Zootaxa: a Review of the Asilid (Diptera) Fauna from Hispaniola

ZOOTAXA

1381

A review of the asilid (Diptera) fauna from Hispaniola with six genera new to the island, fifteen new species, and checklist

AUBREY G. SCARBROUGH & DANIEL E. PEREZ-GELABERT

Magnolia Press Auckland, New Zealand AUBREY G. SCARBROUGH & DANIEL E. PEREZ-GELABERT A review of the asilid (Diptera) fauna from Hispaniola with six genera new to the island, fifteen new species, and checklist (Zootaxa 1381) 91 pp.; 30 cm. 14 Dec. 2006 ISBN 978-1-86977-066-2 (paperback) ISBN 978-1-86977-067-9 (Online edition)

FIRST PUBLISHED IN 2006 BY Magnolia Press P.O. Box 41383 Auckland 1030 New Zealand e-mail: [email protected] http://www.mapress.com/zootaxa/

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ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition) Zootaxa 1381: 1–91 (2006) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA 1381 Copyright © 2006 Magnolia Press ISSN 1175-5334 (online edition)

A review of the asilid (Diptera) fauna from Hispaniola with six genera new to the island, fifteen new species, and checklist

AUBREY G. SCARBROUGH1 & DANIEL E. PEREZ-GELABERT2 1Visiting Scholar, Department of Entomology, University of Arizona, Tucson, AZ 85741. E-mail: [email protected] 2Department of Entomology, National Museum of Natural History, Smithsonian Institution, P. O. Box 37012, Washington, DC 20013-7012. E-mail: [email protected]

Table of contents

Abstract ...... 5 Introduction ...... 5 Methods and materials ...... 6 Systematics ...... 7 Subfamily Apocleinae ...... 7 Efferia Coquillett ...... 7 Efferia fortis (Walker) ...... 7 Efferia fulvibarbis (Macquart) ...... 8 Efferia pachychaeta (Bromley) ...... 8 Efferia stylata (Fabricius) ...... 8 Proctacanthella Bromley ...... 11 Proctacanthella taina sp. nov...... 12 Proctacanthus Macquart ...... 14 Proctacanthus darlingtoni Curran ...... 14 Proctacanthus dominicana Curran ...... 15 Proctacanthus vittatus (Olivier) ...... 15 Wilcoxius Martin ...... 16 Subfamily Laphriinae ...... 17 Tribe Andrenosomatini ...... 17 Andrenosoma Rondani ...... 17 Andrenosoma chalybeum Williston ...... 17 Andrenosoma ruficaudum (Williston) ...... 18 Dasyllis Loew ...... 19 Dasyllis rufa (Bromley) ...... 20 Neophoneus Williston ...... 20

Accepted by N. Evenhuis: 24 Oct. 2006; published: 14 Dec. 2006 3 ZOOTAXA Pilica Curran ...... 21 1381 Pilica sp...... 21 Subfamily Laphriinae ...... 21 Tribe Atomosini ...... 21 Atomosia Macquart ...... 21 Key to the known species of Atomosia from Hispaniola ...... 22 Atomosia anacaona sp. nov...... 22 Atomosia ciguaya sp. nov...... 25 Atomosia jagua sp. nov...... 28 Atomosia jimagua sp. nov...... 31 Atomosia yurabia sp. nov...... 33 Atoniomyia Hermann ...... 35 Atoniomyia mikii (Williston) ...... 36 Atoniomyia sp...... 38 Cerotainia Schiner ...... 39 Cerotainia sola sp. nov...... 39 Eumecosoma Schiner ...... 42 Eumecosoma caerulum sp. nov...... 42 Rhopalogaster Macquart...... 44 Rhopalogaster albidus sp. nov...... 45 Subfamily Ommatiinae ...... 46 Ommatius Wiedemann ...... 46 Ommatius bastardoanus Scarbrough ...... 46 Ommatius cinnamomeus Scarbrough ...... 47 Ommatius elusivus Scarbrough ...... 49 Ommatius geminus sp. nov...... 51 Ommatius gwenae Scarbrough ...... 55 Ommatius hispaniolae Scarbrough ...... 56 Ommatius inflatus Scarbrough ...... 57 Ommatius laticrus sp. nov...... 58 Ommatius lucidatus Scarbrough ...... 62 Ommatius maculosus sp. nov...... 62 Ommatius nigellus Scarbough ...... 66 Ommatius praelongus sp. nov...... 66 Ommatius lucidatus species group ...... 70 Key to the known species of the Ommatius lucidatus species group from Hispaniola ...... 70 Subfamily Stenopogoninae ...... 72 Tribe Stenopogonini ...... 72 Plesiomma Macquart ...... 72 Key to Plesiomma in Hispaniola ...... 73 Plesiomma angustum (Macquart) ...... 73 Plesiomma inflatum Hull ...... 76 Plesiomma lineata (Fabricius) ...... 80 Plesiomma simile sp. nov...... 81 Subfamily Trigonomiminae ...... 84 Tribe Damalini ...... 84 Holcocephala Jaennicke ...... 84

Abstract

The robber fly fauna of the 6 subfamilies Apocleinae, Asilinae, Laphrinae, Ommatiinae, Stenopogoninae, and Trigonomiminae of Hispaniola with special reference to the Dominican Republic is reviewed in light of new collections. This paper reports 6 genera (Cerotainia Schiner, Eumecosoma Schiner, Holcocephala Jaennicke, Pilica Curran, Proctacanthella Bromley, and Rhopalogaster Macquart) new to the island, increasing the number to 20. Within the West Indies, Eumecosoma, Holcocephala, Pilica, and Proctacanthella are only known from Hispaniola. Also, 33 species are reported, including 15 new species (Atomosia anacaona, A. ciguaya, A. jagua, A. jimagua, A. yurabia, Cerotainia sola, Eumecosoma caerulum, Holcocephala indigena, Ommatius geminus, O. laticrus, O. maculosus, O. praelongus, Proctacanthella taina, Plesiomma simile, and Rhopalogaster albidus), increasing the number to 62 species plus 2 species in Dominican amber. The male of O. cinnamomeus Scarbrough & females of Plesiomma inflatum and Proctacanthus darlingtoni Curran are reported for the first time. The Ommatius lucidatus species group is established with 8 extant and 2 fossil species. Plesiomma angustum (Macquart) and Atoniomyia mikii (Williston) are rediscovered and redescribed. Plesiomma lineata (Fabricius) is removed from the distribution list for Cuba and Jamaica and Neophoneus is removed from the list of West Indies asilids because of an error in identification. Neophoneus flavotibis Bigot tentatively diagnosed as belonging to the genus Efferia. Plesiomma macra Loew is removed from synonymy. Significant structures of most species are illustrated and keys to selected species are included. At least one species of Atoniomyia and Pilica remain undescribed from this survey. New distribution records for most species and a checklist of the Hispaniolan fauna are also included.

Key words: Asilidae, new species, distribution records, Hispaniola

Introduction

The Hispaniolan asilid fauna, as well as all of the West Indies, have been sparsely sampled, and are poorly known. Most new discoveries have resulted from systematic surveys conducted over a period of years, such as those in Dominica and Hispaniola. Prior to 1900, Bigot (1875, 1878), Fabricius (1775), Johnson (1894), Macquart (1848), Olivier (1789), Walker (1885), Williston (1885, 1886) described the first 10 Hispaniolan species. Six species were added by Bromley (1928, 1931), Curran (1951), and Hull (1962) during the next 78 years for a total of 16 species. Subsequently, Scarbrough (1984,1997, 2001) and colleagues (Scarbrough & Poinar, 1993; Scarbrough & Perez-Gelabert, 2003, 2005) have added the remaining 46 species, plus 2 in Dominican amber, for a total of 20 genera and 64 species. Of the 165 species recorded in the West Indies, 153 or 86% are endemic species. The

REVIEW OF ASILIDS © 2006 Magnolia Press 5 ZOOTAXA Greater Antilles, with the greater land masses and habitat types, have the vast majority of 1381 species with 47 from Cuba, 62 from Hispaniola, 26 from Jamaica+Cayman Islands, and 24 from Puerto Rico+Virgin Islands. Of these, eleven species, Andrenosoma chalybeum (Williston), Dasyllis cressa Bromley, Apachekolos clavipes (Johnson), Atomosia incisularis Macquart, Efferia stylata (Fabricus), Cerotainia sola sp. nov., Leptogaster bahamiensis Scarbrough, L. obscuripes Loew, Ommatius lineolatus Scarbrough, O. marginellus (Fabricus), and O. setiferous Scarbrough, have been taken from 2 or more islands. Shared species by country are: Cuba with 10, Hispaniola 5, Puerto Rico-Virgin Islands 4, and Jamaica-Cayman Islands one. The Bahamas and Turks and Caicos Islands total 12 species of which 4 are taken from nearby islands or southeastern North America. The Lesser Antilles has 8 species with 2 shared. This study reviews the asilid subfamilies Apocleinae, Asilinae, Laphrinae, Ommatiinae, Stenopogoninae, and Trigonomiminae from Hispaniola with special reference to the Dominican Republic in light of new collections. Six genera new to the island are reported and 33 species are reviewed, 15 of which are new to science. However, several species remain undiscovered or undescribed because males have not been found or the genera are under revision. Furthermore, the asilid fauna is probably only 75–80% complete, especially considering that Haiti has been sparsely sampled with most species being taken in the southern areas.

Methods and materials

Descriptions are composites, based on all available specimens. Morphological terminology follows McAlpine (1981). Terminalia were treated in warm KOH, washed in 70% ETOH and treated with a drop of acetic acid, transferred to glycerin, and subsequently illustrated. Later, terminalia were placed in microvials with glycerin for permanent storage and attached to the specimen pin. Penciled illustrations were scanned and completed using Adobe Photoshop software. Measurements were made using an ocular micrometer in the eyepiece of a Meiji dissecting microscope. Body length was measured from the foremost protrusion of the face to the apex of the terminalia. Wing length was measured from the articulated base to its apex. Data were recorded in a standard format with each line denoted by a forward slash [/] and a second label by an ~ symbol, preceded and followed by a blank space. When these symbols appear on the data label, they are recorded without blank spaces. Spellings of locations on data labels are corrected whenever possible, such as Monte Cristi=Montecristi. Countries appear only at the beginning of a series in bolded, capital letters. Bold letters are also used to designate provinces and types. Brackets [] denote useful additional information not found on labels. Materials for this study may be found in the following institutions. Codons of institutions follow Evenhuis & Samuelson (2006).

6 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT AMNH American Museum of Natural History, New York, USA ZOOTAXA BMNH Natural History Museum, London, England. 1381 CMNH Carnegie Museum of Natural History, Pittsburgh, Pennsylvania, USA. EMF Eric M. Fisher Collection, Sacramento, California, USA. FSCA Florida State Collection of Arthropods, Gainesville, Florida, USA. IIBZ Instituto de Investigaciones Botánicas y Zoológicas de la Universidad Autonoma de Santo Domingo, Santo Dominigo, Dominican Republic. MCZ Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA. USNM National Museum of Natural History, Smithsonian Institution, Washington, DC, USA. MHND Museo Nacional de Historia Natural, Santo Domingo, Dominican Republic. MNHN Museum National d'Histoire Naturelle, Paris, France. MAIC Michael A. Ivie Collection, Montana State University, Bozeman, Montana, USA UMO Oxford Museum of Natural History, Oxford, England. ZMUC Zoological Museum, Copenhagen, Denmark.

Systematics

Subfamily Apocleinae

Efferia Coquillett

Efferia Coquillett, 1893: 175. Type species: candida Coquillett (Coquillett, 1910: 536). Martin & Papavero, 1970: 62. Catalogue. Nerax Hull, 1962: 476. Type species: Asilus aestuans Linneaus (orig. des.). Erax, authors, not Scopoli.

Remarks. This is one of the largest genera in the Americas with 118 named species (Martin & Papavero, 1970), and many more are undescribed. Of the described species, 18 are from the West Indies of which 4 are from Hispaniola. A revision of the West Indian Efferia is under way (Scarbrough & Perez-Gelabert, in prep.) which attempts to establish the distribution, identity and validity of several species. Efferia species inhabit open, sunny areas usually perching on and foraging from bare soil and low ground objects, such as grasses, fallen branches and trees, rocks, and shrubby woody plants.

Efferia fortis (Walker) (Table 1)

Erax fortis Walker, 1855: 623. Type locality: 'Santo Domingo', type: % (BMNH).

REVIEW OF ASILIDS © 2006 Magnolia Press 7 ZOOTAXA Efferia fortis (Walker). Martin & Papavero, 1970: 65. New Comb. 1381 Erax rufitibia Macquart, 1848: 187 (1848: 27). Type locality: 'Haiti ', type: & (UMO). Curran, 1953: 4. Efferia rufitibia (Macquart). Martin & Papavero, 1970: 65. Syn., New Comb. (Macquart 's Haiti specimens).

Remarks. Macquart (1848) described Erax rufitibia from two females from Brazil (Rio Negro) and southern Haiti. Later, Walker (1855) described E. fortis from a male collected from Santo Domingo and separated it from E. rufitibia largely based upon its hyaline wings. Curran (1953) recognized that Macquart 's type series consisted of two species and the difficulty of identifying females, especially based upon Macquart's description. He proposed that the name E. rufitibia be restricted to the Rio Negro specimen, leaving the Haitian specimen without a name. The latter was subsequently synonymized with E. fortis (Martin 1970).

Efferia fulvibarbis (Macquart) (Table 1)

Erax fulvibarbis Macquart, 1848: 188. (1848: 28). Type locality: Haiti, type: &. Efferia fulvibarbis (Macquart). Martin & Papavero, 1970: 65. New Comb.

Remarks. Macquart's type is apparently 'lost', and thus its identity and distribution is problematic.

Efferia pachychaeta (Bromley) (Table 1)

Erax pachychaeta Bromley, 1928: 1. Type locality: Haiti, La Moliniére, type: % (AMNH). Efferia pachychaeta (Bromley). Martin & Papavero, 1970: 66. New Comb.

Remarks. A large, reddish species from southern Haiti.

Efferia stylata (Fabricius) (Table 1)

Asilus stylata Fabricius, 1775: 795. Type locality: St. Croix, U. S. Virgin Islands, type: & (ZMUC). Erax haitiensis Macquart, 1848: 188 (1848: 28) Type locality: Haiti, type: %. Curran, (1931) 9:9. Efferia stylata (Fabricius). Martin & Papavero, 1970: 68. New comb. Distribution, Cuba, Haiti, Puerto Rico, Santo Domingo, U. S. Virgin Islands (St. Croix, St. Thomas).

8 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT TABLE 1. A check list of asilids of Hispaniola (Haiti=H, Dominican Republic=DR). ZOOTAXA 1381 Taxon Species Author, year Country APOCLINAE Efferia Coquillett, 1893 Efferia fortis (Walker, 1855) H Efferia fulvibarbis (Macquart, 1848) H Efferia pachychaeta (Bromley, 1928) H Efferia stylata (Fabricius, 1775) H Proctacanthella Bromley, 1934 Proctacanthella taina Scarbrough & Perez-Gelabert, nov. sp. DR Proctacanthus Macquart, 1838 Proctacanthus darlingtoni Curran, 1951 DR Proctacanthus dominicana Curran, 1951 DR Proctacanthus vittatus (Oliver, 1789) H/DR Wilcoxius Martin, 1975 Wilcoxius planus Scarbrough & Perez-Gelabert, 2005 DR Wilcoxius tumidus Scarbrough & Perez-Gelabert, 2005 DR LAPHRIINAE Andrenosoma Rondani, 1856 Andrenosoma chalybea Williston, 1885 DR Andrenosoma ruficaudum (Williston, 1885) H/DR Dasyllis Loew, 1851 Dasyllis rufa (Bromley, 1931) DR Atomosia Macquart 1838 Atomosia anacaona Scarbrough & Perez-Gelabert, nov. sp. DR Atomosia ciguaya Scarbrough & Perez-Gelabert, nov. sp. DR Atomosia jagua Scarbrough & Perez-Gelabert, nov. sp. DR Atomosia jimagua Scarbrough & Perez-Gelabert, nov. sp. DR Atomosia yurabia Scarbrough & Perez-Gelabert, nov. sp. DR Atoniomyia Hermann, 1912 Atoniomyia mikii (Williston, 1886) DR Atoniomyia sp. DR Cerotainia Schiner, 1866 Cerotainia sola Scarbrough & Perez-Gelabert, nov. sp. DR Eumecosoma Schiner, 1866 Eumecosoma caerulum Scarbrough & Perez-Gelabert, nov. sp. DR to be continued.

