Dinosaurs, Pollen, and the Cretaceous-Tertiary Boundary in the San Juan Basin Robert M
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A Neoceratopsian Dinosaur from the Early Cretaceous of Mongolia And
ARTICLE https://doi.org/10.1038/s42003-020-01222-7 OPEN A neoceratopsian dinosaur from the early Cretaceous of Mongolia and the early evolution of ceratopsia ✉ Congyu Yu 1 , Albert Prieto-Marquez2, Tsogtbaatar Chinzorig 3,4, Zorigt Badamkhatan4,5 & Mark Norell1 1234567890():,; Ceratopsia is a diverse dinosaur clade from the Middle Jurassic to Late Cretaceous with early diversification in East Asia. However, the phylogeny of basal ceratopsians remains unclear. Here we report a new basal neoceratopsian dinosaur Beg tse based on a partial skull from Baruunbayan, Ömnögovi aimag, Mongolia. Beg is diagnosed by a unique combination of primitive and derived characters including a primitively deep premaxilla with four pre- maxillary teeth, a trapezoidal antorbital fossa with a poorly delineated anterior margin, very short dentary with an expanded and shallow groove on lateral surface, the derived presence of a robust jugal having a foramen on its anteromedial surface, and five equally spaced tubercles on the lateral ridge of the surangular. This is to our knowledge the earliest known occurrence of basal neoceratopsian in Mongolia, where this group was previously only known from Late Cretaceous strata. Phylogenetic analysis indicates that it is sister to all other neoceratopsian dinosaurs. 1 Division of Vertebrate Paleontology, American Museum of Natural History, New York 10024, USA. 2 Institut Català de Paleontologia Miquel Crusafont, ICTA-ICP, Edifici Z, c/de les Columnes s/n Campus de la Universitat Autònoma de Barcelona, 08193 Cerdanyola del Vallès Sabadell, Barcelona, Spain. 3 Department of Biological Sciences, North Carolina State University, Raleigh, NC 27695, USA. 4 Institute of Paleontology, Mongolian Academy of Sciences, ✉ Ulaanbaatar 15160, Mongolia. -
What Can Be Learned from a Fossil Tooth? an Inquiry Into the Life & Diet of a Small Theropod Dinosaur
What Can Be Learned From A Fossil Tooth? An Inquiry Into The Life & Diet Of A Small Theropod Dinosaur Karolyn Hamlin The Hell Creek and Lance Formations of the animals alive at the time, paleontologists Montana, North Dakota, South Dakota, and strive to understand why the K-Pg extinction Wyoming (Fig. 1) are valuable sources of seemed to affect dinosaurs so heavily Maastrichtian age (Late Cretaceous) fossils. compared to other organisms. Diverse The Maastrichtian is the last age before the vertebrate remains have been found at these well-known Cretaceous-Paleogene (K-Pg) sites, including fish, sharks, amphibians, extinction that marked the demise of the non- mammals, reptiles, and birds. The phrase avian dinosaurs. Studying the fossils of these “non-avian dinosaurs” hints that birds are formations is not only key to understanding considered dinosaurs, the avian dinosaurs, and the effects the K-Pg extinction had on this paper will focus on the group that biodiversity, but understand the biodiversity includes birds, their close ancestors, and near trends leading up to it. Through understanding cousins: the theropods. these trends as well as the characteristics of . Figure 1: Map of the Hell Creek and Lance Formations. (Modified from Johnson, 1992) 53 Figure 2: Recreation of the Late Cretaceous Western Interior Seaway. The Hell Creek and Lance Formations cover areas that were very near water, making fish available as prey animals. (Blakey, 2006) If you have ever been to a museum with a mouths that flat teeth for grinding are useful dinosaur exhibit, you may have seen a for consuming fibrous plant material, and recreation of a large theropod such as sharp teeth for puncturing or shearing are Tyrannosaurus rex or Allosaurus. -
