68 PACIFIC SCIENCE, Vol. V, January, .1951

Form 3. Leaves pubescent; flower parts 6--7; Specimensexamined cells of drupe 7-9. LAN AI: Puuwaawaa, Rock 3800 (BISH, NY). In Koa dry forest, Lanai, Munro 61 (BISH , several respects this form is intermediate be­ NY); plateau on lee side ofisland, alt. 600 m. tween vat. Fauriei and ssp. St.Johnii. (?), Wilder 90 (BISH ); xerophytic forest near Kanepuu, Fosberg 12551 (BISH); K anepuu, 5. M yoporum sandwicense ssp. sandwi­ K aa, alt. 1,600 ft., St. j ohn, Eames, & Hosaka cense vat. lanaiense Webster, vat. nov. 18803 (BISH) ; same locality, alt. 1,700 fr., St. j ohn & Cowan 22624 (BISH) ; flats above . PIs. II, 21, 33; III, 43 head of Hawaiilanu i Gulch, alt. 500 m., Fos­ Drupa depressi-conica; endocarpiurn non berg 12525 . (BISH); Poomai J =Paoma?], costatum, Munro (BISH); Mahana Valley, Munro 171Y2 Leaves lanceolate, acuminate, entire, gla­ (BISH); M ahana forehills, Rock 8119 (BISH , brous, 6-13.5 em. long, 0.8-1.8 em. wide. GH , W); mountains near Koele, Forbes 91.L Flowers 2- 4 per axil; pedicels 0.7-1 em. long. (BISH) . Calyx lobes 5, 2-3 mm . long, ovate-acumin­ 6. Myoporum sandwicense ssp. sandwi­ ate. Corolla 5-10beq, 5-8 mm. long, glabrous cense vat. stellatum Web ster, vat. nov. or pubescent, the tube 2.5-4 mm . long. Ovary PIs. I, 1; II, 20, 32; III, 42 about 2 .mm. long ; style 1.8- 3 mm. long. Drupe prob ably whitish in color; endocarp Prinastrum cauliflorum Nutt. ex Gray, Am. depressed-conic or hem ispheric in outline, Acad. Sci. 6: 52. 1866 (herbarium name pub­ lished in synonymy). not or scarcely ridged , 3-4.5 mm . high , 6--9 mm. broad, 5- 7-celled. Folia hirtella, ramuli hirtelli; pili ramosi. Branchlets and leaves pubescent with TYPE: Mahana forehills , Lanai, July 1910, branched hairs, somewhat glabrescent with Rock 8119, in the Bishop Museum Herbarium . age ; leaves elliptic-lanceolate, usually entire, Isotypes are in the Gray Herbarium and 4-9 em. long, 0.7- 2.2 em. broad. Calyx mostly Naturhistorisches Museum , Vienna. 5-10bed, pubescent, sepals 1.2- 2.5 mm. long. This variety, named for the island on which Corolla 5-10bed (rarely 6-10bed), 5-7 mm. it occu rs, is apparently now restricted to the long, glabrous or pubescent within , the dry forest near Kanepuu at the northwest end glandular dots rather inconspicuo us. Stamens of th e island . Ordinarily it is easily distin- mostly 4 or 5, if 5 then often one reduced. .guished from vat. sandwicense by its strongly Ovary 2-2.5 mm. long ; style 2-2.8 mm.long. depressed endocarp. Rarely, plants of vat. Drupe creamy-white; endocarp sub globose, sandwicense, such as Webster 1081 from Oahu ridged, 3- 5 mm. in diameter, 5- 7-celled. and Forbes 477.M from Maui, may have de­ T YPE: Oahu, Ewa coral plain near Barbers pressed end ocarps which appear superficially Point, alt. 10 ft., Webster 1243, in the Bishop similar, but these are easily distinguished in Museum Herbarium (isotype in the Univer­ cross section by the ir thinner walls (compare sity of Texas Herbarium ). Figs. 39 and 43). This distinctive variety is now restricted to . The status of vat. sandwicense on Lanai ·is the low plain of emergent coral at the south­ rather uncertain.Judging from the few collec­ western corner of Oahu . It is known from tions at hand, it seems to occupy a lower alti­ Kauai on the basis of a single collection by tudinal zone than var. lanaiense, but further Nuttall, but has not been collected on that collecting will be necessary before its distri­ island in the century since then. If Nuttall's butional relationship to var. lanaiense will be­ rather questionable locality was correct, the come clear. variety has evidently become extinct on PolynesianSpecies of M yOpOf Um-WEBSTER 69