REVIEW OF ASILIDS © 2006 Magnolia Press 9 ZOOTAXA TABLE 1 (continued). 1381 Taxon Species Author, year Country Pilica Curran, 1931 Pilica sp. DR Rhopalogaster Macquart, 1834 Rhopalogaster albidus Scarbrough & Perez-Gelabert, nov. sp. DR LEPTOGASTERINAE Apachekolos Martin, 1957 Apachekolos clavipes (Johnson, 1894) DR Apachekolos flaventis Scarbrough & Perez-Gelabert, 2005 DR Apachekolos invasus Scarbrough & Perez-Gelabert, 2005 DR Apachekolos magnus Scarbrough & Perez-Gelabert, 2005 DR Apachekolos volubilis Scarbrough & Perez-Gelabert, 2005 DR Beameromyia Martin, 1957 Beameromyia dicrana Scarbrough, 1997 DR Beameromyia melana Scarbrough, 1997 DR Beameromyia vultus Scarbrough & Perez-Gelabert, 2005 DR Eurhabdus Aldrich, 1923 Eurhabdus sororius Scarbrough & Perez-Gelabert, 2005 DR Leptogaster Meigen, 1803 Leptogaster hyacinthina Scarbrough, 1996 DR Leptogaster lineatus Scarbrough, 1996 DR Psilonyx Aldrich, 1923 Psilonyx zephyrus Scarbrough & Page, 2005 DR OMMATINAE Ommatius Wiedemann, 1821 Ommatius atrosus Scarbrough, 1997 DR Ommatius bastardoanus Scarbrough, 2003 DR Ommatius cinnamomeus Scarbrough, 1997 H/DR Ommatius elusivus Scarbrough, 2003 DR Ommatius fimbrillus† Scarbrough, 1992 DR Ommatius flavescens Scarbrough, 2003 DR Ommatius geminus Scarbrough & Perez-Gelabert, nov. sp. DR Ommatius gwenae Scarbrough, 1984 DR Ommatius haitiensis Scarbrough, 1984 H/DR Ommatius hierroi Scarbrough, 2003 DR Ommatius hispaniolae Scarbrough, 1984 H/DR to be continued.

10 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT TABLE 1 (continued). ZOOTAXA Taxon Species Author, year Country 1381 Ommatius inflatus Scarbrough, 2003 DR Ommatius laticrus Scarbrough & Perez-Gelabert, nov. sp. DR Ommatius lucidatus Scarbrough, 1997 DR Ommatius maculosus Scarbrough & Perez-Gelabert, nov. sp. DR Ommatius nigellus Scarbrough, 1984 H/DR Ommatius praelongus Scarbrough & Perez-Gelabert, nov. sp. DR Ommatius pulverius Scarbrough, 1997 DR Ommatius russelli Scarbrough, 1984 DR Ommatius stramineus Scarbrough, 1984 H/DR Ommatius subtus† Scarbrough, 1992 DR Ommatius vitreus Bigot, 1875 H/DR Ommatius vivus Scarbrough, 1997 DR STENOPOGONINAE Plesiomma Macquart, 1838 H/DR Plesiomma angustum (Macquart, 1848) H/DR Plesiomma inflatum Hull, 1962 H/DR Plesiomma simile Scarbrough & Perez-Gelabert, nov. sp. DR TRIGONOMIMINAE Holcocephala Jaennicke, 1838 Holcocephala indigena Scarbrough & Perez-Gelabert, nov. sp. DR Total Genera 20 Total Species 64

Remarks. Efferia stylata was described in the genus Asilus and later transferred to Efferia. Presently it is reported as widespread in the West Indies. The identity and distribution of the species is problematic since the type, a female, is essentially destroyed. No new records are reported here.

Proctacanthella Bromley

Proctacanthella Bromley, 1934: 96. Type species: Asilus cacopilagus Hine (orig. des.). Hull, 1962: 498; Martin & Papavero, 1970: 82. Catalogue.

Remarks. Proctacanthella is a small genus thus far limited to western United States and northern Mexico (E. Fisher, pers. com. 2006). The 5 species in this genus are small to median size [11–25 mm] flies with a gradually inclined face, moderately produced

1381 (Fisher, 1997). The anatergite is bare, vein R5 meets the wing margin below the apex, and the female ovipositor is round in cross-section and bears a circlet of spines. Superficially, members of the genus are a smaller version of Proctacanthus Macquart [see remarks section under this genus] but are distinguished from the latter by its smaller size, wing venation and terminalia. Here we report the genus from the Dominican Republic for the first time.

Proctacanthella taina sp. nov. (Table 1, Figs. 1–2, 118)

Male. Measurements, body 11.8 mm, wing 9.4 mm. Mostly blackish with whitish vestiture. Head: Whitish tomentose, mostly whitish bristles and setae. Face gradually produced, well exposed beyond margin of eyes; oral margin with several stout bristles medially, thinner bristles laterally, dense white setae dorsally. Palpus and proboscis white setose, former with sparse black setae apically. Antenna black setose; scape, flagellum, and style twice or more as long as pedicel. Frons with setae as long as scape. Ocellar tubercle low, with short, weak whitish setae. Occiput with numerous stout bristles dorsally, bristles thinner, longer laterally. Thorax: Pronotum with abundant, long whitish setae and numerous stout bristles. Scutum mostly yellowish-gray tomentose; 2 narrow longitudinal brown tomentose stripes medially and 2 lateral spots; setae short, bristly mostly black dorsally, narrow anterior and side white setose; 4–5 thin, black dorsocentral bristles present with sparse bristles between rows; lateral bristles stout, whitish, supra-alar bristles black. Scutellum yellowish-gray tomentose with scattered bristles of varying thicknesses and lengths dorsally and apically, one or 2 pairs slightly thicker than others; preapical crease absent. Anatergite bare. Pleura black, gray tomentose with white vestiture. Halter whitish. Wing: Hyaline, veins brown, those basally yellow or brownish-yellow. Costal margin

white setose. Vein R4 without spur vein; R5 joining wing margin below wing apex; spur r4

vein absent. Crossvein r-m just beyond middle of discal cell. Cell m3 petiolate basally. Legs: Mostly black, with whitish setae and bristles. Coxae yellowish-gray tomentose with abundant setae and bristles. Femora black, narrow base yellow; bristles white, usually stout; fore femur with thin, long bristles ventrally. Tibiae with basal 1/3 or 1/2 yellowish, black bristles present apically on all tibiae, anteriorly on fore tibiae, mixed black and whitish on hind tibiae. Tarsi blackish with whitish setae, mixed stout black and white bristles. Abdomen: Mostly blackish, segment 6 apically, segments 7 and 8 entirely reddish or brownish-yellow; yellowish-gray tomentose, with whitish setae and bristles, bristles longer and stouter on tergites apically and laterally; tergites with large brownish tomentose

12 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT spot dorsally. ZOOTAXA Terminalia (Figs. 1–2): Brownish-yellow, whitish setose; epandrium narrow, 1381 gonocoxa large, concealing aedeagus. Hypandrium simple.

FIGURES 1–2. Proctacanthella taina sp. nov., male terminalia, 1. lateral view, 2. gonostylus. Abbreviations: Ae=aedeagus, Ep=epandrium, Gc=gonocoxa, Gs=Gonostylus, Hp=hypandrium, S8=sternite 8, T8=tergite 8.

Female. Measurements, body 12.3–12.5 mm, wing 7.5–8.3 mm. Head: Ocellar tubercle with black setae. Wing: Crossvein well before middle of discal cell. Vein R4 usually with minute spur vein. Fore and middle femora with yellow extending to basal 1/3 ventrally, or entirely yellow ventrally; apical 1/2–2/3 of femora yellow. Abdomen: Black. Tergites with small brown tomentose spot medially. Terminalia: Black, round in cross- section, with apical spines. Types. DOMINICAN REPUBLIC: Pr. Montecristi, 4 km. N. of / Botoncillo 25.iv.2000 / R.E.Woodruff, hand (holotype %, FSCA). Paratypes, pr. / Montecristi, 10 km. S. / Montecristi, 2.x.85 / Woodruff & Stange / Blacklight trap (1 &, FSCA); RD-186 Rd. to Playa Buen / Hombre, Montecristi prov. / 132 m, 19o46.213'N 71o23. / 996'W, 6.xii.2003, D. / Perez, R.Bastardo, A. Mar- / molejos (1 &, USNM); Bahoruco [province], 5.8 km SW. / Neiba eastern playa / of Lago Enriquillo / 10-25-17N 71-26-38W [10o25.17'N 71o26.38'W] ~ -5 m, 3 Apr 2004 J / Rawlins, R.Davidson, C.Young Salt scrub / on sandy playa hand / collected, Sample 50143 ~ Carnegie Museum / Specimen Number / CMNH 260,729 (1 %, CMNH); same data except Specimen Number / CMNH 307,313 (1 &, CMNH). Distribution. Captured in sandy areas at low elevation during April, May, October, and

REVIEW OF ASILIDS © 2006 Magnolia Press 13 ZOOTAXA December in Bahoruco and Montecristi provinces, Dominican Republic. 1381 Etymology. Noun in apposition, taina referring to the species being an aboriginal of the West Indies. Remarks. Proctacanthella taina is readily distinguished from the 3 Proctacanthus species by the gradually produced face, whitish vestiture of the body, and the combined characters of the terminalia (Figs. 1–2). This is the first record of Proctacanthella in the West Indies.

Proctacanthus Macquart

Proctacanthus Macquart, 1838: 120 (1839: 236). Type species: philadelphicus Macquart (Coquil- lett, 1910: 595). Hull, 1962: 487; Martin & Papavero, 1962: 77. Catalogue.

Remarks. A largely New World genus, only 6 WI species are known, and 3 of these are from Hispaniola. Here we add Azua, La Altagracia, La Vega, National District, Peravia, and San Cristóbal provinces in the Dominican Republic and Haiti to its distribution in Hispaniola. The species are recognized by a large body (20–45 mm), abruptly produced

face, long tapered abdomen that extends beyond the wings, and veins R4 and R5 meeting the wing margin before the apex. Species perch on bare soil or low on vegetation. Ovipositors have conspicuous spines to deposite eggs in the soil.

Proctacanthus darlingtoni Curran (Table 1)

Proctacanthus darlingtoni Curran, 1951: 6. Type locality: Dominican Republic, Barahona; holotype: % (MCZ). Hull, 1962: 488; Martin & Papavero, 1970: 78.

New records. DOMINICAN REPUBLIC: Dunas de Baní, Peravia prov. / 10.xi.2003, R H. Bastardo (1 &, USNM); DOMINICAN REPUBLIC: / prov. Peravia / 1 km S.E. Galeón / 19.vi.1976 / E. Grissell / E. Marcano, on / Casearia iliciflora (1 %, FSCA); NE of Jarabacoa / 31.v.1989, S.W.Dunkle (1 %, FSCA); prov. Peravia, Las / Salinas 19.v.85 / Woodruff & Stange / Hand catch (1 %, 2 &, FSCA); Peravia prov.Bahia ~ 5 m / W. of Baní 18o17.460' N / 70o23.090'W, 90 m, 8.viii. / 2006 D.Perez, R.Bastardo (6 %, [Apis mellifera prey] USNM, MHND). Distribution. Captured during May, June, and November at low altitude in Barahona and Peravia provinces. Remarks. Proctacanthus darlingtoni was originally described from Barahona province. This study reports the first records of the species from Peravia province. Proctacanthus darlingtoni is distinguished from congeners by pale yellowish or white

14 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT vestiture of the head and thorax, the black ground color and grayish tomentum of the ZOOTAXA thorax, and black femora. Contrastingly, the vestiture of the head is largely yellow, the 1381 cheeks black, and the thoracic dorsum and femora are red in P. dominicana and P. vittatus. Also, the postpronotum and anepisternum anteriorly have white vestiture in P. darlingtoni whereas it is black in P. dominicana and P. vi tt at u s.

Proctacanthus dominicana Curran (Table 1)

Proctacanthus dominicana, Curran 1951: 9. Type locality: Dominican Republic, Sánchez [Samaná province]; holotype: %, (AMNH). Hull. 1962: 488; Martin & Papavero. 1970: 78.

New records. DOMINICAN REPUBLIC: prov. Azua, 2 km Hato Nuevo, 23.x. / 1986, R.E.Woodruff / & P.E.Skelley, (1 %, FSCA); RD-277 Guaraguao PN del / Este / La Altagracia prov. / 18o20.296 N 68o48.907'W, / near sea level, 19–20.vii.2004, / D.Perez (d/n) (1 %, USNM); La Altagracia prov., trail / Guaraguao – Cueva del Puente, / 18o19.532'N 68o48.525'W, sea / level, 24.viii.2006, D.Perez, B.Hierro (day/night) (1 %, USNM); Pedernales prov., Pasosena, / on rd. to Los Arroyos, 18o04. / 829'N 71o44.091'W, 69 m, / 17.viii.2006, D.Perez, R.Bastardo (5 %, USNM); Peravia prov., Arroyo Grande / bridge diving Honduras / Angostura, 18o21.383'N 70o / 25.993'W, 230 m, 8.viii.2006, / D.Perez, R.Bastardo (1 %, 2 &, USNM); prov. San Cristóbal / Palenque, 22.x.1986 / R.Woodruff / & J.H.Frank (1 %, FSCA); Santo Domingo [Santo Domingo province] / 3 Sept. 1971 ~ T.E.Rogers / Coll. ~ Proctacanthus / dominicana / Curran / det. J.Wilcox 1979 (1 %, 1 &, FSCA). Distribution. Captured at low elevations during May and July-October in Azua, La Altagracia, Pedernales, Peravia, San Cristóbal, Sánchez, Santo Domingo provinces. Remarks. Proctacanthus dominicana is distinguished by the reddish ground color of the thorax and black setose anepisternum. The color of the fronal setae are variable, ranging from entirely yellow, black, or mixed. The species was originally report from Sánchez province. Its distribution is expanded to include six additional provinces.

Proctacanthus vittatus (Olivier) (Table 1)

Asilus vittatus Olivier, 1789: 263. Type locality: San Domingo, type lost. Martin & Papavero, 1970: 91. Unplaced species. Proctacanthus rufiventris Macquart, 1838: 123 (1839:239). Type locality: 'Santo Domingo', type: % (MNHM). Hull, 1962: 488; Martin & Papavero, 1970: 79. Proctacanthus vittatus Oliver. Curran 1951: 7. Hull, 1962: 488.

REVIEW OF ASILIDS © 2006 Magnolia Press 15 ZOOTAXA New records. DOMINICAN REPUBLIC: La Altagracia prov. PN del / Este Guaraguao 1381 / 18o19.568'N 68o48.500'W / 0–5 meters 21 July 2004 / N.E.Woodley (2 %, USNM); RD- 277 Guaraguao PN del / Este Guaraguao / La Altagracia prov. / 18o20.296 N 68o48.907'W, / near sea level, 19–20.vii.2004, / D.Perez (d/n) (2 %, MHND); prov., La Altagracia / Parque Nac. del Este / Bayahibe Park ser. office / 07AUG1999 / M.A.Ivie & K.A.Guerrero (1 &, MAIC); prov. La Vega / nr. Manabao / 03.Sep. 1988 ~ M.A.Ivie, T.K. Philips / & K.A.Johnson (1 %, USNM); prov. / National District, Boca / Chica 22.VI.1998 / R.E.Woodruff (1 %, 1 &, FSCA); HAITI: Fond Parisien, February 11–18, 1922 (1 %, AMNH); Darien, November 14, 1934, P. Darlington (1 &, AMNH); Port-au- Prince, September 2, 1934, M.Bates (1 &, AMNH). Distribution. Captured at or near sea level during June, July, September, and November in La Altagracia, La Vega and National District provinces, Dominican Republic, and in Haiti. Remarks. Proctacanthus vittatus is the most widespread of the 3 species of Proctacanthus in Hispaniola, occurring at low altitude in southern Haiti and southern and northern Dominican Republic. It is most similar to P. dominicana, differing primarily by the dense setae on sternite 9 being undivided medially, and the apical fringe of setae on sternite 8 is shorter, finer, and doubled rowed (Curran, 1951).

Wilcoxius Martin (Table 1)

Wilcoxius Martin, 1975: 71. Type species: truncus Martin (orig. des.).