Theropod Teeth from the Upper Maastrichtian Hell Creek Formation “Sue” Quarry: New Morphotypes and Faunal Comparisons
Theropod teeth from the upper Maastrichtian Hell Creek Formation “Sue” Quarry: New morphotypes and faunal comparisons TERRY A. GATES, LINDSAY E. ZANNO, and PETER J. MAKOVICKY Gates, T.A., Zanno, L.E., and Makovicky, P.J. 2015. Theropod teeth from the upper Maastrichtian Hell Creek Formation “Sue” Quarry: New morphotypes and faunal comparisons. Acta Palaeontologica Polonica 60 (1): 131–139. Isolated teeth from vertebrate microfossil localities often provide unique information on the biodiversity of ancient ecosystems that might otherwise remain unrecognized. Microfossil sampling is a particularly valuable tool for doc- umenting taxa that are poorly represented in macrofossil surveys due to small body size, fragile skeletal structure, or relatively low ecosystem abundance. Because biodiversity patterns in the late Maastrichtian of North American are the primary data for a broad array of studies regarding non-avian dinosaur extinction in the terminal Cretaceous, intensive sampling on multiple scales is critical to understanding the nature of this event. We address theropod biodiversity in the Maastrichtian by examining teeth collected from the Hell Creek Formation locality that yielded FMNH PR 2081 (the Tyrannosaurus rex specimen “Sue”). Eight morphotypes (three previously undocumented) are identified in the sample, representing Tyrannosauridae, Dromaeosauridae, Troodontidae, and Avialae. Noticeably absent are teeth attributed to the morphotypes Richardoestesia and Paronychodon. Morphometric comparison to dromaeosaurid teeth from multiple Hell Creek and Lance formations microsites reveals two unique dromaeosaurid morphotypes bearing finer distal denticles than present on teeth of similar size, and also differences in crown shape in at least one of these. These findings suggest more dromaeosaurid taxa, and a higher Maastrichtian biodiversity, than previously appreciated. -
KENNETH CARPENTER, Ph.D. Director and Curator Of
KENNETH CARPENTER, Ph.D. Director and Curator of Paleontology Prehistoric Museum Utah State University - College of Eastern Utah 155 East Main Street Price, Utah 84501 Education May, 1996. Ph.D., Geology University of Colorado, Boulder, CO. Dissertation “Sharon Springs Member, Pierre Shale (Lower Campanian) depositional environment and origin of it' s Vertebrate fauna, with a review of North American plesiosaurs” 251 p. May, 1980. B.S. in Geology, University of Colorado, Boulder, CO. Aug-Dec. 1977 Apprenticeship, Smithsonian Inst., Washington DC Professional Museum Experience 1975 – 1980: University of Colorado Museum, Boulder, CO. 1983 – 1984: Mississippi Museum of Natural History, Jackson, MS. 1984 – 1986: Academy of Natural Sciences of Philadelphia, Philadelphia. 1986: Carnegie Museum of Natural History, Pittsburgh, PA. 1986: Oklahoma Museum of Natural History, Norman, OK. 1987 – 1989: Museum of the Rockies, Bozeman, MT. 1989 – 1996: Chief Preparator, Denver Museum of Nature and Science, Denver, CO. 1996 – 2010: Chief Preparator, and Curator of Vertebrate Paleontology, Denver Museum of Nature and Science, Denver, CO. 2006 – 2007; 2008-2009: Acting Department Head, Chief Preparator, and Curator of Vertebrate Paleontology, Denver Museum of Nature and Science, Denver, CO. 2010 – present: Director, Prehistoric Museum, Price, UT 2010 – present: Associate Vice Chancellor, Utah State University Professional Services: 1991 – 1998: Science Advisor, Garden Park Paleontological Society 1994: Senior Organizer, Symposium "The Upper Jurassic Morrison Formation: An Interdisciplinary Study" 1996: Scientific Consultant Walking With Dinosaurs , BBC, England 2000: Scientific Consultant Ballad of Big Al , BBC, England 2000 – 2003: Associate Editor, Journal of Vertebrate Paleontology 2001 – 2003: Associate Editor, Earth Sciences History journal 2003 – present: Scientific Advisor, HAN Project 21 Dinosaur Expos, Tokyo, Japan. -
