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Raulerson, L. 2006 Vascular Plant Distribution in the Mariana Islands. X = present; indigenous; O = present, introduced, as of 1995; ? = presence uncertain; + = fossil. Guam Rota Aguihan Tinian Saipan Marpi Anatahan Sarigan Guguan Alamagan Pagan Agrihan Asuncion Maug Uracas PTERIDOPHYTA PSILOTACEAE Psilotum complanatum Sw. X X X Psilotum nudum (L.)Beauv. X X X X X X X LYCOPODIACEAE Lycopodium cernuum L. X X X X X X Lycopodium phlegmaria var. longifolium Spring X X Lycopodium squarrosum Forst.f. X SELAGINELLACEAE Selaginella ciliaris (Retz.) Spring X X X X X Selaginella erythropus O OPHIOGLOSSACEAE Helminthostachys zeylanica (L.) Hook. X Ophioglossum nudicaule L.f. X X X Ophioglossum pendulum L. X X X Ophioglossum reticulatum L. X MARATTIACEAE Angiopteris evecta (Forst.f.) Hoffm. X X X X X X HYMENOPHYLLACEAE Trichomanes brevipes (Presl) Baker X X Trichomanes javanicum var. boryanum (Kunze) X X Fosb. Trichomanes minutum Bl. X X PARKERIACEAE Ceratopteris gaudichaudii Brongn. X OSMUNDACEAE Osmunda japonica Thunb. X X GLEICHENIACEAE Guam Rota Aguihan Tinian Saipan Marpi Anatahan Sarigan Guguan Alamagan Pagan Agrihan Asuncion Maug Uracas Gleichenia linearis (Burm.f.) C.B.Cl. X X X X X SCHIZAEACEAE Lygodium auriculatum (Willd.) Alston X Lygodium microphyllum (Cav.) R.Br. X X X Schizaea dichotoma var. sellingii Fosb. X X CYATHEACEAE Alsophila australis R.Br. O Cyathea aramaganensis Kaneh X X X X X X Cyathea lunulata (Forst.f.) Copeland X DICKSONIACEAE Cibotium chamissoi Kaulf. O POLYPODIACEAE Acrostichum aureum L. X X X X X X Acrostichum speciosum Willd. ? ? Adiantum philippense L. X Adiantum tenerum L. O Adiantum trapeziforme O Antrophyum plantagineum (Cav.) Kaulf. X X X Arachniodes aristata (Forst.f.) Tindale X Asplenium laserpitifolium Lam. X X Asplenium nidus L. X X X X X X X X X X X X Asplenium pellucidum Lam. var. pellucidum X X X Asplenium polyodon Forst.f. X X X X X Asplenium unilaterale Lam. X X X X X X Belvisia spicata (L.f.) Mirb. ex Copel X X X X X Blechnum cartilaginum Sw. X Blechnum orientale L. X X Bolbitis quoyana (Gaud.) Ching in C. Chr. X X X X Cheilanthes tenuifolia (Burm.f.) Sw. X X X X X Ctenitis dissecta (Forst.f.) Copel. X Ctenitis lepigera (Baker) Ito X Ctenitis subglandulosa (Hance) Ching X Ctenitis subtripinnata var. leucostipes (Bak.) Ito X Davallia solida (Forst.f.) Sw. X X X X X X X X X Guam Rota Aguihan Tinian Saipan Marpi Anatahan Sarigan Guguan Alamagan Pagan Agrihan Asuncion Maug Uracas Diplazium proliferum (Lam.) Thouars X Diplazium sorzogonense Presl X Diplazium woodwardioides (Presl) Morton X Diplazium sp. X Drynaria quercifolia (L.) J.Sm. O Heterogonium pinnatum (Copel.) Holtt. X X Histiopteris incisa (Thunb.) J.Sm. X X Humata banksii Alst. X Humata heterophylla (J.E.Sm.) Desv. X X Humata repens (L.f.) Diels X X Lepisorus boninensis (Christ) Ching Lindsaea ensifolia Sw. X X X X X X Lindsaea obtusa J.Sm. X X Lindsaea repens var. lingulata Kramer X X Lunathyrium bonincola (Nakai) H.Ohba X X Lunathyrium petersonii (Kunze) H.Ohba X Microlepia manilensis Presl X Microlepia speluncae (L.) T. Moore