Remarks. This small Neotropical genus consists of eleven species, of which 6 are from Cuba and the Dominican Republic (Scarbrough & Perez-Gelabert, 2005). These are small to medium sized species that perch on rocks and fallen vegetation on the ground, low branches in sunny areas of open scrubby or second growth vegetation (Fisher in press). Of the 2 species from the Dominican Republic, W. planus Scarbrough & Perez-Gelabert (2005) were captured in road-side vegetation in Azua province. Wilcoxius tumidus Scarbrough & Perez-Gelabert (2005) was captured at low elevation in scrub vegetation near streams in Azua, San Juan, and Santo Domingo provinces, and open, grassy, pine forests at higher elevations in Pedernales province.

Andrenosoma Rondani

Andrenosoma Rondani, 1856: 160. Type species: Laphria atra. Hull, 1962: 349; Martin & Papav- ero, 1970: 42.

Remarks. This large genus is distributed worldwide (Fisher, in press). About 35 described species are recorded in the Americas with most being in the Neotropics, of which 6 extend into the West Indies. Two of these are reported here for the first time from Hispaniola. Species are found in mature forests, perching on leaves and fallen logs near forest breaks. The immature stages develop in fallen logs. Dr. Eric Fisher, California Department of Food and Agriculture, Sacramento, California, is revising a part of this genus.

Andrenosoma chalybeum Williston (Table 1)

Andrenosoma chalybeum Williston, 1885: 56. Type locality: Cuba, type:?. Fisher, 1986: 90, distribution Cuba, Hispaniola, Puerto Rico. Andrenosoma chalybea Williston. Martin & Papavero, 1970: 42.

New records. DOMINICAN REPUBLIC: RD-152 Barreras, Azua prov., / 18o19.527’ N, 70o54.411'W 174 m 14.vii.2003, D.Perez / R.Bastardo, B.Hierro (night) (2 %, USNM); RD-154 Busú, El Curro / Sierra Martín Garcia, Azua prov. / 18º17.819'N 70º57.287'W, 771 m, 16- / 17.vii.2003 / D.Perez, R.Bastardo, B.Hierro (1 %, MHND); Azua [province] / East side of crest, / Sierra Martín Garcia, / 7 km WNW Barrero. / 18–21N, 70–58W. [18º21'N, 21º58'W]. 860 m ~ 25–26 July 1992 / C.Young, R.Davidson, / S.Thompson, J.Rawlings / Cloud forest adjacent / to disturbed forest ~ Carnegie Museum / Specimen Number / CMNH-306,582 (8 %, 5 &, CMNH); RD-153 La Poza de Agua Nueva / El Curro, Sierra Martin Garcia / Azua prov. 18o18.324'N, 70o57 / 176'W~800 m, 15–16.vii.2003, / D.Perez, R.Bastardo, B.Hierro, day/night) (2 %, EMF); same location and collectors, 17–18.vii.2003 (2 %, USNM, MHND); prov. / Barahona, nr. Filipines / Larimar Mine: 26-vi-7- / vii.1992: R.E.Woodruff / & P.E.Skelley, (1 &, FSCA); Hato Mayor [province]. Parque Los / Haitises, 3 km W Cueva / de Arena, 19-04N, 69-29W [19º04'N, 69º29'W] ~ 20 m. 7–9 July 1992 / R.Davidson, J.Rawlins / S.Thompson, C.Young / Mesic lowland forest ~ Carnegie Museum / Specimen Number / CMNH- 305,527 (1 %, CMNH); prov. La Altagracia / 1 km N. San Rafael, / de Yuma or 1 km E. / La Pinita, 28.v.85 ~ L.A.Stange / Coll. (1 &, FSCA); RD-277 Guaraguao, P N del / Este, La Altagracia prov., / 18o20.296'N 68o48.907'W, / near sea level, 19–20.vii.2004 /

REVIEW OF ASILIDS © 2006 Magnolia Press 17 ZOOTAXA D.Perez (d/n) (1 %, 4 &, USNM); La Altagracia prov. PN del / Este, Guaraguao, / 1381 18o19.568'N 68o48.500'W, / 0–5 meters 21 July 2004 / N.E.Woodley (1 %, &, EMF); RD- 267 Rio Pedernales, nr / Fuerte Banano, Pedernales / prov., 280 m, 12.vii.2004, / D.Perez (d) (4 %, &, USNM); Pedernales prov., P N Sierra de / Barohuco, Las Abejas / 18o09.011'N 71o37.342'W. / 1150 meters 11 July 2004 / N.E. Woodley (1 &, EMF); RD- 266 Las Abejas, P.N. / Sierra de Bahoruco, Peder- / nales prov., 1,310 m, / 18o09.011'N, 71o37.342'W 11.vii 2004, D.Perez (d/n) (1 %, USNM); prov. Pedernales / 24 km N Cabo Rojo, 610 m / 19 AUG 1988, wet forrest / M.Ivie, Philips & Johnson (1 %, 2 &, CMNH); Pedernales [province], 26 km / N Cabo Rojo. / 18-06N, 71-38W [18º06'N, 71º38'W]. 730 m / 19–25 July 1990 / L.Masner, J.Rawlins / C.Young. Wet deciduous / forest; intercept trap ~ Carnegie Museum / Specimen Number / CMNH-306,386 (1 %, CMNH); prov. Pedernales / 24 km N. Cabo Rojo, 610 m / 20 AUG-09 SEP 1988, wet forest / Malaise trap, MA.Ivie, / T.K.Philips & K.A.Johnson (1 %, CMNH). PUERTO RICO:Orocovis / Bosque Estatal del / Toro Negro, 1.2 km W / Cerro Dona Juana, / Cordillera Central ~ 18- 10-30N, 66-29-37W [18º10.25'N, 66º29.07'W] / 600 m, 8 June 1996 / R.Davidson, J.Rawlins, / C.Young, S.Thompson / M Klinger, W.Zanol (1 %, CMNH); Cayey / Bosque Estatal de / Carite, 4.2 km SE / Campamento Guavate / 18-05-25N, 66-02-07W [18º05.25'N, 66º02.07'W] / 580 m, 7 June 1996, / C.Young R.Davidson, J.Rawlins, S.Thompson / M.Klingler, W.Zanol (1 &, CMNH). Distribution. Cuba, Hispaniola, Puerto Rico. Captured during May, June, July, August- September from sea level-1150 m in elevation in Azua, Barahona, La Altagracia, Hato Mayor, and Pedernales provinces in the Dominican Republic. Remarks. Andrenosoma chalybeum is distinguished by the bluish-black ground color and the straight proboscis with an obtuse apex (Fisher, 1986). All specimens examined from the Dominican Republic and Puerto Rico fit the description of this species and is the first record of the species in the former location. Yet at least three size groups exists and probably represent different species. The series from the Dominican Republic consist of two distinct size groups, one moderately large and the other much smaller. Furthermore, the specimens from Puerto Rico are much larger than any of those from the Dominican Republic. Specimens from the Dominican Republic were captured on follen trees at or near sea level in moist, deciduous forests in Azua, Barahona, La Altagracia, Hato Mayor, and Pedernales provinces of Dominican Republic and in Puerto Rico.

Andrenosoma ruficaudum (Williston) (Table 1)

Laphria ruficauda Williston, 1885: 55. Type locality: Santo Domingo, type ?. Andrenosoma ruficaudum (Williston). Martin & Papavero, 1970: 38. New comb.

New records. DOMINICAN REPUBLIC: RD-155 Charquito Prieto / Curro, Sierra

18 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT Martín García, / Azua prov. 18º18.324'N 70º / 57.176'W, 731 m, 17–18.vii. / 2003 ZOOTAXA D.Perez, R.Bastardo, / B.Hierro (day/night) (1 %, USNM); La Altagracia prov., PN del / 1381 Este, Boca de Yuma / 18o21.508'N 68o36.956''W. / 3–20 meters 20 July 2004 / N.E. Woodley (1 %, 2 &, USNM); RD-277 Guaraguao, P N del / Este, La Altagracia prov., / 18o20.296'N 68o48.907'W, / near sea level, 19–20.vii.2004 / D.Perez (d/n) (1 %, 1 &, EMF); RD-276 Boca de Yuma PN / del Este, La Altagracia prov., / 18o19.554'N 68o48.503'W. / near sea level, 19–20.vii.2004, / D.Perez (d/n) (1 %, MHND); La Altagracia prov., PN del / Este, Guaraguao / 18º19.568'N 68º48.500'W / 0–5 meters 2 July 2005 / S.W. Lingafelter (1 &, MHND); La Altagracia province / Punta Cana near Ecological / Reserve 0–5 meters / 18º30.477'N, 68º22.499'W / 12 June 2005 N.E.Woodley (1 &, USNM); prov., La Altagracia / Boca de Yuma entr. Par. Nac / del Este, 06Aug1999, 12 m / 18º21.904’N 68º37.094’W / M.A.Ivie, beating vegetation (1 &, MAIC); La Altagracia [province] 2 km N / Bayahibe, 18-23N, 68-51N / 3 m, 3 July 1992 ~ C.Young, R. Davidson / S. Thompson, J.Rawlins / Dry seasonal forest / on limestone (1 &, CMNH); prov. Pedernales / 24 km N. Cabo Rojo, 610 m / Mike Ivie, Philips & Johnson (1 %, CMNH); El Seibo [province], / El Cocal, Enero 1972 / C.Diaz Carela, Marcano number 12418 (1 &, IIBZ); Boca de Yuma [Altagracia province] / 23 Junio 1973 / J.Cicero, L.Ariza J., Marcano number 13683 (1&, IIBZ). Distribution. Captured during June and July from sea level to 731 m in elevation in Azua, El Seibo, La Altagracia, Pedernales, and Santo Domingo provinces. Remarks. Andrenosoma ruficaudum is distinguished by the reddish face, antenna, legs, and the apical 1/2 or more of the abdomen. It was captured near sea level in forests in the Dominican Republic. The type locality of the species is Santo Domingo [province]. Its distribution now includes Azua, El Seibo, La Altagracia, and Pedernales provinces.

Dasyllis Loew

Laphria Loew, 1851: 20. Type species: Laphria haemorrhoa Wiedemann (orig. des.). Andrenosoma Martin & Papavero, 1970: 45. Dasyllis Hull, 1962: 358. Fisher, 1986: 105.

Remarks. This Neotropical genus currently consists of 26 medium-sized to large species with a black or reddish to yellowish integument (Fisher 1986, Fisher in press). The species are distinguished from all other Andrenosomini by the following combination of characters: broad, wedge-shaped proboscis; proboscis apically parallel-sided, apex with distinct V-shaped notch; gonostylus bilobed with longitudinal 'comb' of stout reclinate setae, and usually preceeded by an apical tooth (Fisher, 1986). Only Dasyllis cressa (McAtee) from Cuba and D. rufa (Bromley) from Hispaniola occur in the West Indies. Dr. Eric Fisher, California Department of Food and Agriculture, Sacramento, CA, will report a new genus of ten species that includes the latter two species. As most Andrenosomini,

Dasyllis rufa (Bromley) (Table 1)

Andrenosoma rufum Bromley, 1931: 132. Type locality: Dominican Republic, Santo Domingo, type: % (BMNH); Martin & Papavero, 1970: 43. Dasyllis rufa, Fisher 1986: 194. New comb.

New records. DOMINICAN REPUBLIC: RD-239~5 km SW Peralta, / Azua prov., / 23.iv.2004, D. / Perez, B.Hierro (d) (1 &, MHND); RD-153 La Poza de Agua / Nueva , El Curro, Sierra / Martín García, / Azua prov.18º18.324'N / 70º57.176'W ~ 800 m, 15- / 16.vii.2003 / D.Perez, R.Bastardo, B.Hierro / (day/night) (1%, USNM); La Altagracia prov., PN del / Este, Boca de Yuma 18o21.508'N 68o36.956'W. / 3–20 meters 20 July 2004 / N.E.Woodley (1 %, USNM); La Altagracia prov., PN del / Este, Boca de Yuma / 18o21.508''N 68o36.956'W / 3–20 meters 20 July 2004 / N.E.Woodley (1 &, USNM); Republica Dominicana / Pedernales [province], Oviedo / Dry forest / 5–8.vi.2001 / H. Takizawa (1 &, MHND). Distribution: Captured in April, June, and July at sea level to 800 meters in Azua, La Altagracia, Pedernales, and Santo Domingo provinces. Fisher (1986) reported 2 specimens from Haiti. Remarks. Dasyllis rufa is distinguished by its thick body, black antenna, broad proboscis, apex subrectangular with an apical notch. Its distribution now includes Azua, La Altagracia, and Pedernales provinces.

Neophoneus Williston

Phoneus Macquart, 1838:79 (1839:195) (preocc. Kaup 1829). Type species servillei Macquart . Neophoneus Williston, 1889: 255. New comb. Hull, 1962: 353; Martin & Papavero, 1970: 44.

Neophoneus is removed from the list of West Indies asilids because of an error in identification in existing literature (Martin & Papavero, 1970). Bigot (1878) described flavotibius from Haiti, probably placing it in Phoneus because Macquart's figure of the type species, servillei, appears somewhat like a female Efferia. Bigot apparently did not realize that Macquart’s genus was a Laphriinae. Diagnosis of P. flavotibius as a species of Efferia is based on photographs of the type specimen (provided by J.M. Ayala), and the original description of the species and figures provided by Bigot (E. Fisher, pers. comm., 2006). The status of the species will be determined following examination of the type P. flavotibius and a review of the species of Efferia from Hispaniola.

Remarks. This Neotropical genus, consisting of 11 described species, is distinguished by a sharply pointed, usually upturned proboscis (Fisher pers. com.). This is the first report of the genus in the West Indies. Dr. Eric Fisher, California Department of Food and Agriculture, Sacramento, California, is revising this genus.

Pilica sp. (Table 1)

New records. DOMINICAN REPUBLIC: Azua [province] / East side of crest, / Sierra Martin Garcia, / 7 km WNW Barrero / 18-21N, 70-58W [18o21N, 70o58'W]. 860 m ~ 25–26 July 1992 / C.Young, R.Davidson, S.Thompson, J.Rawlins / Cloud Forest adjacent / to disturbed forest (1 %, CMNH); RD-153 La Poza de Agua Nueva / El Curro, Sierra Martín Garcia, / Azua prov., 18º18.324'N / 70º57.176’W about 800 m, 15–16. / vii.2003 / D.Perez, R.Bastardo, B.Hierro / (day/night) (1 %, 1 &, USNM, MHND); La Altagracia province / El Veron, Hoyo Azul / 18o33.610'N 68o26.881'W. / 25–40 meters 22 July 2004 / N.E.Woodley (1 %, USNM). Remarks. This slender bodied species with a black antenna and red abdomen was captured from 25 to 860 meters in mature forests, often near cloud forests, in Azua and La Altagracia provinces.

Subfamily Laphriinae

Tribe Atomosini

Atomosia Macquart

Atomosia Macquart, 1838: 73 (1839: 189). Type species: incisuralis Macquart (Coquillett 1910: 512) = puella (Wiedemann).

Remarks. This large primarily Neotropical genus includes about 50 species. Previously, six species were reported from the West Indies, 4 from Cuba (Bromley 1929) and 2 from Puerto Rico (Curran 1930). We report Atomosia from the Dominican Republic for the first time and 5 new species are described. The genus is distinguished by small robust bodies [5–12 mm], a strongly convergent vertex, and the antenna is without an apical style. The flagellum is longer than the scape and pedicel combined and has a dorsal excised notch at

REVIEW OF ASILIDS © 2006 Magnolia Press 21 ZOOTAXA or beyond the middle that bears a minute spine-like bristle. The abdomen consists of 6 1381 exposed segments with the sixth cup-shaped and concealing the terminalia. The dorsal surface of the tergites is coarsely punctate (Hull 1964). Species are very similar and often difficult to separate. Identification of species relies heavily on color patterns of the cuticle, tomentum, and vestiture. Male terminalia thus far have not been studied extensively and are usually not illustrated. Species perch on sunlit rocks, trees, branches, and leaves, and erect objects such as dead or live tree trunks, sides of buildings and fence supports. Perch selection is documented in only a few species.