A Phylogenetic Analysis of the Basal Ornithischia (Reptilia, Dinosauria)
A PHYLOGENETIC ANALYSIS OF THE BASAL ORNITHISCHIA (REPTILIA, DINOSAURIA) Marc Richard Spencer A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements of the degree of MASTER OF SCIENCE December 2007 Committee: Margaret M. Yacobucci, Advisor Don C. Steinker Daniel M. Pavuk © 2007 Marc Richard Spencer All Rights Reserved iii ABSTRACT Margaret M. Yacobucci, Advisor The placement of Lesothosaurus diagnosticus and the Heterodontosauridae within the Ornithischia has been problematic. Historically, Lesothosaurus has been regarded as a basal ornithischian dinosaur, the sister taxon to the Genasauria. Recent phylogenetic analyses, however, have placed Lesothosaurus as a more derived ornithischian within the Genasauria. The Fabrosauridae, of which Lesothosaurus was considered a member, has never been phylogenetically corroborated and has been considered a paraphyletic assemblage. Prior to recent phylogenetic analyses, the problematic Heterodontosauridae was placed within the Ornithopoda as the sister taxon to the Euornithopoda. The heterodontosaurids have also been considered as the basal member of the Cerapoda (Ornithopoda + Marginocephalia), the sister taxon to the Marginocephalia, and as the sister taxon to the Genasauria. To reevaluate the placement of these taxa, along with other basal ornithischians and more derived subclades, a phylogenetic analysis of 19 taxonomic units, including two outgroup taxa, was performed. Analysis of 97 characters and their associated character states culled, modified, and/or rescored from published literature based on published descriptions, produced four most parsimonious trees. Consistency and retention indices were calculated and a bootstrap analysis was performed to determine the relative support for the resultant phylogeny. The Ornithischia was recovered with Pisanosaurus as its basalmost member. -
The Geology of New Mexico As Understood in 1912: an Essay for the Centennial of New Mexico Statehood Part 2 Barry S
Celebrating New Mexico's Centennial The geology of New Mexico as understood in 1912: an essay for the centennial of New Mexico statehood Part 2 Barry S. Kues, Department of Earth and Planetary Sciences, University of New Mexico, Albuquerque, New Mexico, [email protected] Introduction he first part of this contribution, presented in the February Here I first discuss contemporary ideas on two fundamental areas 2012 issue of New Mexico Geology, laid the groundwork for an of geologic thought—the accurate dating of rocks and the move- exploration of what geologists knew or surmised about the ment of continents through time—that were at the beginning of Tgeology of New Mexico as the territory transitioned into statehood paradigm shifts around 1912. Then I explore research trends and in 1912. Part 1 included an overview of the demographic, economic, the developing state of knowledge in stratigraphy and paleontol- social, cultural, and technological attributes of New Mexico and its ogy, two disciplines of geology that were essential in understand- people a century ago, and a discussion of important individuals, ing New Mexico’s rock record (some 84% of New Mexico’s surface institutions, and areas and methods of research—the geologic envi- area is covered by sediments or sedimentary rocks) and which were ronment, so to speak—that existed in the new state at that time. advancing rapidly through the first decade of the 20th century. The geologic time scale and age of rocks The geologic time scale familiar to geologists working in New The USGS did not adopt the Paleocene as the earliest epoch of the Mexico in 1912 was not greatly different from that used by modern Cenozoic until 1939. -