X X X Nephrolepis acutifolia (Desv.) Christ X Nephrolepis biserrata (Sw.) Schott var. biserrata X X X X X X X Nephrolepis biserrata var. furcans Hort. O Nephrolepis cordifolia var. tuberosa Bak. O Nephrolepis exaltata (L.) Schott O Nephrolepis hirsutula (Forst.f.) Presl X X X X X X X X X X X X Nephrolepis multiflora (Roxb.) Jarrett ex Morton X X X X X Pityrogramma calomelanos (L.) Link O O O O Platycerium grande (Fee) J. Sm. ex Presl O Polypodium alternifolium Willd. X Polypodium cyathoides Sw. X X Polypodium punctatum (L.) Sw. X X X X X Polypodium scolopendria Burm.f. X X X X X X X X X X X X Pteris boninensis H.Ohba X X X X Pteris ensiformis Burm.f. X Pteris ensiformis var. victoriae O Pteris excelsa Gaud. X Pteris fauriei Hieron. ? X X X X X Pteris quadriaurita Retz. s.l. X X X X X X X X X Pteris tripartita Sw. X X X X Guam Rota Aguihan Tinian Saipan Marpi Anatahan Sarigan Guguan Alamagan Pagan Agrihan Asuncion Maug Uracas Pteris vittata L. O O O O O Pyrrosia lanceolata (L.) Farw. X X X X Sphenomeris chinensis ssp. biflora (Kaulf.) Jotani X X X X X X X X X X &Ohba Stenosemia aurita (Sw.) Presl X Tectaria crenata Cav. X X X X Thelypteris dentata (Forsk.) E.St.John X X X Thelypteris gretheri (Wagner) Stone X X Thelypteris guamensis (Holtt.) Fosb. & Sachet X X X X Thelypteris heterocarpa (Bl.) Morton X Thelypteris interrupta (Willd.) Iwats. X Thelypteris maemonensis (Wagner & Grether) X X Stone Thelypteris ogasawarensis (Nakai) H.Ito ex Honda X Thelypteris opulenta (Kaulf.) Fosb. X X X X X X Thelypteris parasitica (L.) Tard. X X X Thelypteris subpubescens (Bl.) Iwats. X X Thelypteris torresiana (Gaud.) Alston X X X X X X X X Thelypteris truncata (Poir.) Moatsuki X X Thelypteris unita (L.) Morton X X X X X X X X Thelypteris warburgii (Kuhn & Christ) Stone X Vittaria incurvata Cav. X X X X X GYMNOSPERMAE CYCADACEAE C.Pers. Cycas L. Cycas circinalis L. X X X O X Cycas revoluta Thunb. O O O ZAMIACEAE Horaninow Zamia L., nom. cons. Zamia furfuracea O ARAUCARIACEAE H.Henkel & W.Hochst. Araucaria Juss. Araucaria columnaris (Forst.f.) Hook. O Araucaria columnaris X heterophylla ? O Araucaria cunninghamii D.Don O Araucaria heterophylla (Salisb.) Franco O Guam Rota Aguihan Tinian Saipan Marpi Anatahan Sarigan Guguan Alamagan Pagan Agrihan Asuncion Maug Uracas PODOCARPACEAE Endl. Podocarpus L’Herit. ex Pers., nom. cons. Podocarpus gracilior Pilger O Podocarpus macrophyllus var. maki Endl. O Podocarpus neriifolius D.Don O PINACEAE J.Lindl. Pinus L. Pinus caribaea var. hondurensis Barret & Golfari O Pinus luchuensis Mayr O Pinus thunbergii Parl. O TAXODIACEAE Warming Cryptomeria D.Don Cryptomeria japonica (L.f.) D. Don O CUPRESSACEAE Bartl. Cupressus L. Cupressus sempervirens L. O Juniperus L. Juniperus chinensis L. O Juniperus virginiana L. O Thuja L. Thuja orientalis L. O O O MONOCOTYLEDONAE PANDANACEAE R.Br. Freycinetia Gaud.