Key to the known species of Atomosia from Hispaniola

1. Scutum white setose, lateral bristles and dorsocentral setae black ...... 2 - Scutum yellow setose, lateral bristles and dorsocentral setae yellow...... 4 2. Face entirely tomentose with white setae; hind femur and hind tibia black, at most narrow bases yellow or reddish; tarsi black setose dorsally, hind tarsus entirely blackish3 - Face in part shining black atomentose, mystax and epistomal margin black setose, basal 1/3 or 1/2 femur and tibia yellow; tarsi mostly yellowish setose dorsally; hind tarsus in part brownish-yellow or reddish; male flagellum and terminalia (Figs. 3–7) ...... anacaona sp. nov. 3. Ocellar tubercle black setose; flagellum (Fig. 18) with dorsal corner of excised notch spur-like, as long or longer than spine-like bristle, sensory area 1/2 as long as flagellum; terminalia (Figs. 19–22)...... jimagua sp. nov. - Ocellar tubercle white setose; flagellum with dorsal corner angular, not long spur-like, much shorter than spine-like bristle, sensory area short, about 1/3 as long as flagellum (Fig. 13); terminalia (Figs. 14–17)...... jagua sp. nov. 4. Anepisternum in part shiny black, atomentose; hind trochanter yellow; basal tarsomere of hind tarsus yellowish basally; fore and middle femora largely yellow, dorsoapical 2/ 3 blackish; basal 1/2 of hind femur and hind tibiae yellow, black apically; male flagellum and terminalia (Figs. 23–27) ...... yurabia sp. nov. - Anepisternum entirely tomentose; hind trochanter and hind tarsus black; femora and hind tibia largely black, narrow apices and bases of femora and narrow bases of hind tibia yellowish or reddish; male flagellum and terminalia (Figs. 8–12, 119)ciguaya sp. nov.

Atomosia anacaona sp. nov. (Table 1, Figs. 3–7)

Male. Black, face in part shining black, black mystax, body white setose and tomentose. Measurements, body 6.7–6.9 mm; wing 5.5–6.1mm. Head: Face largely white tomentose,

22 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT white setose, mystax shining black, atomentose with black vestiture, epistomal margin ZOOTAXA sparsely black. Palpus, proboscis, and occiput white setose. Antenna mostly or entirely 1381 black setose, flagellum (Fig. 3) about 1/3 as long as scape and pedicel combined, scape slightly longer than pedicel, pedicel wide, 2.5 times basal width; flagellum blackish-brown tomentose, scape grayish; flagellum with short spine at or beyond apical 1/3, apex of spine sometimes white; sensory area oval, tapered apically, about 1/4–1/3 as long as flagellum. Frons white tomentose, white setose, sparse black setae sometimes present. Ocellar tubercle with 2 long, apically diverging setae, black, sometimes one black, or both white. Occiput mostly sparse brownish tomentose, white laterally and ventrally, postocular bristles dull white. Thorax: Black, white tomentose, white setae and bristles. Scutum with bluish reflections dorsally, narrow apical and lateral margins white tomentose; setae sparse, short, white, less than length of pedicel, longer setae posteriorly, forming triangular pattern medially; lateral bristles and dorsocentral setae black, latter about as long as scape. Scutellum with bluish reflections and long, decumbent white setae; 2 stout pale yellow marginal scutellar bristles plus one or 2 shorter, thinner ones on each side, stout bristles about as long as scutellum medially. Anatergite with 6–7, stout, whitish or pale yellowish bristles. Pleura entirely dense white tomentose, white vestiture; thick pleural bristle about as long as anepisternum. Halter creamy yellow, base dull brown. Wing: Hyaline, iridescent, mostly brownish microtrichose, basal cells partly amicrotrichose. Cell r4 with base just beyond d cell, wide at wing margin; cell r5 usually open at wing margin, diameter about as great as length of r-m crossvein, rarely petiolate apically. Cell m1 ventrally slightly convergent apically; m3 just before or in line with apex of cell d. Legs: Whitish setae and bristles. Coxae black, white tomentose, white setose. Trochanters brownish-yellow dorsally, apical margin of hind trochanter, especially ventrally, blackish. Femora and tibiae mostly black, setae and bristles yellowish; fore and middle femora basally, and apex of all femora narrowly yellow, fore and middle femora sometimes pale brownish-yellow anteriorly; hind femur with basal 1/3 or 1/2 yellow; basal 1/3–2/3 yellow, or grading from yellow to black; hind tibia with sparse brown bristles. Tarsi blackish with mostly yellowish setae dorsally, sparse long black setae apically on anterior pairs and entirely black on hind tarsus dorsally; fore and middle tarsi with mostly pale yellow bristles, reddish or black bristles anteriorly; hind tarsus with only black or sparse amber colored bristles. Basal tarsomere of hind tarsus mostly brownish-yellow or yellowish-brown. Abdomen: Black with blue reflections, twice as long as scutum and scutellum combined, sides parallel, junction of tergite 2 and 3 slightly constricted; short black, recumbent setae dorsally, setae longer, white or yellowish laterally; tergites 1–3 with stout whitish bristles laterally. Narrow apical margins of tergites 2–5 white tomentose; narrow apical margin of tergite 1 black, white tomentose laterally. Brownish-gray tomentose,

FIGURES 3–7. Atomosia anacaona sp. nov., 3. male flagellum, 4. epandrium, lateral view, 5. terminalia, ventral view, 6–7. aedeagus dorsal and lateral views. Abbreviations: ea=ejaculatory apodeme, Gc=gonocoxa, Hp=hypandrium, sa=sensory area, sp=spine.

Terminalia (Figs. 4–7): Gonocoxa narrow apically. Aedeagus longer than wide dorsally, trifid apex declinate. Ejaculatory apodeme clavate, slightly curved forward apically. Hypandrium triangular. Female. Measurements, body 7.5–8.7mm; wing 6.7–7.5mm. Flagellum about 1/3 longer than pedicel and scape combined; spine at apical 1/3; sensory spot 1/3 as long as flagellum. Legs: More extensively yellow with base of femora and tibiae often yellow on

24 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT basal 1/4 or 1/2; hind tibia with apical 1/4 blackish; basal tarsomere of hind tarsus mostly ZOOTAXA to entirely yellowish or brownish-yellow. Abdomen: Tergite 1 only slightly wider than 1381 tergite 2, tergites 3–6 usually slightly wider, constriction at junction of tergite 2 and 3 absent. Types. DOMINICAN REPUBLIC: RD-243 Babosico, on way to / Jánico, Santiago prov., 515 m / 19o20.955'N, 70o47.503'W, 27. / iv.2004 D.Perez, B.Hierro, / R.Bastardo (d) (holotype %, USNM). Paratypes, same data (6 %, &, USNM, MHND). Distribution. Known only from type locality in Santiago province, Dominican Republic. Collected at 515 meters in elevation, in April, from the trunks of palm trees. Etymology. Named after Anacaona, also called 'golden flower', who was an Indian 'queen' and wife of Caonabo, one of the five 'caciques' of the island of Santo Domingo. Remarks. Atomosia anacaona is distinguished from congeners in the West Indies by the black mystax wide yellow base of the hind femur, brownish-yellow basal tarsomere of the hind tarsus, and combined characters of the antenna and terminalia (Figs. 3–7). Though similar to A. parva Bigot (Bigot 1856, Bromley 1929, Curran 1935) in the black mystax, A. anacaona is distinguished by the yellow base of the femora and tibiae, the short stout marginal scutellar bristles which are about as long as the scutellum medially, the black lateral scutal bristles, the white setose palps and thoracic dorsum, and the black setose abdominal tergites. In A. parva, the face and palps are entirely black setose, the legs are largely black, the scutellar bristles are much thinner and longer than the scutellum medially, the lateral scutal bristles are largely yellowish, only the notopleural bristle is black, and the dorsum of the thorax and abdomen are yellowish setose.

Atomosia ciguaya sp. nov. (Table 1, Figs. 8–12, 119)

Male. Black, thorax and abdomen dorsally yellow setose, black tarsi with yellowish setae and bristles. Measurements, body 5.4–7.5 mm; wing 4.3–6.3 mm. Head: Whitish or yellowish tomentose, whitish setose. Antenna black, brownish tomentose, scape basally brownish-yellow or whitish; black setose, scape ventrally whitish; flagellum (Fig. 8) 1/ 5–1/3 longer than pedicel and scape combined, spine white, at apical 1/4, sensory area small, usually white, sometimes brownish, about 1/5 as long as flagellum; pedicel wide apically about twice as wide as base, about 3/4 as long as scape. Frons, ocellar tubercle laterally, and occiput dorsally yellowish-white tomentose. Ocellar tubercle with 2 long white setae. Occiput dorsally with stout white or yellowish postocular bristles. Thorax: White to slightly yellowish tomentose. Pronotal bristles and setae white or slightly yellowish. Scutum dorsally atomentose, narrow margin anteriorly and laterally white; setae and bristles yellow, setae decumbent, about as long as pedicel; dorsocentrals erect, longest setae longer than scape and pedicel combined; notopleural sometimes black.

REVIEW OF ASILIDS © 2006 Magnolia Press 25 ZOOTAXA Scutellum with yellowish setose and bristles, setae decumbent, about as long as scape; 2 1381 stout marginal bristles, 2 or 4 thinner, shorter bristles. Numerous anatergal bristles and sparse setae, yellowish. Pleura entirely tomentose, white, sometimes yellowish, vestiture white, stout yellowish pleural bristle about as long as anepisternum. Halter creamy white, base dull brown.

FIGURES 8–12. Atomosia ciguaya sp. nov., 8. male flagellum, 9. epandrium, lateral view, 10. terminalia, ventral view, 11–12. aedeagus dorsal and lateral views. Abbreviations: ea=ejaculatory apodeme, Gc=gonocoxa, Hp=hypandrium, sa=sensory area, sp=spine.

Wing: Hyaline, iridescent, brown microtrichose. Cell r4 narrow apically, diameter at wing margin as great or slighter greater than length of r-m crossvein, base well beyond

apex of cell d. Crossvein r-m just before middle of cell d. Cell m1 with parallel sides,

sometimes slightly convergent apically. Apex of cell m3 at or beyond base of d cell. Legs. Bristles and setae pale yellow or whitish. Coxae and trochanters dark, black or

26 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT yellowish-brown. Femora and tibiae mostly blackish, narrow apex of all femora and basal ZOOTAXA 1/3 of fore and middle femora brownish-yellow or yellow; basal 1/5 or 1/4 of hind femur 1381 and hind tibia pale yellow. Tarsi blackish, bristles and setae yellowish, sparse long black setae on apical one or 2 tarsomeres; hind tibia sometimes with sparse black bristles. Abdomen: Black with bluish reflections, densely punctate, slightly narrower than thorax, sides parallel, twice as long as scutum and scutellum combined; setae yellow, short dorsally, longer, paler laterally; stout whitish bristles on tergites 1–3, sometimes much shorter, thinner bristle on tergite 4; tergite 7 wide, about 1/4 as long as wide. Sternites black, tomentum sparse, grayish or brownish, sparsely setose. Terminalia (Figs. 8–12): Gonocoxa narrow apically. Aedeagus wider than long dorsally, apex declinate; ejaculatory apodeme clavate apically. Hypandrium oval, pointed medially. Female. Measurements, body 6.6–6.8 mm; wing 4.3–5.5 mm. Flagellum 1/4 to 1/3 longer than scape and pedicel combined, spine at apical 1/3, sensory area 1/4 as long as long as flagellum. Abdomen 2–3 times combined length of scutum and scutellum. Types. DOMINICAN REPUBLIC: RD-232, Road San Juan- / Vallejuelo near river, San / Juan prov., 449 m 18o41. / 754'N, 71o11.725'W, 21.iv. / 2004, D.Perez, B.Hierro, / R.Bastardo (d) (holotype %, USNM). Paratypes, RD-229 – 2 km road to San / Juan, near bridge on Viajama / river, Azua prov., 294 m, 18o / 35.245'N 70o58.085'W, 20.iv. / 2004, D.Perez, B.Hierro, R. / Bastardo (d) (1 &, USNM); Bara- / hona [province] 6 km NW Paraiso, Rio / Nizao. 18-02N, 71-12W [18o02'N, 71o12'W] 170 m / 25–26 July 1990. C.Young, / J.E.Rawlins, S.A.Thompson (1 &, CMNH); Montecristi [province] 5 km / NNE Botoncillo / 50 m 19-46N, 71-24W [19o46'N, 71o24'W] ~ 29–30 November 1992 / R.Davidson, M.Klinger / S.Thompson, J.Rawlins / Arid thornscrub (2 &, CMNH); prov. Montecristi Guayubin / Cerro Gordo Abajo 78 m / 258302 m E 2173910 mN, / N.Bastardo, F.Ortiz, W.Ortiz, / R.Bastardo Potrero (1 %, MHND); RD-235 - 2 km road to Pedro / Santana, Elias Piña prov. / 314 m, 19o05.484'N, 71o41. / 738'W, 21.iv.2004, D.Perez, / B.Hierro, R.Bastardo (n) (1 &, USNM); prov. Independencia / Los Rios / Lago, Enriquillo / 25-v-1986 / R.Miller & L.Stange (1 &, FSCA); RD-224 5 km on road to / Botoncillo, Montecristi prov., / [19o] 46.220’N, 71o24.001'W, 16.iv. / 2004, D.Perez, B.Hierro (d/n) (1 %, USNM); RD-186 Rd. to Playa Buen / Hombre, Montecristi prov./ 132 m, 19o46.213'N, 71o23. / 996'W, 6–7.xii.2003, D. / Perez, R.Bastardo, A.Mar- / molejos (4 %, 4 &, USNM); prov. Montecristi / 9 km N. Villa Elisa / 4-vi-1986, R.Miller / & Stange (1 &, FSCA); Pedernales prov., km 13 Rd. / Cabo Rojo, Aceitillar, 18o01. / 866'N 71o38.806'W, 123 m, / 16.viii.2006, D.Perez, R.Bastardo (night) (1 %, USNM); RD- 057 – 2 km N of Baní, / Peravia prov., dry scrub, / 18o18.489'N, 70o20.599'W, / 16.xi.2002, D.Perez, R. / Bastardo, B.Hierro (night) (1 %, USNM); Peravia prov., Bahía, ~5 km / W of Baní,18o17.460'N / 70o23.090'W, 90 m, 8.viii. / 2006, D.Perez, R.Bastardo (12 %, 20 &, USNM); Peravia prov., Arroyo Grande / bridge diving Honduras / Angostura, 18o21.383'N 70o / 25.993'W, 230 m, 8.viii.2006, / D.Perez, R.Bastardo (2 %, 1

REVIEW OF ASILIDS © 2006 Magnolia Press 27 ZOOTAXA &, USNM); Arroyo Margajita, Mina de / Oro de Pueblo Viejo, Sánchez / Ramírez prov., 1381 373–634 m E / 2094-137 m N, 2.vii.2003, / R.H.Bastardo (1 &, USNM); same data as holotype (11 %, 20 &, USNM, MHND); RD-238 1 km SE Bastida, / foothills near canal of Yaque / del Sur River, San Juan prov., / 23.iv.2004, D.Perez, B.Hierro (1 %, 1 &, USNM); prov. San Juan, El / Capa, 17 m, N.E. / Vallejuelo, 27-v- / 1986, R.Miller & / L.Stange (1 %, 2 &, FSCA). Distribution. Azua, Barahona, Elias Piña, Independencia, Montecristi, Pedernales, Peravia, Sánchez Ramírez, and San Juan provinces. Collected among riverine, lake, and arid cactus-scrub vegetation at low altitudes (50–634 m) during April–July and November- December. Etymology. Named after the Ciguayos, the Taino group that inhabited present day San Juan de la Maguana province in west-central Dominican Republic. Remarks. Atomosia ciguaya is distinguished from congeners by the characters in the key, especially the yellow setose dorsum of the thorax and abdomen, yellowish or whitish setae of the ocellar tubercle, and the combined characters of the flagellum and terminalia (Figs. 8–12). It will key to A. danforthi (Curran 1935) but is distinguished from that species by the largely blackish legs, pale ocellar setae, and terminalia.

Atomosia jagua sp. nov. (Table 1, Figs. 13–17)

Male. Black, white tomentose, white setose, legs black. Measurements, body 5.7–6.3 mm; wing 4.7–5.1 mm. Head: White tomentose, mostly white setose. Antenna black, mostly or entirely black setose, scape medioventrally with one long bristly seta; scape slightly longer than pedicel, flagellum (Fig. 13) about 1/3 longer than pedicel and scape combined; flagellum incised at apical 1/3 dorsally, dorsal corner usually angular, sometimes with short scale-like process; flagellum apically narrowed and slightly curved ventrally, white sensory pad elongate and tapered apically, about 1/3 as long as flagellum. Ocellar tubercle with 2 long, apically divergent, black setae. Occiput thin, brownish-gray tomentose dorsally, white setose; postocular bristles, stout, whitish. Thorax. Black, white tomentose, white setose, bristles and setae mostly whitish. Scutum and scutellum with blue reflections, minutely punctate with short, white, recumbent setae, setae about as long as pedicel, none longer or denser posteriorly; scutum with narrow anterior and lateral margins gray tomentose; lateral bristles and dorsocentral setae black; latter about as long as pedicel; postalar callus reddish-brown. Scutellum with 2 stout whitish marginal bristles, slightly shorter than scutellum medially. Anatergal bristles and sparse setae white, or mixed black and white. Pleura entirely white tomentose, vestiture white; one long, stout, black or white pleural bristle, as long as anepisternum. Halter yellowish, base brownish narrowly. Wing: Hyaline, iridescent; brown microtrichose, basal cells partly amicrotrichose;

28 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT ZOOTAXA veins brownish, yellowish-brown. Cell r4 wide apically, vein R4 at or just above wing tip; 1381 cell r5 narrowly open, diameter usually less than length of r-m, sometimes petiolate with veins R5 and M1 fused at or just before wing margin. Crossvein r-m just before middle of d cell. Cell m1 with sides parallel to wing margin, vein M2 slightly convergent apically; cell m3 with apex at, just before or beyond base of m1.