Implications for Predatory Dinosaur Macroecology and Ontogeny in Later Late Cretaceous Asiamerica
Canadian Journal of Earth Sciences Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous Asiamerica Journal: Canadian Journal of Earth Sciences Manuscript ID cjes-2020-0174.R1 Manuscript Type: Article Date Submitted by the 04-Jan-2021 Author: Complete List of Authors: Holtz, Thomas; University of Maryland at College Park, Department of Geology; NationalDraft Museum of Natural History, Department of Geology Keyword: Dinosaur, Ontogeny, Theropod, Paleocology, Mesozoic, Tyrannosauridae Is the invited manuscript for consideration in a Special Tribute to Dale Russell Issue? : © The Author(s) or their Institution(s) Page 1 of 91 Canadian Journal of Earth Sciences 1 Theropod Guild Structure and the Tyrannosaurid Niche Assimilation Hypothesis: 2 Implications for Predatory Dinosaur Macroecology and Ontogeny in later Late Cretaceous 3 Asiamerica 4 5 6 Thomas R. Holtz, Jr. 7 8 Department of Geology, University of Maryland, College Park, MD 20742 USA 9 Department of Paleobiology, National Museum of Natural History, Washington, DC 20013 USA 10 Email address: [email protected] 11 ORCID: 0000-0002-2906-4900 Draft 12 13 Thomas R. Holtz, Jr. 14 Department of Geology 15 8000 Regents Drive 16 University of Maryland 17 College Park, MD 20742 18 USA 19 Phone: 1-301-405-4084 20 Fax: 1-301-314-9661 21 Email address: [email protected] 22 23 1 © The Author(s) or their Institution(s) Canadian Journal of Earth Sciences Page 2 of 91 24 ABSTRACT 25 Well-sampled dinosaur communities from the Jurassic through the early Late Cretaceous show 26 greater taxonomic diversity among larger (>50kg) theropod taxa than communities of the 27 Campano-Maastrichtian, particularly to those of eastern/central Asia and Laramidia. -
A Revision of the Ceratopsia Or Horned Dinosaurs
MEMOIRS OF THE PEABODY MUSEUM OF NATURAL HISTORY VOLUME III, 1 A.R1 A REVISION orf tneth< CERATOPSIA OR HORNED DINOSAURS BY RICHARD SWANN LULL STERLING PROFESSOR OF PALEONTOLOGY AND DIRECTOR OF PEABODY MUSEUM, YALE UNIVERSITY LVXET NEW HAVEN, CONN. *933 MEMOIRS OF THE PEABODY MUSEUM OF NATURAL HISTORY YALE UNIVERSITY Volume I. Odontornithes: A Monograph on the Extinct Toothed Birds of North America. By Othniel Charles Marsh. Pp. i-ix, 1-201, pis. 1-34, text figs. 1-40. 1880. To be obtained from the Peabody Museum. Price $3. Volume II. Part 1. Brachiospongidae : A Memoir on a Group of Silurian Sponges. By Charles Emerson Beecher. Pp. 1-28, pis. 1-6, text figs. 1-4. 1889. To be obtained from the Peabody Museum. Price $1. Volume III. Part 1. American Mesozoic Mammalia. By George Gaylord Simp- son. Pp. i-xvi, 1-171, pis. 1-32, text figs. 1-62. 1929. To be obtained from the Yale University Press, New Haven, Conn. Price $5. Part 2. A Remarkable Ground Sloth. By Richard Swann Lull. Pp. i-x, 1-20, pis. 1-9, text figs. 1-3. 1929. To be obtained from the Yale University Press, New Haven, Conn. Price $1. Part 3. A Revision of the Ceratopsia or Horned Dinosaurs. By Richard Swann Lull. Pp. i-xii, 1-175, pis. I-XVII, text figs. 1-42. 1933. To be obtained from the Peabody Museum. Price $5 (bound in cloth), $4 (bound in paper). Part 4. The Merycoidodontidae, an Extinct Group of Ruminant Mammals. By Malcolm Rutherford Thorpe. In preparation. -