-Beaupre Freycinetia reineckei Warb. X X X X X X X X Pandanus S.Parkinson Pandanus amaryllifolius O Pandanus dubius Spreng. var. dubius X X X X Pandanus tectorius Park. X X X X X X X X X X X CYMODOCEACEAE N.Taylor Halodule Endl. Halodule uninervis (Forssk.) Aschers. X X Guam Rota Aguihan Tinian Saipan Marpi Anatahan Sarigan Guguan Alamagan Pagan Agrihan Asuncion Maug Uracas POTAMOGETONACEAE Dumort. Potamogeton L. Potamogeton lucens L. X Potamogeton mariannensis C. & S. X Ruppia L. Ruppia maritima var. pacifica St. John & Fosb. X ALISMATACEAE Vent. Sagittaria L. Sagittaria subulata var. kurziana (Gluck) Bogin O HYDROCHARITACEAE Juss. Enhalus L.Rich. Enhalus acoroides (L.f.) Royle X X X Halophila Du Petit-Thouars Halophila minor (Zoll. ex Miq.) den Hart. X X Hydrilla L.Rich. Hydrilla verticillata (L.f.) Royle O Thalassia Banks & Sol. ex Koenig Thalassia hemprichii (Ehrenb.) Aschers. ? TRIURIDACEAE G.Gardner Sciaphila Blume Sciaphila X TYPHACEAE Juss. Typha L. Typha sp. O POACEAE Barnhart Andropogon L. Andropogon brevifolius var. paradoxus (Buse) X Ohwi Andropogon fragilis R.Br. X Arundinaria Michaux Arundinaria(Pleioblastus distichus (mitord)Nakai) O Arundo L. Guam Rota Aguihan Tinian Saipan Marpi Anatahan Sarigan Guguan Alamagan Pagan Agrihan Asuncion Maug Uracas Arundo donax L. var. donax O Avena L. Avena sativa L. O Axonopus Pal. de Beauv. Axonopus compressus (Sw.) Beauv. O Bambusa Schreber, nom. cons. Bambusa blumeana Schultes f. O O Bambusa multiplex (Lour.) Raeusch. O O Bambusa polymorpha Munro O Bambusa tulda Roxb. O Bambusa vulgaris Schrad. ex Wendl. O O O O O O O Cenchrus L. Cenchrus brownii R.&S. O O O O O O O Cenchrus echinatus L. O O O O O O O Centotheca A.Desv., nom. orth.cons. Centotheca lappacea (L.) Desv. X X X X Chloris Sw. Chloris gayana Kunth O Chloris inflata Link O O O O O O O O O Chloris radiata (L.) Sw. O O O O O Chloris virgata Sw. O O Chrysopogon Trin., nom. cons. Chrysopogon aciculatus (Retz.) Trin. X X X X X X X X X Coix L. Coix lacryma-jobi L. O O O O Cymbopogon Sprengel Cymbopogon citratus (DC. ex Nees) Stapf O O Cymbopogon nardus (L.) Rendle O ? Cynodon L.Rich., nom. cons. Cynodon dactylon (L.) Pers. var. dactylon O O O O O O O O O O Cynodon dactylon var. maritimus (HBK) Hack. O Cynodon dactylon var. parviglumis (Ohwi) Fosb.& X X X X X X Sachet Cyrtococcum Stapf Cyrtococcum oxyphyllum (Hochst. ex Steud.) X Stapf Cyrtococcum patens (L.) Camus X Dactyloctenium Willd. Dactyloctenium aegyptium (L.) Willd. O O O O O O O Guam Rota Aguihan Tinian Saipan Marpi Anatahan Sarigan Guguan Alamagan Pagan Agrihan Asuncion Maug Uracas Dendrocalamus Nees Dendrocalamus strictus (Roxb.) Nees O Dichanthium Willemet Dichanthium bladhii (Retz.) Clayton O O O O O Dichanthium caricosum (L.) A. Camus O O O Digitaria Hallier, nom.cons. Digitaria bicornis (Lam.) R.&S. O O O O O O Digitaria ciliaris (Retz.)Koel. X ? X X ? ? X X X X Digitaria gaudichaudii (Kunth) Buse X X X X X X X X Digitaria insularis (L.) Mez O O O Digitaria mariannensis Merr. X Digitaria mezii Kaneh. X X Digitaria radicosa (J.S. Presl) Miq. X X X X X X X X Digitaria setigera Roth in R.&S. X X X X X X X X Digitaria setigera var. calliblepharata (Henr.)Veldk. X X X Digitaria violascens Link O O Dimeria R.Br. Dimeria chloridiformis (Gaud.) K.Schum. & Laut. X Dimeria ornithopoda var.tenera (Trin.) Hack.
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