FIGURES 13–17. Atomosia jagua sp. nov., 13. male flagellum, 14. epandrium, lateral view, 15. terminalia, ventral view, 16–17. aedeagus, dorsal and lateral views. Abbreviations: ea=ejaculatory apodeme, Gc=gonocoxa, Hp=hypandrium, sa=sensory area, sp=spine.

Legs: Blackish, vestiture pale, sparse black bristles on tibiae, mostly hind tibia. Coxae gray tomentose. Narrow base and apex of femora and narrow base of femora and tibiae

REVIEW OF ASILIDS © 2006 Magnolia Press 29 ZOOTAXA reddish or yellow; basal 1/3 of fore tibia sometimes pale. Tarsi black with mostly black 1381 vestiture; fore and middle tarsi with 2 yellowish bristles basally, dorsal setae black, except laterally on basal tarsomere of fore and middle tarsi; dense ventral setae brownish-yellow. Abdomen: About twice as long and slightly narrower than thorax, sides parallel; black with blue reflections. Tergites coarsely punctate with short recumbent white setae dorsally, black setae medially, longer white setae laterally and ventrally; narrow apical margins of tergites 2–5 gray tomentose; tergites 1–3 or 4 with sparse whitish bristles laterally. Tergite 7 about 1/3 as long as wide, base deeply notched medially. Sternites black white or brownish tomentose with scattered white setae. Terminalia (Figs. 14–17): Gonocoxa wide apically. Aedeagus with lateral apodemes, somewhat narrow apically, trifid distiphallus declinate, ejaculatory apodeme clavate, curved apically. Hypandrium triangular, apex wide. Female. Measurements, body 6.4–7.0 mm; wing 5.4–5.7 mm. Face with long setae just below antenna often brownish. Flagellum about twice as long as scape and pedicel combined; spine just beyond apial 1/3; sensory area 1/4 as long as flagellum. Types. DOMINICAN REPUBLIC: RD-272 Caseta 1, PN Sierra / de Bahoruco, Independencia / prov. 18o16.038'N71o32.691'W / 1,239 m, 14.vii.2004, D.Perez (holotype %, USNM). Paratypes, prov., Pedernales / ca. 35 km. Cabo Rojo / Las Abejas, 1250 m / 26 Aug – Sep. 1988 ~ M.A.Ivie, T.K.Phillips / & K.A.Johnson colrs. ~ Malaise / trap (1 %, MAIC); Pedernales [province] / ca. 35 km N Cabo Rojo 1250 m / Las Abejas, 26 AUG. 1988 / beating veg., M.A.Ivie / T.K.Philips & K.A.Johnson (3 %, MAIC); same location and collectors except date 09 SEP1988 (1 %, MAIC); Pedernales [province], 26 km N Cabo / Rojo, 18-06N, 71-38W [18o 06'N 71o38’W] / 730 m 16 July 1992 ~ C.Young, R.Davidson / S.Thompson, J.Rawlins / mesic deciduous forest / with scattered pines (1 %, CMNH); Pedernales [province] / ca. 35 km NNW Cabo Rojo / 1430 m, El Aceitillar / 09SEP1988, pine forest / M.Ivie, Phillips & Johnson (1 %, MAIC); same location and collectors except 1370 km (1 %, MAIC); prov. Pedernales / ca. 35 km N Cabo Rojo / El Aceitillar Las Abejas / 1250–1430 m 23 AUG1988, M.Ivie, Philips & Johnson (1 &, MAIC); Pedernales / prov. El Aceitillar / ca. 35 km NNW Cabo Rojo / 23 AUG 1988,1370–1430 m ~ M.A.Ivie, T.K.Philips / & K.A.Johnson colls (1 %, 1 &, MAIC); Pedernales [province], Las Abeja[s] / 38 km NNW Cabo Rojo, / (18-09W, 71-38W) [18o09'W 71o38'W] ~ 1440 m 13–16 July 1987 / Robert L.Davidson and J.Rawlins (1 &, CMNH); same location and collectors except 1250 m, 15 July 1987 (2 &, CMNH); Peder- / nales prov. 'Las Abejas' 7.3 km NNE Mercedes 18o19'N, 71o38''W 1100- / 1150 m 23 August 1983 ~ F.M.Harrington / L.D.Weintraub / Collectors (1 &, FSCA); Pedernales prov., Pasosena, / on rd. to Los Arroyos, 18o04. / 829'N 71o44.091'W, 69 m, / 17.viii.2006, D.Perez, R.Bastardo (1 %, 3 &, USNM); Pedernales prov., km 13 Rd. / Cabo Rojo, Aceitillar, 18o01. / 866'N 71o38.806'W, 123 m, / 16.viii.2006, D.Perez, R.Bastardo (night) (9 %, 7 &, USNM); Pedernales prov., 0.7 km N / Cruce Aguas Negras, Mencia, / 18o06.956'N 71o43.0388'W, 362 m, / 17.viii.2006, D.Perez, R.Bastardo (1 %, USNM);

30 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT Peravia prov., Bahía, ~5 km / W of Baní, 18o17.460'N / 70o23.090'W, 90 m, 8.viii. / 2006, ZOOTAXA D.Perez, R.Bastardo (3 %, USNM). HAITI: Departement / du Sud, Villa Formon / 31 km 1381 NW Les Cayes, / S slope Formon / Massif de La Hotte, ~ 18-20N 74-01W [18o20'N 74o0'W] / 1405 m, 7–8 Sept 1995 R.Davidson, G.Onore, / J.Rawlins, Disturbed / Forest and Field (1 %, CMNH). Distribution. Independencia, Pedernales, and Peravia provinces in the Dominican Republic and Departement / du Sud, Haiti. Collected in deciduous and pine forests between 730 and 1440 m in elevation from July to September. Etymology. From the Taino jagua, referring to the black color of aged fruit tree sap [jagua] that was used for body paint and dying cotton material. The name refers to the ground color of this species. Remarks. Atomosia jagua keys to A. danforthi Curran (1935) but is distinguished by the 2 black ocellar dorsocentral setae and black lateral scutal bristles, reddish-brown postalar callus, black legs, white setose body, and brownish-yellow or reddish base of the hind tarsus. It is similar to A. jimagua in the whitish setose dorsum of the thorax and abdomen but is distinguished by the characters above, those in the key, the long tergite 7, and by the combined characters of the flagellum and terminalia (Figs. 13–17). In A. jimagua, the sensory area on the flagellum is about 1/2 as long as flagellum, the spine-like dorsal corner of excised notch is as long or longer than the ventral spine, the legs are in part yellow, tergite 7 is strap-like, about 1/4 as long as wide, the gonocoxa is narrow apically, the hypandrium is triangular, and the ejaculatory apodeme is capitate and strongly curved forward (Figs. 19–22).

Atomosia jimagua sp. nov. (Table 1, Figs. 18–22)

Because A. jimagua is quite similar to A. jagua, only significant differences are given in the following description. Male. Black, white setose, black legs; spur-like flagellar notch and long sensory pad. Measurements, body 7.3–8.3 mm; wing 6.0–7.0 mm. Head: Scape 1/3 longer than pedicel, flagellum (Fig. 18) about 1/4 or 1/3 longer than pedicel and scape combined; flagellum incised at apical 1/4, dorsal corner long, spur-like, spine below as long or shorter than spur, sensory area elongate, at least 1/2 as long as flagellum. Ocellar tubercle with 2 long, apically diverging, white setae. Thorax: Anatergal bristles usually mixed black and white, sometimes entirely black or white. Long pleural bristle usually white, sometimes black.

Wing: Cell r5 narrowly open at wing margin, never petiolate, diameter slightly greater that length of r-m. Crossvein r-m just before or at middle of cell d. Legs: Black, narrow apices of femora, narrow bases of tibiae, yellowish or reddish;

REVIEW OF ASILIDS © 2006 Magnolia Press 31 ZOOTAXA vestiture whitish, hind tibia with sparse black bristles. Tarsi with black vestiture; fore and 1381 middle basal tarsomeres with 2 whitish bristles, setae usually black dorsally. Abdomen: About twice as long as scutum and scutellum combined; tergites 1–3 with sparse whitish bristles laterally; tergite 7 short, strap-like, about 1/4 as long as wide. Terminalia (Figs. 19–22): Gonocoxa narrow apically. Aedeagus with lateral apodemes separated, trifid distiphallus decumbent, ejaculatory apodeme strongly curved forward, capitate apically. Hypandrium triangular. Female. Measurements, body 7.4–8.5 mm; wings 6.2–8.7 mm.

FIGURES 18–22. Atomosia jimagua sp. nov., 18. male flagellum, 19. epandrium, lateral view, 20. terminalia, ventral view, 21–22. aedeagus dorsal and lateral views. Abbreviations: ea=ejaculatory apodeme, Gc=gonocoxa, Hp=hypandrium, sa=sensory area, sp=spine.

32 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT Types. DOMINICAN REPUBLIC: Pedernales prov. PN Jaragua / trail to Carlitos ZOOTAXA ca. 6 km S of / Highway 44, 106 meters / 17o48.932'No 71028.271'W / 8 July 2004 N. 1381 E.Woodley (% holotype, USNM). Paratypes, same data as holotype (2 &, USNM); prov. Pedernales / 13.5 km N Cabo Rojo, 140 m / cactus-thorn scrub, flight / intercept trap, M.A. Ivie, / T.K.Philips & K.A.Johnson ~ 21 AUG-10 / SEP 1988 (3 %, 1 &, MAIC); prov. Pedernales / 13.5 km N. Cabo Rojo, 140 m / 21 AUG10-SEP1988 flight / intercept trap, M.Ivie, / T.K.Philip & K.A.Johnson (2 %, MAIC); prov. Pedernales 24 km N. Cabo Rojo, 610 m / Aug 19 1988, wet forest / M.Ivie, Philips & Johnson (2 %, 2 &, MAIC); Pedernales prov. S. / end of Lago de Oviedo / 26.v.1986, R.Miller / & L.Stange (3 %, FSCA). Distribution. Pedernales province in the Dominican Republic. Captured during May and August-September in cactus-thorn scrub vegetation at low elevations (106–140 m) and in wet forests at higher elevations (610 m). Etymology. From the Taino language, jimagua meaning 'twins, those that are very similar', referring to the fact that A. jimagua and A. jagua are very similar species. Remarks. Atomosia jimagua runs to A. danforthi in Curran's key (1935) but is distinguished by the darker legs and white setae on the thoracic and abdominal dorsum. It differs from A. jagua by the characters discussed in the remark section of that species.

Atomosia yurabia sp. nov. (Table 1, Figs. 23–27)

Male. Black, yellowish setose dorsally, anepisternum partly atomentose. Measurements, 5.0–6.0 mm; wing 3.8–4.4 mm. Head: Face and frons white tomentose including ocellar tubercle laterally, white setose, sometimes slightly yellowish. Palpus and proboscis black, white setose. Antenna black setose, basal 1/2 of scape ventrally white setose; pedicel 2/3 as long as scape; flagellum (Fig. 23) 1.5 times as long as scape and pedicel combined, excised notch at apical 1/3, sensory pad white, small, 1/3 or 1/4 as long as flagellum. Ocellar tubercle with 2 whitish or yellowish setae. Occiput thin, yellowish tomentose with yellowish or whitish setae dorsally, dense white tomentose along eye margin and ventrally with mostly white setae; postocular bristles whitish or yellowish. Thorax: Black, mostly white tomentose, including narrow anterior and lateral margins of scutum, pleural setae and pronotal bristles and setae usually white. Scutum and scutellum dorsally atomentose, with minute puntuations and short, decumbent, yellow, sometimes whitish, setae, latter about as long as pedicel and evenly distributed, none especially dense; dorsocentral setae and lateral scutal, 2 scutellar, anatergal, and anepisternal bristles, yellow or yellowish, latter about as long as anepisternum; dorsocentral setae about as long as scape; scutellar margin with 2 setae, 1/2 or 2/3 as long as pristles. Anepisternum partly shining black medially. Halter creamy yellow, base dull

apically, brownish-yellow or brown. Cells r5 and m1 open, slightly narrowed apically, former diameter at wing margin equal to or slighter greater than r-m length. Vein r-m just

before or at middle of cell d. Apex of m3 just before or beyond base of m1.

FIGURES 23–27. Atomosia yurabia sp. nov., 23. male flagellum, 24. epandrium, lateral view, 25. terminalia, ventral view, 26–27. aedeagus, dorsal and lateral views. Abbreviations: ea=ejaculatory apodeme, Gc=gonocoxa, Hp=hypandrium, sa=sensory area, sp=spine.

Legs: Yellowish setae and bristles. Coxae black, white tomentose with whitish setae. Trochanters brownish-yellow or yellow, apex of hind trochanter usually blackish at least in part. Femora mostly yellow, apical 1/3 or 1/2 dorsally blackish; hind femur with wide blackish band extending from just beyond middle to narrow yellow apex. Fore and middle tibiae with apices brown or blackish, yellow anteriorly, brownish-yellow posteriorly; hind

34 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT tibia mostly blackish, at least with basal 1/2 anteriorly and basal 1/3 posteriorly yellow. ZOOTAXA Tarsi mostly blackish, bristles mostly blackish, at least 2 or more pale yellowish; basal 1381 tarsomere of fore and middle tarsi yellow basally, that of hind tarsus narrowly reddish; setae yellowish dorsally except 3 long black setae on all apical tarsomeres and entire hind tarsus dorsally. Abdomen: Sides parallel, base subequal in width to thorax, 1.5 times as long as scutum and scutellum combined; densely punctate dorsally with short decumbent whitish setae, longer setae laterally; narrow apical margin of tergites 1–5 white tomentose; tergites 1–3 or 4, with whitish bristles laterally. Sternites brownish, thinly whitish tomentose, white setose. Terminalia (Figs. 24–27). Gonocoxa narrow, longer than wide, apex pointed in lateral view. Aedeagus with declinate trifid distiphallus, lateral apodemes sheet-like; ejaculatory apodeme long and narrow. Hypandrium wider than long, triangular. Female. Measurements, body 5.3–6.3 mm; wings 4.9–5.3 mm. Thoracic and abdominal dorsa entirely golden yellow setose. Specimens examined. DOMINICAN REPUBLIC: RD-233 Few km E Valle- / juelo, San Juan prov., 704 m, / 18o40.122'N 71o18.446'W, / 21.iv.2004, D.Perez, B. / Hierro, R.Bastardo (d) (holotype %, USNM). Paratypes, same data as holotype (4 %, 7 &, USNM, MHND); Pedernales prov., 0.7 km N. / Cruce Aguas Negras - Mencia / 18o06.956'N 71o43.388'W, 326 m / 17.viii.2006, D.Perez, R.Bastardo (1 &, USNM). Distribution. Only known from the type locality, at 704 m in San Juan and Pedernales provinces. Etymology. From the Taino language, yurabia for small place, alluding to the fact that the type series was captured at one location on the same day. Remarks. Atomosia yurabia runs to A. rica (Curran, 1935) but is distinguished by its smaller body (5.0–6.3 mm), swollen, wider, pedicel, which is about 2/3 as long as the scape, yellow scutal bristles, largely yellow femora, largely yellow setae on fore and middle tarsi, and combined characters of the flagellum and male terminalia (Figs. 23–27). Atomosia rica is a larger (6.5–8.0 mm) species with at least black notopleural bristles, and largely blackish femora with only the basal 1/4 or 1/3 of the femora yellow. The apex of the gonocoxa is wider and rounded in lateral view, and the ejaculatory apodeme is significantly wider on the apical 1/2. Contrastingly, the dorsum of the thorax and abdomen is whitish setose in the male (yellow in the female), apex of the gonocoxa is narrower, more pointed in lateral view and the ejaculatory apodeme is significantly narrower on the apical 1/2.