Mesozoic Stratigraphy at Durango, Colorado
160 New Mexico Geological Society, 56th Field Conference Guidebook, Geology of the Chama Basin, 2005, p. 160-169. LUCAS AND HECKERT MESOZOIC STRATIGRAPHY AT DURANGO, COLORADO SPENCER G. LUCAS AND ANDREW B. HECKERT New Mexico Museum of Natural History and Science, 1801 Mountain Rd. NW, Albuquerque, NM 87104 ABSTRACT.—A nearly 3-km-thick section of Mesozoic sedimentary rocks is exposed at Durango, Colorado. This section con- sists of Upper Triassic, Middle-Upper Jurassic and Cretaceous strata that well record the geological history of southwestern Colorado during much of the Mesozoic. At Durango, Upper Triassic strata of the Chinle Group are ~ 300 m of red beds deposited in mostly fluvial paleoenvironments. Overlying Middle-Upper Jurassic strata of the San Rafael Group are ~ 300 m thick and consist of eolian sandstone, salina limestone and siltstone/sandstone deposited on an arid coastal plain. The Upper Jurassic Morrison Formation is ~ 187 m thick and consists of sandstone and mudstone deposited in fluvial environments. The only Lower Cretaceous strata at Durango are fluvial sandstone and conglomerate of the Burro Canyon Formation. Most of the overlying Upper Cretaceous section (Dakota, Mancos, Mesaverde, Lewis, Fruitland and Kirtland units) represents deposition in and along the western margin of the Western Interior seaway during Cenomanian-Campanian time. Volcaniclastic strata of the overlying McDermott Formation are the youngest Mesozoic strata at Durango. INTRODUCTION Durango, Colorado, sits in the Animas River Valley on the northern flank of the San Juan Basin and in the southern foothills of the San Juan and La Plata Mountains. Beginning at the northern end of the city, and extending to the southern end of town (from north of Animas City Mountain to just south of Smelter Moun- tain), the Animas River cuts in an essentially downdip direction through a homoclinal Mesozoic section of sedimentary rocks about 3 km thick (Figs. -
Recent Advances on Study of Hadrosaurid Dinosaurs in Heilongjiang ( Amur) River Area Between China and Russia
Global Geology,14( 3) ∶ 160-191( 2011) doi: 10. 3969 /j. issn. 1673-9736. 2011. 03. 03 Article ID: 1673-9736( 2011) 03-0160-32 Recent advances on study of hadrosaurid dinosaurs in Heilongjiang ( Amur) River area between China and Russia Pascal Godefroit1 ,Pascaline Lauters1 ,Jimmy Van Itterbeeck2 , Yuri L. Bolotsky3 ,DONG Zhiming4 ,JIN Liyong5 ,WU Wenhao6 , Ivan Y. Bolotsky3,6 ,HAI Shulin7 and YU Tingxiang7 1. Royal Belgian Institute of Natural Sciences,Department of Palaeontology,B - 1 000 Bruxelles,Belgium; 2. Afdeling Historische Geologie,Katholieke Universiteit Leuven,B - 3000 Leuven,Belgium; 3. Amur Natural History Museum,Institute of Geology and Nature Exploration FEB RAS,675000 Blagoveschensk, Amurskaya Oblast',Russia; 4. IVPP,Beijing 100044 / Research Center of Paleontology and Stratigraphy,Jilin University,Changchun 130026,China; 5. Museum of Jilin University,Changchun 130026,China; 6. Research Center for Paleontology and Stratigraphy,Jilin University,Changchun 130061,China; 7. The Geological Museum of Heilongjiang,Harbin 150036,China Abstract: Four main dinosaur-bearing sites have been investigated in latest Cretaceous deposits from the Amur / Heilongjiang Region: Jiayin and Wulaga in China ( Yuliangze Formation) ,Blagoveschensk and Kundur in Rus- sia ( Udurchukan Formation) . More than 90% of the bones discovered in these localities belong to hollow-cres- ted lambeosaurine hadrosaurids: Charonosaurus jiayinensis at Jiayin,Amurosaurus riabinini at Blagoveschensk, Olorotitan arharensis at Kundur,and Sahaliyania elunchunorum at -