Kauai . On Oahu it originally grew from the Branchlets glabrous. Leaves mostly ellip­ mouth of Pearl Harbor to Barbers Point, but tic, less commonly lanceolate, pubescent, the it has been exterminated over much of this hairs unbranched, entire or serrate, 4- 9 em. range by the encroachment of military es­ long, 1-2.5 em. broad. Flowers 5-6 per axil. tablishments. In the vicinity ofthe lighthouse Calyx lobes 5, glabrous or pubescent, 1.5­ it is still quite abundant, however, and will 3.5 mm. long. Corolla glabrous or pubescent continue to persist unless the vegetation is within, 5-7.5mm. long. Stamens 5, rarely 4 entirely removed . Ecologically, it must be or 6. Ovary 2-3 .5 mm. long; style 2-5 mm. highly specialized, as it has become adapted long. Drupe color unknown, presumably to a calcareous substrate quite different from white; endocarp 5-9 mm. long, 5-7-celled. the lateritic or basaltic soils on which the TYPE : East Maui, north mauka of Ulupala­ other varieties grow. kua, Degener 12183, in the Bishop Museum Although this variety superficially resem­ Herbarium. bles var. Degeneri in the pubescent leaves, it This variety occurs on the dry leeward has been independently derived, probably slopes of Haleakala, East Maui, from near from Oahu Form 5 of var. sandwicense. This Ulupalakua to the Kaupo Gap; a form from relationship is especially evident in the fact Molokai is referred here for convenience. It that var. stellatum shows the tendency toward is named for Mr . Otto Degener, long a stu­ the loss of the fifth stamen which is fully ex­ dent of the Hawaiian naio, who illustrated it pressed in the form from the Waianae Range. in his book, Plants ofHawaii N ational Park. The name of this variety alludes to the ap­ Specimens examined pearance of the branched hairs, a character MOLOKAI: unique in the genus as far as I can determine. Probably near Kaluaaha, Degener 9738 Specimens examined (DEG, NY). KAUAI: EAST MAUl: Atooi [=Kauai], Nuttall (K) . North mauka ofUlupalakua, Degener 12183 OAHU: (BISH, DEG); Auhi [=Auwahi], Ulupala­ Honouliuli, Barbers Point coral plain, alt. kua, Munro 385 (BISH); Kaapilopilo and Ka­ 10 ft., Cowan 787, 788 (BISH); same locality, pakahawai gulches, Forbes 1940.M(BISH); Degener & Park 9727 (DEG), Egler 37-268 Waiopaa [=Waiopai] Ranch , Forbes 1875.M (BISH), Hosaka 1359 (BISH), Pollock (F. (BISH); Haleakala, Kaupo Gap, Forbes Brown's) 1278 (BISH) , Rock 17038 (BISH, 1105.M (BISH) ; Haleakala, south of Kuiki labeled with an unpublished varietal name), along east side of Kaupo Gap, Degener Setchell (GH) , Webster 1243, 1244, 1250 17600 (NY). (BISH, T), 1245 (BISH) , 1249 (T); between Although the plant from Molokai is placed Barbers Point and Pearl Harbor, along beach, in var. Degeneri for convenience, it probably Degener & Park 9733 (DEG); Ewa, below has arisen independently of the Maui plants Magnetic Observatory toward beach, Egler & and may represent a different variety. How­ Hosaka 37-419 (BISH); Puuloa, 1909, Stokes ever, until the confusing forms from Molokai (BISH) ; east ·side of Pearl Harbor, Stokes are better understood, it seems unwise to de­ (BISH). scribe any new varieties from there. 7. Myoporum sandwicense ssp. sandwi­ 8. Myoporum sandwicense ssp. St.-Johnii cense var. Degeneri Webster, var. nov. Webster, ssp. nov. PI. I, 10-11 PIs. II, 22, 34-35 ; III, 44 Folia elliptica hirtella serrata; pili simplici; Petala et sepala plerumque 6-8; stylus ramuli glabri. hirtellus; drupa plerumque 7-12-locularis.