Atoniomyia Hermann

Atonia Williston, 1889: 257. (preocc. Gistel, 1848). Type species: mikii Williston (Williston, 1901: 316).

Remarks. This largely Neotropical genus consists of 16 species, one of which is Neartic (Martin & Papavero, 1970). It is distinguished by its small (6–7 mm) robust body, strongly punctate scutum and abdominal tergites, short, appressed setae, and the terminalia are concealed by the cup-shaped tergite 6. The flagellum has an apical, down-turned scape with microsegment at its base. Species perch on soil near streams or low on vegetation near ground level (Fisher and Hespenheide, 1992; Fisher, in press). We report 2 females of an undescribed species and A. mikii (Williston) from Azua and Barahona provinces in the Dominican Republic.

Atoniomyia mikii (Williston) (Table 1, Figs. 28–32)

Atomosia mikii, Williston 1886: 290. Type locality: Dominican Republic, Santo Domingo, type: &?. Atonia mikii (Williston), 1889: 257. Atoniomyia mikii (Williston). Hull, 1962: 61; Martin & Papavero, 1970: 554.

Redescription, male. Black, silvery-white tomentose, white setose, robust body. Measurements, body 7.0–8.0 mm; wing 6.1–7.2 mm. Head: Silvery-white tomentose white setose. Face gradually produced, ventral 1/2 well exposed beyond eye; entirely or largely white setose, abundant ventrally, 2 narrow stripes dorsally. Palpus brownish- yellow. Proboscis black. Antenna black, brownish-yellow tomentose, black setose; flagellum (Fig. 28) flat and widest on apical 1/2, constricted about apical 1/3 ventrally, with long apical style, about 1/3 as long as flagellum, and short microsegment; scape slightly shorter than pedicel, latter 2 combined about 2/3 as long as flagellum. Ocellar tubercle with 2 long black setae, about as long as antenna. Occiput usually grayish tomentose, eye margin denser; setae and bristles pale, sparse, setae most abundant ventrally, dorsal postocellar bristles sometimes black. Thorax: Black, silvery-gray tomentose, bristles and setae mostly white. Scutum and scutellum black with minute punctuations, atomentose, shining black with recumbent white setae, setae about as long as pedicel; dorsocentral setae black, longest 3/4 as long as scape and pedicel combined; sparse, erect, black setae laterally; lateral bristles black. Apical margin of scutellum with numerous, long, whitish or yellowish setae, about as long as flagellum. Anatergite with narrow vertical stripe of black, spine-like bristles, and short bristly setae. Pleura mostly dense silvery-white tomentose with white vestiture; anespisternum partly atomentose, shining black, extending ventrally on to katepisternum as narrow line. Halter creamy white, base dull brown. Wing: Hyaline, mostly brown-yellow microtrichose, basal cells somewhat sparse. Cell

36 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT ZOOTAXA r4 with base beyond apex of cell d, wide apically with R4 above and R5 well below wing 1381 apex. Crossvein r-m just before middle of cell d. Apex of cell m3 well beyond cell d.

FIGURES 28–32. Atoniomyia mikii (Williston), 28. male flagellum, 29. epandrium, lateral view, 30. terminalia, ventral view, 31–32. aedeagus, dorsal and lateral views. Abbreviations: Gc=gonocoxa, Ea=ejaculatory apodeme, Hp=hypandrium, sa=sensory area, sp=spine, St=style.

Legs: Black, white setose, with mostly whitish bristles; coxae silvery-white tomentose. Tibiae with narrow bases reddish, sparse black bristles laterally, and abundant flat or erect white setae ventrally; middle tibia narrow, somewhat capitate apically, with 2–3 unusually long bristles, about 2/3 as long as tibia; gradually wider from base to apex, apex about

REVIEW OF ASILIDS © 2006 Magnolia Press 37 ZOOTAXA twice as wide as base. Tarsi with basal 4 tarsomeres largely pale, yellow on fore and 1381 middle tarsi, yellowish-brown or red on hind tarsus, each slightly brownish apically, apical tarsomere entirely blackish; setae largely black dorsally, yellowish fore tarsus, black and yellowish on middle tarsus, entirely black on hind tarsus; bristles black, sparse yellow on fore and middle tarsi, outer bristles about 1/2 as long as tarsi. Abdomen: Black, sides parallel, twice as long as scutum and scutellum combined; tergites coarsely punctate, abundant short, brown setae medially, much longer whitish setae laterally and apically on tergites 6 and 7; tergites 1–5 laterally with sparse, stout, yellowish, spine-like bristles. Sternites brownish with scattered, short, pale yellow to whitish setae. Terminalia (Figs. 29–32): Black, white setose. Gonocoxa narrow. Aedeagus yellow, wider laterally than long in dorsal view; trifid distiphallus slightly curved horizontally; ejaculatory apodeme wide on apical 1/2. Hypandrium rectangular. Female: Measurements, body 8.0–9.2 mm, wing 7.0–8.3 mm. Tomentum and setae yellowish-gray or yellow. Head: Dorsal 1/2 of face, entire frons, vertex and occiput medially yellowish-gray to golden yellow tomentose; sparse brown setae present on dorsal 1/2 of face, frons sometimes reddish yellow. Flagellum without obvious constriction . Legs: Middle tibia slender, apex not strongly swollen. Tarsi paler than in male, setae largely yellowish. New records. DOMINICAN REPUBLIC: Azua [province], 8 km NE Padre / Las Casas, Rio Las / Cuevas ~18-24 N, 70-53 W. [18o24'N, 70o53'W] 580 m / 7 August 1990 / J.Rawlings, S.Thompson (10 %, 18 &, CMNH); Bara / hona [province], 6 km NW Paraiso, Rio / Nizao 18o02'N 71o12'W, 170 m / 25–26.vii.1990, C.Young, J.E.Rawlins, S.A.Thompson, (1 %, 2 &, CMNH). Distribution. Captured between 170 and 580 m in elevation during July and August in Azua, Barahona, and Santo Domingo provinces, Dominican Republic. Remarks. Atoniomyia mikii is distinguished by the shape of the flagellum and length of the style (Fig. 28), anepisternum partly atomentose, the unusually long bristles on the tibiae and tarsi, and long setae laterally and brown setae medially on most tergites. The male is further recognized by the combined characters of the terminalia (Figs. 29–32) and the black setose, apically enlarged middle tibiae. Williston described A. mikii from specimens captured in Santo Domingo province. Azua and Barahona provinces are added to its distribution. This is the first report of the species since the original publication in the late 1800's.

Atoniomyia sp. (Table 1)

New records. DOMINICAN REPUBLIC: prov. / Barahona, nr. Filipinas Larimar Mine: 20 26- / vi.1992: R.E.Woodruff / P.E.Skelley, at light (1 &, FSCA); same locality,

38 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT collectors (1 &, FSCA). ZOOTAXA Distribution. Barahona province, Dominican Republic. 1381 Remarks. Material of a second species captured in Barahona province is available but only in the female sex. In addition, the flagellum is absent in both specimens. The species is distinguished from Atoniomyia mikii by its smaller body, entirely tomentose pleura, black tarsi, wing venation, and setation pattern of the body. Because the antenna is an important element as well as male characters, especially the terminalia, in the diagnosis of the species, we leave it underscribed.

Cerotainia Schiner

Cerotainia Schiner, 1866: 662 (in key), 673 (1868: 170; second erection of genus). Type species: Laphria xanthoptera Wiedemann (orig. des.). Hull, 1962: 394; Martin & Papavero, 1970: 52. Ceratotaenia Lynch Arribálzaga, 1880: 52, emend.

Remarks. This largely Neotropical genus is widely distributed from Connecticut and Michigan in North America southward to Argentina and Chile with 30 species. Three species are reported from the West Indies, one each from Cuba (Bromley 1929), Jamaica (Johnson 1919, Farr 1963), and Dominica (Scarbrough & Knutson 1989). The genus is distinguished by the small body, strongly divergent frons and vertex, the antenna is as long as the height of the head, scape 3–4 times as long as pedicel, and the thoracic dorsum and abdominal tergites are punctuated, and the sixth abdominal tergite is cup-shaped, concealing the terminalia. The species are usually found on the exposed, tips of branches and leaves of trees, shrubs, herbaceous plants and tall grasses.

Cerotainia sola sp. nov. (Table 1, Figs. 33–36, 120)

Male. Black, scutum and abdomen dorsally with minute punctations and short, decumbent setae. Measurements, body 4.3–5.8 mm; wing 3.3–5.0 mm. Head. Mostly yellowish-gray tomentose, yellowish setose. Face with narrow margin of eye yellow, mystax whitish or yellowish setose, 3–6 long setae blackish. Palpus and proboscis yellowish setose. Antenna mostly black setose, scape with one contrastingly long, sometimes yellow, seta ventrally; scape 10 times longer than pedicel; flagellum longer than combined length of pedicel and scape; apical 1/3 of flagellum with small spur-like process dorsally and elongate, oval, white sensory area on inner surface. Frons yellowish-gray tomentose. Vertex yellowish- gray tomentose anteriorly, bare or sparsely brownish tomentose posteriorly and on to occiput dorsally, surface shiny black in dorsal view. Occiput mostly gray tomentose, shiny area dorsally bordered ventrally with yellow or brownish tomentum; white setose; stout yellowish or brownish-yellow bristles dorsally.

REVIEW OF ASILIDS © 2006 Magnolia Press 39 ZOOTAXA Thorax. Black, with bluish reflections dorsally; length greater than wide. Pronotum 1381 yellow tomentose. Scutum with short decumbent setae, anterior broadly, lateral and posterior margins narrowly yellow setose, medially setae blackish in lateral view, brownish or yellowish in dorsal view; lateral bristles yellow or brownish-yellow. Scutellum yellowish setose; mediotergite and anatergite brownish tomentose; anatergite bristles short, stout, and black or brownish-yellow. Pleura yellowish-gray or gray tomentose, vesititure usually yellowish, sometimes whitish; anepisternum atomentose with long yellow setae. Halter creamy yellow or white. Wing: Brownish, costal cell darkest, anal lobe hyaline basally; entirely brownish

microtrichose; iridescent. Vein R4 abruptly curved anteriorly before apex; R5 almost

straight. Cell m1 narrowly convergent at wing margin. Cell m3 usually without M3 vein apically, if present short, spur-like, never reaching wing margin. Legs. Black, tibiae and tarsi rarely uniformly brown, with whitish or yellowish setae and yellow bristles. Abdomen: About twice as long as thorax, narrow, 5 times as long as segment 2. Ground color black with blue reflections; yellow setose, short setae dorsally mixed with black, setae much longer laterally, those on tergites 1–2 laterally whitish. Narrow margins of tergites 2–4 whitish tomentose laterally; apical corner of tergite 5 narrowly and wide margin of 6 dull yellowish. Sternites dull grayish tomentose. Terminalia (Figs. 33–36): Aedeagus yellow or brownish-yellow, apex short, trifid. Female. Measurements, body 5.4–6.0 mm, wing 4.2–4.6 mm. Types. DOMINICAN REPUBLIC: Pedernales prov. 25 km N / of Cabo Rojo / 18o06.769'N, 71o37.245'W / 679 meters 19 June 2005 / N.E.Woodley (holotype %, USNM). Paratypes, Hato Mayor [province], Parque Los / Haitises, 3 km W Cueva / de Arena, 19-04N, 69-29W) [19o04'N, 69o29'W]. ~ 20 m. 7–9 July 1992 / R.Davidson, J.Rawlins / S.Thompson, C.Young / Mesic lowland forest (1 %, CMNH); RD-148 Up 100 m from El Sitio / del Agua, cloud forest N Los Bolos, / Sierra de Neiba, Independencia prov./ 18o39.339'N, 71o39.279'W, 1,520 m, / 9.vii.2003, D.Perez, R.Bastardo, B.Hierro. (day/night) (2 %, USNM); La Altagracia prov., P.N. del / Este, Boca de Yuma / 18o21.508'N, 68o36.956'W / 3–20 meters, 20 July 2004 / N.E.Woodley (1 %, USNM); La Altagracia prov., P.N. del / Este, Boca de Yuma / 18o21.508'N, 68o36.956'W / 3–20 meters, 6 July 2006 / N.E.Woodley (1 %, USNM); same data as holotype (1 %, 1 &, USNM); Pedernales [province], Las Abejas / 38 km NNW Cabo Rojo, / 18-09N, 71-38W [18o09'N, 71o38'W]. ~ 1250 m. 15 July 1987 / John E.Rawlins and / Robert L.Davidson (2 %, CMNH); Pedernales [province], 26 km / N Cabo Rojo, / (18-06N, 71-38W) [18o06'N, 71o38'W] ~ 13–20 July 1990 / L.Masner, J.Rawlins / C.Young. Wet deciduous / forest: intercept trap (1 %, 3 &, CMNH); same data except 19–25 July 1990 (2 &, CMNH); RD- 264 km 25 rd. Cabo Rojo / - Aceitillar, Pedernales prov., / 678 m, 10.vii.2004, D.Perez (2 %, 2 &, MHND); RD-262 La Charca, km 30 / road Cabo Rojo – Aceitillar, / Pedernales prov., 9.vii.2004, / D.Perez (d) (4 %, 8 &, USNM); RD-266 Las Abejas, P N / Sierra de

40 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT Bahoruco, Peder- / nales prov., 1,310 m, 18o09.011'N, 71o37.342'W, / 11.vii.2004, ZOOTAXA D.Perez (d/n) (2 %, 1 &, USNM). PUERTO RICO: Rio Abajo / State Forest / 18o18'N, 1381 66o45'W / 22 June 2002 / N.E.Woodley (1 %, 1 &, USNM).

FIGURES 33–36. Cerotainia sola sp. nov., 33. epandrium, lateral view, 34. male terminalia, ventral view, 35–36. aedeagus, dorsal and lateral views. Abbreviations: Ep=epandrium, ea=ejaculatory apodeme, Hp=hypandrium.

Distribution. Captured at low to high (20–1250) elevations during June and July from Hato Mayor, Independencia, La Altagracia, and Pedernales provinces in the Dominican Republic and Puerto Rico. Etymology. Latin sola referring to it being the only known species of the genus in

REVIEW OF ASILIDS © 2006 Magnolia Press 41 ZOOTAXA Hispaniola and Puerto Rico. 1381 Remarks. Cerotainia sola is similar to C. laticeps Bromley (1929) but differs in the yellowish tomentum of the head, blackish setae on the face and the scutum, yellowish setae of the head, thorax, and legs, black legs, brownish wings, narrow abdomen and terminalia (Figs. 33–36). In C. laticeps the frons is silvery tomentose, the setae of the scutum is entirely brassy, the tibiae are brown with narrow apices black, the hyaline wing, and broad abdomen.

Eumecosoma Schiner

Eumecosoma Schiner, 1866: 673. Type species: Laphria pleuritica Wiedemann, 1828, by original designation. Hull, 1962: 379; Martin & Papavero, 1970: 49.

Remarks. This Neotropical genus consists of 11 species described from southern Mexico to southern Brazil. These small dark flies have a slightly diverging vertex, the elongated, oval flagellum has an acute apex and a preapical spine. The abdomens are smooth or minutely punctuate with 7 exposed tergites in the male, 8 in females. In males, the seventh tergite is about 1/2 as long and wide as the sixth, and the terminalia are exposed. Species usually perch on the tops of leaves in shadows of small sunlit spots of nearly mature forests (Fisher and Hespenheide, 1992; Fisher, in press). Here we report the first record of the genus from the West Indies.

Eumecosoma caerulum sp. nov. (Table 1, Figs. 37–39)

Male. Body black with bright blue reflections. Measurements, body 5.7–5.9 mm; wing 4.9–5.0 mm. Head: Black, mostly black vestiture. Face narrow with parallel sides, width 1/5–1/6 as wide as head, dorsal 2/3 flat, scarcely visible in lateral view, yellowish tomentose with sparse, moderately long, black setae; lower 1/3 abruptly produced, visibly laterally, white tomentose with several long, black, seta-like bristles and numerous white setae. Palpus minute. Proboscis unusually short, about as long as flagellum, white setose ventrally. Flagellum (Fig. 37) elongate, as long as pedicel and scape combined, oval, apex acute, with preapical spine; surface black, without obvious delineated sensory area; pedicel oval, wider and longer than scape. Frons and vertex brown or black tomentose, may appear brownish-yellow at some angles, wide median brownish-yellow tomentose line extending from ocellar tubercle posteriorly onto occiput. Ocellar tubercle with 2 long diverging bristles, length subequal to antennae. Occiput mostly brown or black tomentose with several short black postocular bristles and sparse black setae dorsally, margins narrowly grayish tomentose with whitish setae.