Dinosaur Eggshells from the Lower Maastrichtian St. Mary River Formation of Southern Alberta, Canada
Canadian Journal of Earth Sciences Dinosaur eggshells from the lower Maastrichtian St. Mary River Formation of southern Alberta, Canada Journal: Canadian Journal of Earth Sciences Manuscript ID cjes-2017-0195.R1 Manuscript Type: Article Date Submitted by the Author: 13-Nov-2017 Complete List of Authors: Voris, Jared; University of Calgary, Geoscience; Zelenitsky, Darla; Department of Geoscience, Tanaka, Kohei; Nagoya Daigaku Hakubutsukan; University of Calgary, DepartmentDraft of Geoscience Therrien, François; Royal Tyrrell Museum of Palaeontology, Is the invited manuscript for consideration in a Special N/A Issue? : Keyword: eggshell, dinosaur, Cretaceous, Maastrichtian, Alberta https://mc06.manuscriptcentral.com/cjes-pubs Page 1 of 47 Canadian Journal of Earth Sciences 1 2 3 4 5 6 7 8 9 Dinosaur eggshells from the lower Maastrichtian St. Mary River Formation of southern 10 Alberta, Canada 11 12 Jared T. Voris, Darla K. Zelenitsky,Draft François Therrien, Kohei Tanaka 13 J. T. Voris, D. K. Zelenitsky, and K. Tanaka. Department of Geoscience, University of 14 Calgary, 2500 University Dr. NW, Calgary, AB T2N 1N4, Canada; [email protected], 15 [email protected], [email protected] 16 K. Tanaka. Nagoya University Museum, Nagoya University Furocho, Chikusa-Ku, Nagoya, 17 464-8601, Japan; [email protected] 18 F. Therrien. Royal Tyrrell Museum of Palaeontology, Box 7500, Drumheller, AB T0J 0Y0, 19 Canada.; [email protected] 20 1 https://mc06.manuscriptcentral.com/cjes-pubs Canadian Journal of Earth Sciences Page 2 of 47 1 2 Abstract–North America is known for its rich uppermost Cretaceous record of dinosaur egg 3 remains, although a notable fossil gap exists during the lower Maastrichtian. -
Vertebrate Paleontology of the Cretaceous/Tertiary Transition of Big Bend National Park, Texas (Lancian, Puercan, Mammalia, Dinosauria, Paleomagnetism)
Louisiana State University LSU Digital Commons LSU Historical Dissertations and Theses Graduate School 1986 Vertebrate Paleontology of the Cretaceous/Tertiary Transition of Big Bend National Park, Texas (Lancian, Puercan, Mammalia, Dinosauria, Paleomagnetism). Barbara R. Standhardt Louisiana State University and Agricultural & Mechanical College Follow this and additional works at: https://digitalcommons.lsu.edu/gradschool_disstheses Recommended Citation Standhardt, Barbara R., "Vertebrate Paleontology of the Cretaceous/Tertiary Transition of Big Bend National Park, Texas (Lancian, Puercan, Mammalia, Dinosauria, Paleomagnetism)." (1986). LSU Historical Dissertations and Theses. 4209. https://digitalcommons.lsu.edu/gradschool_disstheses/4209 This Dissertation is brought to you for free and open access by the Graduate School at LSU Digital Commons. It has been accepted for inclusion in LSU Historical Dissertations and Theses by an authorized administrator of LSU Digital Commons. For more information, please contact [email protected]. INFORMATION TO USERS This reproduction was made from a copy of a manuscript sent to us for publication and microfilming. While the most advanced technology has been used to pho tograph and reproduce this manuscript, the quality of the reproduction is heavily dependent upon the quality of the material submitted. Pages in any manuscript may have indistinct print. In all cases the best available copy has been filmed. The following explanation of techniques is provided to help clarify notations which may appear on this reproduction. 1. Manuscripts may not always be complete. When it is not possible to obtain missing pages, a note appears to indicate this. 2. When copyrighted materials are removed from the manuscript, a note ap pears to indicate this. 3.