with English summary). 1994; Atıcı & Yıldız 1996; Demirsoy 1996; Aykulu et al. AKKÖZ C. & OBALI O. 1998. The diatoms of Besgöz Lake 1999; Yardımcı & Temel 2000; Atıcı 2002), Susurluk (Sarayönü, Konya). In: XIV Ulusal Biyoloji Kongresi, (Skuja 1937; Gezerler 1992; Gezerler et al. 1994; Dere pp. 282–291. Ondokuz Mayıs Üniversitesi, Samsun (in et al. 2002; Karacaoğlu et al. 2004), West Black Sea Turkish). River Basin (Öztürk 1994; Cirik 1997), Yeşilırmak (Yazıcı AKSIN M., ÇETIN K. & YILDIRIM V. 1999. The algae of & Gönülol 1994; Altuner & Pabucçu 1996; Kır & Polat Keban Stream (Elazığ-Turkey). Fırat Üniversitesi Fen ve 1996; Gönülol & Obalı 1998). Mühendislik Bilimleri Dergisi 11(1): 59–65 (in Turkish with English summary). GENERAL DISTRIBUTION AND ECOLOGY. Tycho- ALBAY M. & AYKULU G. 1994a. Algal features of Göksu planktonic and meso-polysaprobous species (Denys stream (İstanbul), I. Planktonic algae. In: XII Ulusal Biy- 1991; Hofmann 1994; Van Dam et al. 1994), clas- oloji Kongresi, pp. 157–165. Trakya Üniversitesi, Edirne sifi ed as an alkaliphilous diatom (Håkansson 1993; (in Turkish). Van Dam et al. 1994). ALBAY M. & AYKULU G. 1994b. Algal features of Göksu Stream (İstanbul), II. Epipelic algae. İstanbul Üniversitesi ACKNOWLEDGEMENTS. We are grateful to Dr. Sofia Su Ürünleri Dergisi 8: 119–128 (in Turkish).

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Received 11 July 2011