42 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT Thorax: Black, scutum bright blue, smooth, without punctations; wide anterior margin ZOOTAXA posteriorly to postpronotum and narrow lateral and posterior margins brown tomentose; 1381 vestiture black, setae abundant to transverse suture, sparse posteriorly, 4–5 dorsocentral 3 bristles laterally, both long, thin. Scutellum bright blue with scattered black setae, apical margin brown tomentose with sparse black setae and 2 long, thick black bristles; apical crease absent. Mediotergite and anatergite brown tomentose, latter with numerous black setae. Pleuron gray tomentose except anepisternum, anepimeron, and katatergite brown or brownish-gray; one anepisternal and 4–5 katatergal bristles, black. Halter dull yellow. Sclerotized metacoxa present.

Wing: Slightly brownish. Discal and m3 cells slightly offset apically. Squama pale yellow, fringe sparse, white. Legs: Shining black, apices of femora and base of tibiae narrowly yellowish-brown. Coxae gray tomentose, vestiture whitish. Bristles black, mostly unusually long and thin. Femora and tibiae with yellowish setae ventrally except sparse exceptionally long, black setae; hind femur and tibia ventrally and middle tibia lateroapically with abundant setae of uniform length. Tarsi black, bristles unusually long and black, setae brownish-yellow or yellow except white on apical 4 tarsomeres ventrally. Abdomen: Sides parallel with segment 2 slightly wider than long and only slightly narrower than segment 5; only 7 tergites present. Black with blue reflections dorsally and minute punctuations, narrow apical margins of segments 2–4 grayish tomentose; vestiture pale, mostly short, yellowish, tergites 1–3 laterally and sternite 1 with 2–5 long bristles. Sternites thinly brownish-gray tomentose. Terminalia (Figs. 38–39): Black, rotated 180o. Epandrium undivided, ventral. Aedeagus trifid apically. Gonocoxa elongate, pointed apically. Hypandrium dorsal. Female. Measurements, body 6.5–7.5 mm; wing 6.5–7.5 mm. Face entirely yellowish- gray with only black bristles. Proboscis slightly longer than flagellum. Eight abdominal tergites present. Bristles thin; tergite 8 with sparse black bristles. Terminalia not dissected. Types. DOMINICAN REPUBLIC: La Vega prov., PN Armando / Bermúdez km 1–3 along trail / W of Ciénaga 900–1100 m / 19o01.753'N, 70o54.654''W / 24 June 2005 N.E.Woodley (holotype %, USNM). Paratypes, same data as holotype (1 %, 1 &, USNM); on way down from cloud forest / ~ 1,300 m, Los Bolos, Sierra de Neiba, Independencia prov. / 9.vii.2003, D.Perez, R. Bastardo, B.Hierro(day) (1 &, MHNH). Distribution. Captured at intermediate to high elevations (900–1300 m) during June and July in Independencia and La Vega provinces. Etymology. Latin caerulum for blue, referring to the bright blue color of the scutum and scutellum. Remarks. Eumecosoma caerulum, a member of the pleuriticum species group (Fisher and Hespenheide 1992), is the first species of this genus reported in the West Indies. It runs to E. metallescens Schiner (1868) from Peru in Curran's (1930) key but is distinguished from congeners by the color of the facial tomentum in the female and male antenna (Fig. 37) and terminalia (Figs. 38–39).

FIGURES 37–39. Eumecosoma caerulum sp. nov., 37. male antenna, lateral, 38–39. male terminalia, lateral and dorsal views. Abbreviations: Ae=aedeagus, Ep=epandrium, ea=ejaculatory apodeme, Gc=gonocoxa, Fl=flagellum, Hp=hypandrium, Pe=pedicel, sa=sensory area, Sc=scape, sp=spine.

Rhopalogaster Macquart

Rhopalogaster Macquart 1834: 279. Hull, 1962: 338; Martin & Papavero, 1970: 40.

Remarks. This Neotropical genus consists of only 7 species, one of which is reported from Cuba. Here we report the genus and a new species from the Dominican Republic for the first time. The genus is distinguished by its size (15–20 mm) laterally flat proboscis, short pile, greatly reduced bristles, strongly constricted abdomen, and very long antenna. The scape is usually 10 times as long as the pedicel, and the flagellum is 1.2–2 times the length of the scape and pedicel combined (Hull, 1962). Species inhabit mature forest below 1500 meters, perching on shaded dead and live tree trunks (Fisher and Hespenheide, 1992; Fisher, in press).

Female. Measurements, body 11.1 mm, wing 7.8 mm. Head: Black. Face and frons dull brownish-gray or brown tomentose; ventral 1/2 of face black setose, dorsal 1/2 with flat, yellowish setae, sparse thin black setae; epistomal margin sparsely white setose laterally. Palpus and proboscis white setose. Antenna, frons, vertex, and dorsal 1/3 of occiput black setose; flagellum flat, apex minutely notched, twice as long as scape. Occiput with ventral 2/3 gray tomentose, white setose. Thorax: Black. Scutum with wide median stripe and 2 lateral spots black tomentose, postpronotal spot, narrow line between stripe and lateral spots, and prescutellum between dorsocentral setae gray or brownish-gray tomentose; lateral margins brownish-gray or yellowish-gray tomentose; setae black, sparse white setae anteriorly. Scutellum black tomentose, sparsely black setae, with numerous long, thin, white marginal setae. Pleura white tomentose, propleuron and mesanepisternum with abundant, long, white setae, katepisternum black and white setose, setae sparse or absent elsewhere. Halter knob yellow, stalk blackish.

Wing: Mostly blackish microtrichose, sparse or absent in basal cells. Cell r5 petiolate apically, R5 and M1 fused just before wing margin. Crossvein r-m at basal 1/4 of discal cell.

Cell m3 long, pointed, apex just before base of m1. Legs: Black, mostly white vestiture. Coxae gray tomentose. Hind femur and tibiae apically, fore tibiae anteriorly, and hind tibiae entirely black setose. Hind femur and hind tibia slightly clubbed, preapex about twice diameter of corresponding bases; latter with abrupt apical notch anteriorly. Tarsi: Black, with mostly black vestiture. Abdomen: Strongly spatulate, segments 2–3 about 2/3 as wide as segment 1 basally, segments 4–5 slightly wider than segment 1; ground color black with blue iridescence, apical margin of tergites yellowish, color especially wide on tergite 2–4; tergite 1 entirely and base of tergite 2 narrowly brown tomentose; mostly black setose, tergites 1–2 whitish setose. Cercus yellow setose. Terminalia: Not dissected. Male. Unknown. Type. DOMINICAN REPUBLIC: La Vega prov., PN Armando / Bermúdez, km 1–3 along trail / W of La Ciénaga 900–1100 m / 19o01.753'N, 70o54.654' W / 17 July 2004 N. E.Woodley (holotype &, USNM). Distribution. Known only from the type locality, La Vega province, Dominican Republic. Etymology. Latin albidus for 'gleaming or white' referring to the abundant, flat, white setae on the face. Remarks. This is the first record of Rhopalogaster (Fig. 121) from the Dominican

REVIEW OF ASILIDS © 2006 Magnolia Press 45 ZOOTAXA Republic and the second from the West Indies. Rhopalogaster albidus is distinguished 1381 from R. bella Bromley (1929) of Cuba by the long, flat, white setae below the antennae and black setae on the dorsal 1/2 of the occiput.

Subfamily Ommatiinae

Ommatius Wiedemann

Remarks. This large, nearly world wide, genus inhabits mostly tropical regions. It is well represented in the Neotropics with 100 varified species now reported. Of these, nearly 1/2 (46) is from the West Indies, with 21 from Hispaniola, largely the Dominican Republic, and the remaining 25 the other islands. We also report the discovery of the male of Ommatius cinnamomeus Scarbrough (1984) and illustrate its terminalia. The genus is distinguished by the pectinate style. Many species prefer tips of bare branches of woody or herbaceous vegetation along margins of sunlit clearings in, paths and roads through forested areas.

Ommatius bastardoanus Scarbrough (Table 1)

Ommatius bastardoanus Scarbrough, In Scarbrough & Perez-Gelabert, 2003: 43. Type locality: Dominican Republic, La Vega province, type: % (USNM).

New Records. DOMINICAN REPUBLIC: RD-266 Las Abejas, P. N, / Sierra de Bahoruco, Peder- / nales prov., 1,310m, / 18o09.011'N 71o37.342'W / 11.vii 2004, D.Perez (d/n) (1 %, MHND); La Vega prov., P.N. Armando / Bermúdez, La Ciénaga de / Manabao, 19o04.022'N 71o51. / 849'W, 1183 m, 19.viii.2006, D.Perez, R.Bastardo, B.Hierro (11 %, 5 & USNM); RD-250 La Ciénaga – Los / Tablones, P N Armando / Bermúdez, La Vega prov., / 19o04.044'N 70o51.789'W. / 1,100 – 1,270 m, 17.vii.2004, / D.Perez (d) (6 %, 3 &, MHND); Republica Dominicana / prov. La Vega, La Sal Reserva / Ćientifica ÉbanoVerde, 2–3.vii. / 1992, S. Naverro, 225 (1 %, IIBZ). Distribution. Captured in montane forests between 1077 and 1270 mm in elevation during July and August in La Vega and Santiago provinces. Remarks. Ommatius bastardoanus is readily recognized by the unique terminalia, especially the somewhat truncate epandrium that bears a short, narrow ventroapical notch. Additionally, the strongly produced sternite 8 with a wide W-shaped apical margin and pyriform spermatheca distinguishes the female (Scarbrough & Perez-Gelabert, 2003). The type series were found in July. Here we extend its distribution to include Pedernales province and the month of August.

Ommatius cinnamomeus Scarbrough, 1984: 147. Type locality: Haiti, La Hotte, type: & USNM; 1997: 11; Scarbrough & Perez-Gelabert, 2003: 41.

FIGURES 40–44. Ommatius cinnamomeus Scarbrough, male terminalia. 40–42. Lateral, dorsal, and ventral views; 43. gonostylus, and 44. aedeagus. Abbreviations: Ce=cercus, Ep=epandrium, Gc=gonocoxa, Gcp=gonocoxal process, Gs=Gonostylus, Hp=hypandrium.

REVIEW OF ASILIDS © 2006 Magnolia Press 47 ZOOTAXA New records. DOMINICAN REPUBLIC: Independencia prov., 200 m / N. 1381 Destacamento Aguacate, / 18o19.681' N 71o41.830'W, / 1050 m, 12.viii.2006, D. / Perez, R.Bastardo, M.Hierro (1 %, 1 &, USNM); RD-264 km 25 rd Cabo Rojo / - Aceitillar, Pedernales prov. / 678 m, 10.vii.2004, D.Perez (1 %, USNM); Pedernales [province] 22.5 km / N. Cabo Rojo / 18-06N, 71-38W [10o06'N, 71o38'W] 540 m ~ 13–19 July 1990 / L.Masner, J.Rawlins, C.Young Deciduous forest: intercept trap (1 %, CMNH); Peder / nales [province] 25 km N. Cabo Rojo / 18-06N, 71-38W [10o06'N, 71o38'W] 730 m / 31 July 1990 J.Rawlins, / C.W.Young, S.Thompson (1 &, CMNH).

FIGURES 45–51. Right wing of seven species of Ommatius. 45. O. elusivus Scarbrough; 46. O. geminus sp. nov., 47. O. maculosus sp. nov., 48. O. inflatus Scarbrough, 49. O. laticrus sp. nov., 50. O. lucidatus Scarbrough, 51 O. praelongus sp. nov.

48 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT Distribution. Captured between 60 and 1050 m in elevation in Pedernales and ZOOTAXA Independencia provinces, Dominican Republic, and the southern peninsula of Haiti. 1381 Remarks. The male of Ommatius cinnamomeus is distinguished by the large body, dilated anterior margin of the wing, and the combined characters of the male terminalia (Figs. 40–44).

Ommatius elusivus Scarbrough (Table 1, Figs. 45, 52)

Ommatius elusivus Scarbrough, In Scarbrough & Perez-Gelabert, 2003: 46. Type locality: Domini- can Republic, PNAB, Santiago province, type: % (USNM).

FIGURES 52–53. Vestiture of abdominal sternites of % Ommatius. 52.O. elusivus Scarbrough showing angular, clubbed, and apically flattened thin bristles; 53. Sternite 6 of O. geminus nov. sp., showing a dense mass of overlapping bristles.

REVIEW OF ASILIDS © 2006 Magnolia Press 49 ZOOTAXA New records. DOMINICAN REPUBLIC: RD-250 La Ciénaga –Los / Tablones, PN 1381 Armando Bermúdez, La Vega prov., 19o04.004'N 70o51.789'W / 1,100–1,270 m, 29.vi.2004 / D.Perez (d) (1 %, 4 &, USNM); La Vega prov., P.N. Armando / Bermúdez, La Ciénaga de / Manabao, 19o04.022'N 71o51. / 849'W, 1183 m, 19.viii.2006, D.Perez, R.Bastardo, B.Hierro (8 & USNM); RD-250 La Ciénaga – Los / Tablones, PN Armando / Bermúdez, La Vega prov., / 19o04.044'N 70o51.789'W. / 1,100–1,270 m, 17.vii.2004, / D.Perez (d) (1 %, 4 & MHND).

FIGURES 54–56. Vestiture of abdominal sternites of % Ommatius. 54. O. inflatus Scarbrough showing distribution of mushroom- and pencil-shaped bristles; 55. O. laticrus sp. nov., showing the distribution of setae and bristles on sternites and the apically produced margins of sternites 6–7; 56. O. lucidatus Scarbrough, showing the distribution of setae and bristles on sternites, especially the dense fine setae and thin hooked bristles of sternite 6 and clustered mass of long, hooked bristles on the apical margin of sternite 7.

50 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT Distribution. Captured between 740 and 1270 m from May through August in La Vega ZOOTAXA and Santiago provinces. 1381 Remarks. Ommatius elusivus is distinguished from congeners by the characters in the key. It is associated with montane forests at intermediate to high elevations. The type series was taken from May through July (Scarbrough & Perez-Gelabert, 2003). Here we extend its distribution through August.

Ommatius geminus sp. nov. (Table 1, Figs. 46, 53, 59–66)

Male. Black, pleura entirely tomentose, hind tibia blackish. Measurements, body 14.0 mm; wing 10.3 mm. Head: Face yellowish tomentose dorsally, white ventrally, sparsely pale setose, mystax white, slightly shorter than proboscis, 6 black bristles present; face at antenna 1/9 as wide as head. Palpus brown basally, paler brown with whitish setae apically. Proboscis black, narrow base paler brown ventrally, sparsely white setose basally. Antenna mostly brown or black, black setose; pedicel with narrow base yellowish, wider than flagellum, apex of stout bristle at apex of flagellum ventrally; scape partly yellow setose dorsally; flagellum black; style with several ventral setae extending from base to apex, none exceptionally short. Frons brownish-yellow tomentose, brown or yellow and brown mixed setose, parallel sided, slightly convergentdorsally. Ocellar tubercle, longer than wide, about 1/2 as wide as vertex, black setose, 2 setae slightly longer than scape and pedicel combined. Occiput yellowish-gray tomentose dorsally, gray ventrally, vestiture pale, setae most abundant ventrally, 5–6 thin blackish postocular bristles, proclinate, apex just beyond posterior margin of eye. Thorax: Prothorax brownish-yellow tomentose dorsally, gray laterally, vestiture pale, 4 pale bristles; prosternum yellow. Scutum brownish or brownish-yellow tomentose dorsally, margins brownish-yellow or yellow; setae sparse, blackish dorsally, pale laterally; 4 long, thin, lateral bristles and 3 dorsocentral bristles blackish, notopleural bristle short, about 1/3 or 1/2 as long as supra-alar and postalar bristles, dorsocentral bristles, thinner, shorter than lateral bristles. Scutellum brownish-yellow tomentose, sparsely yellow setose, 2 blackish marginal bristles and 2–3 thin, marginal setae sometimes present. Pleura entirely tomentose, brownish-yellowish anteriorly, yellowish- gray or gray posteriorly, vestiture white or yellowish, anepisternum sparsely setose posteriorly, anepimeron with long, stout, yellowish bristle, metacoxa blackish medially, yellow laterally. Halter yellowish, stalk whitish.

Wing (Fig. 46): Hyaline, costal cell brownish; cells r4 and m1 moderately long; base of r4 beyond apex of cell d, about 2/3 as wide as cell at wing margin, veins divergent just before apex; base of m1 narrow, about 1/7 as wide as cell at basal 1/3, divergent apically, much wider at wing margin, base of M1 angular. Cells d and m3 narrow, former 1/3 as long

1381 slightly beyond base of cell m3.

FIGURES 57–58. Vestiture of abdominal sternites of % Ommatius. 57. O. maculosus sp. nov., showing unusual cuticular structures on sternites 4 and 5, transverse row of flat bristles on sternite 5, and dense vestiture on sternite 6; 58. O. praelongus, nov. sp., showing the distribution and variation of setae and bristles on sternites 4–7.

Legs: Fore coxa yellow, middle coxa mostly black, hind coxa blackish anteriorly; coxae grayish tomentose, vestiture yellowish, apical bristles thin. Fore and middle trochanters yellow, narrow margins brown; hind trochanter brown. Fore and middle femora mostly yellow, blackish anteriorly and posteriorly; hind femur mostly black, yellow ventrally and narrow ventral surfaces anteriorly and poseriorly; setae black, bristles mostly thin, short, and yellowish ventrally. Fore femur narrow apically, abruptly wider than hind femur, swollen posteriorly with sigmoid black bristle, and pale brownish-yellow callus-like groove. Middle femur with only 2 black bristles anteriorly. Hind femur narrow, 7 times as long as wide, 2 long yellowish posteroventral bristles basally. Tibiae with only black bristles, 2–3 on middle tibiae long and thin, dense setae yellowish; fore and middle tibiae yellow except basal 1/3 anteriorly and entire anterior surface, respectively; hind tibia

52 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT black except basal 1/3 posteriorly. Tarsi black, black setose, bristles black except one on ZOOTAXA fore tarsus; basal tarsomere of fore and middle tarsi yellowish, narrow apex reddish or 1381 brownish; hind tarsus with basal tarsomere about 1/3 wider than second tarsomere, 3 stout bristles present apically. Abdomen: Black, narrow apical margins of basal 4–5 segments and lateral margins of tergites yellow, grayish tomentose, basal 2–3 segments and lateral margins of tergites dense, setae largely yellow, long, somewhat stout on tergites 2–5, those laterally on sternites 3–5 somewhat bristly in 2 parallel rows; sternite 6 (Fig. 53) with dense, cluster of long, overlapping, brown bristles apically, row of thin setae posteriorly; apical margin of sternite 7 produced ventrally with abundant bristly setae. Terminalia (Figs. 59–63): Black, yellow and black setose. Epandrium narrow, apex round, not pointed. Gonostylus yellow, with large spatulate lobes basally. Aedeagus wide basally, narrowed apically, curved dorsally, apex expanded. Hypandrium broadly rounded. Female. Measurements, body 13.0–13.1 mm; wing 10.0–10.5mm. Face yellowish tomentose, sparse brown setae sometimes dorsally, width at antenna 1/8 as wide as head. Scutellum yellow tomentose, sparse brown setose; pleura yellow or yellowish tomentose, anepimeral bristle yellow or brown; metacoxa largely black. Legs normal, middle coxa black; middle and hind femur with black anteroventral bristles; fore and middle tibiae black anteriorly, bristles black. Tarsi blackish or black except basal tarsomeres of fore and middle tarsi brownish-yellow; bristles black; hind tarsus with basal 2 tarsomeres wider than apical 3tarsomeres. Abdomen without unusual vestiture; segments 6–8 black, sternite 7 black setose. Terminalia (Figs. 64–66). Types. DOMINICAN REPUBLIC: La Vega prov., P.N. Armando / Bermúdez km 1.3 along trail / W of La Ciénaga 900–1100 m / 19o01.753'N 70o54.654'W / 22 June 2005 ~ S.W.Lingafelter / collector/ UV light (holotype %, USNM). Paratypes, same data as holotype (2 &, USNM); RD-251 Los Tablones – La / Laguna, P.N. Armando Ber / múdez La Vega prov., 1270 / -1,980 m. 30.vi.2004 D. / Perez (d) (1 %, MHND); RD-256 La Compartición - / Los Tablones, P.N. Armando / Bermúdez, La Vega prov., 2,450–1,110 m, 3.vii 2004, D.Perez (d) (1 &, USNM). Distribution. Captured in June and July at intermediate to high elevations (900 and 2450 m) in National Park Armando Bermúdez, La Vega province. Etymology. Latin geminus for twin, referring to the similarity of this species and O. praelongus. Remarks. Ommatius geminus is distinctive from congeners by the characters in the key, the largely black middle coxa, dense cluster of bristles in the apical corner of sternite 6 (Fig. 53), the produced sternite 7, and the combined characters of the terminalia (Figs. 59–66). Ommatius geminus is further distinguished from O. praelongus by the partially blackish metacoxa, shorter cell m3 (Fig. 46), the black palpus and proboscis, and sparse anepisternal setae. The male is also distinguished by the long sigmoid black bristle and grooved yellowish callus on the fore femur posteriorly, 2–3 apical bristles on the hind

REVIEW OF ASILIDS © 2006 Magnolia Press 53 ZOOTAXA basotarsomere, and the numerous yellow bristles on tergites 3–5 laterally. In O. 1381 praelongus, these characters are absent. The pale yellow callus is narrow and limited to the dorsal margin of the fore femur and a capitate bristle is present on the apex of the hind basomere.

FIGURES 59–63. Ommatius geminus sp. nov., terminalia. Male, 59–61. lateral, dorsal, and ventral views; 62. gonostylus, 63. aedeagus. Abbreviations: Ae=aedeagus, Ce=cercus, Ep=epandrium, Ea=ejaculatory apodeme, Gc=gonocoxa, Gs=Gonostylus, Hp=hypandrium.

FIGURES 64–66. Ommatius geminus sp. nov., terminalia. Female, 64. sternite 8; 65. genital fork, 66. spermatheca. Abbreviations: me=membranous, thinly sclerotized.

Ommatius gwenae Scarbrough (Table 1)

Ommatius gwenae Scarbrough, 1984: 139. Type locality: Constanza, Dominican Republic, type % (USNM); 1997: 12; Scarbrough & Perez-Gelabert, 2003: 48.

New Records. DOMINICAN REPUBLIC: Independencia prov., 200 m / Destacamento Aguacate, / 18o19.681'N 71o41.830'W, 1050 m, 12.viii.2006, D.Perez, R.Bastardo, B.Hierro (2 %, 2 & USNM); La Altagracia prov., trail / Guaraguao – Cueva del Puente, / 18o19.532'N 68o48.525'W, sea / level, 24.viii.2006, D.Perez, B.Hierro (day/night) (1 %, USNM); Santo Domingo Norte [Province], Sierra / Prieta, 9.viii.2006, D.Perez, / R.Bastardo, B.Hierro, S. / Medrano (night) (1 %, 1 &, USNM); La Bomba, Sierra Prieta, / Santo Domingo prov., 24. / vi.2004, R. H. Bastardo (1 %, USNM); Peravia prov., Matadero, by / aqueduct, 18o24.129’N 70o / 25.277'W, 455 m, 8.viii.2006, D.Perez, R.Bastardo (1 %, USNM); RD-248 Entrance to Sabana / Iglesia, Santiago prov., 372 m / 19o20.196'N 70o45.483'W, / 29.iv.2004, D.Perez, B.Hierro, / R.Bastardo (d) (3 %, 1 &, USNM); RD-244 Cerro Santo Tomás, / Parque Botánico de Jánico, / Santiago prov., 493 m, 19o19. / 382'N 70o49.240'W, 27.iv.2004, / D.Perez, B.Hierro, R.Bastardo (d) (2 %, USNM).

REVIEW OF ASILIDS © 2006 Magnolia Press 55 ZOOTAXA Distribution. Largely restricted to coastal plains and lowland mesic forests below 500 1381 m in elevation from May through August. Four specimens captured at 1040 m. Range includes Distrito Nacional, Hato Mayor, Independencia, Isla Beata, La Altagracia, La Vega, M.T. Sánchez, Pedernales, Peravia, Sánchez Ramirez, Santiago, San Cristóbal, San Pedro de Macorís provinces. Remarks. Ommatius gwenae is readily distinguished by the male terminalia, especially the strongly produced median process of ventral lamella (Scarbrough 1997). This wide spread species was previously recorded from May through July taken in 10 provinces from May through July. Its range now includes Independencia, Peravia, and Santiago provinces and the month of August.

Ommatius hispaniolae Scarbrough (Table 1)

Ommatius hispaniolae Scarbrough, 1984: 137. Type locality: Barahona, Dominican Republic, holotype % (MCZ); 1997: 13.

New records. DOMINICAN REPUBLIC: RD-241 Entrance to Boca / Vieja Marina, near Biyeya / beach, Azua prov., 23.iv.2004, D.Perez, B.Hierro (n) (12 %, 12 &, USNM, MHND); RD-229 – 2 km road to San / Juan, near bridge on Viajama / river, Azua prov., 294 m, 18o / 35.245'N 70o58.085'W, 20.iv. / 2004, D.Perez, B.Hierro, R. / Bastardo (d) (4 %, 8 &, MHND); Azua prov. Monte Río 18o / 24.119'N 70o42.671'W, 18 m, / 16.viii.2006, D.Perez, R.Bastardo (1 &, USNM); Barahona prov., around Basurero de Cabral, 13.viii. / 2006, D.Perez, R.Bastardo, B.Hierro (9 %, 8 &, USNM); Independencia prov., P N / Sierra de Baoruco, 1 km E of / Puerto Escondido, 450 meters / 18o19.652’N 71o34.156'W / 14 July 2004 N.E.Woodley (1 %, USNM); Independencia prov., Puerto / Escondido, 18o19.372'N, 71o / 34.014'W, 427 m, 12.viii.2006, / D.Perez, R.Bastardo, B.Hierro (13 %, 19 & USNM); RD-273 km 28, Rd. Duvergé / - Jimaní, S of Lake Enriquillo, / Independencia prov., 14.vii.2004, / D.Perez (d) (1 &, USNM); Independencia prov., Puerto / Escondido, canal on western / end of town, 18o19.442'N / 71o34.485'W, 415 m, 13.viii, / 2006, D.Perez, R.Bastardo, B.Hierro (2 %, 2 &, USNM); RD-225 100 m N Playa Buen / Hombre, Montecristi prov., / near sea level, 19o51.797'N / 71o24.181'W, 17.iv. 2004, / D.Perez, B.Hierro (d) (3 %, 3&, MHND); RD-226 2 km E Montecristi, / Montecristi prov., 44 m, 19o / 50.127'N 71o37.252'W, 17. / iv.2004, D.Perez, B.Hierro (d) (1 %, USNM); RD-259 ~ 5 km W Baní / Peravia prov., dry forest, 6. / vii.2004, D.Perez (d) (1 &, MHND); RD-249 Cerro Gordo, Baní / Peravia prov., 18o16.337'N / 70o21091'W, 28.vi.2004 / D.Perez (d) (3 %, 6 &, USNM); Peravia prov., Arroyo Grande / bridge diving Honduras / Angostura, 18o21.383'N 70o / 25.993'W, 230 m, 8.viii.2006, / D.Perez, R.Bastardo (1 %, USNM); Peravia prov., Bahía, ~5 km / W of Baní, 18o17.460'N /

56 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT 70o23.090'W, 90 m, 8.viii. / 2006, D.Perez, R.Bastardo (14 %, 10 &, USNM); RD-269 km ZOOTAXA 3 Rd. Cabo / Rojo – Aceitillar, Pedernales / prov., near sea level, 13.vii. / 2004, D.Perez 1381 (d) (1 &, MHND); RD-262 La Charca, km 30 / road Cabo Rojo – Aceitillar, / Pedernales prov., 9-vii-2004, / D.Perez (d) (4 %, 3 &, MHND); RD-267 Rio Pedernales, nr / Fuerte Banano, Pedernales / prov., 280 m, 12.vii.2004, / D.Perez (d) (1 &, MHND); Pedernales prov., Pasosena, / on rd. to Los Arroyos, 18o04. / 829'N 71o44.091'W, 69 m, / 17.viii.2006, D.Perez, R.Bastardo (2 %, 2 &, USNM); Pedernales prov., 0.7 km N / Cruce Aguas Negras, Mencia, / 18o06.956'N 71o43.038'W, 362 m, / 17.viii, 2006, D.Perez, R.Bastardo (1 %, 1 &, USNM); Pedernales prov., km 13 Rd. / Cabo Rojo, Aceitillar, 18o01. / 866'N 71o38.806'W, 123 m, / 16.viii.2006, D.Perez, R.Bastardo (night) (1 &, USNM); RD-215 ~2 km on trail to / Carlitos, Parque Nacional / Jaragua, Pedernales prov., / 124 m, 18o06.591'N 71o37. / 234'W, 7.iv.2004, D.Perez, / B.Hierro, R.Bastardo (d) (1 &, USNM); RD-227 Rd. Inoa – El Caimito, / nr San José de las Matas, San- / tiago prov., 552 m, 19o22.225'N / 71o00.661'W, 28.iv.2004, D. / Perez, B.Hierro, R.Bastardo (d/n) (1 %, 1 &, USNM); RD-243 Babosico, on way to / Jánico, Santiago prov., 515 m, /19o20.955'N 70o47.503'W, 27. / iv.2004, D.Perez, B.Hierro / R.Bastardo (d) (4 %, 4 &, USNM); Sabana Grande de Palenque, / San Cristóbal prov., 23.xii. / 2003, R.H. Bastardo (1 &, USNM); RD-232 Road San Juan –Vallejuelo, near river, San / Juan prov., 449 m, 18o41. / 754'N 71o11.725'W, 21.iv. / 2004, D.Perez, B.Hierro, / R.Bastardo (d) (3 %, 4&, MHND); RD- 233 Few kms E Valle- / juelo, San Juan prov., 704 m, / 18o40.122'N 71o18.446'W, / 21.iv. 2004, D.Perez, B. / Hierro, R.Bastardo (d) (4 %, 1 &, USNM); RD-230 ~2 km road Guanito / - Bohechio, San Juan prov., / 409 m, 18o44.383'N 71o05. / 357'W, 20.iv.2004, D.Perez, / B.Hierro, R.Bastardo (n) (1 %, USNM). Distribution. Captured between sea level and 552 m in elevation during April, July- August, and December in Azua, Barahona, Independencia, Montecristi, Pedernales, Peravia, Santiago, San Cristóbal, and San Juan provinces. Vegetation zones include scattered scrub or pine in grassy areas, and deciduous forests. Remarks. The largely pale hind tarsomere and terminalia (Scarbrough, 1997) distinguishes Ommatius hispaniolae from congeners. This lowland species is usually one of the most abundant species in collections.

Ommatius inflatus Scarbrough (Table 1, Figs. 48, 54)

Ommatius inflatus Scarbrough, in Scarbrough & Perez-Gelabert, 2003: 50. Type locality: 9 km N. Montecristi, Dominican Republic, holotype: % (FSCA).

New records. DOMINICAN REPUBLIC: RD-189 Near Laguna Cabral, / Barahona prov., 55 m, 11.xii. / 2003, D.Perez, B.Hierro, / R.Bastardo (d) (1 &, USNM); RD-233

REVIEW OF ASILIDS © 2006 Magnolia Press 57 ZOOTAXA Few kms E Valle- / juelo, San Juan prov., 704 m, / 18o40.122'N 71o18.446'W, / 1381 21.iv.2004, D.Perez, B. / Hierro, R.Bastardo (d) (1 &, USNM); RD-224, 5 km on road to / Botoncillo, Montecristi prov., / 46o220'N 71o24.001'W, 16.iv. / 2004, D.Perez, B.Hierro (d/n) (4 &, USNM); RD-186 to Playa Buen / Hombre Montecristi province, / 132 m. 19o46.213'N / 71o23. 996'W, 6–7.xii.2003, D. / Perez, R.Bastardo, A.Mar- / molejos (2 &, USNM); RD-187 near Uvero, Rd. to / Playa Buen Hombre [Montecristi province], 200 m. / 7.xii.2003, D.Perez, R.Bas- / tardo, A.Marmolejos (1 %, USNM); Republica Dominicana / Prov. Montecristi, Guayubín, / Cerro Gordo Abajo 78 m (1%, IIBZ); RD-243 Babosico, on way to / Jánico, Santiago prov., 515 m, / 19o20.955'N 70o47.503'W, 27. / iv.2004, D.Perez, B.Hierro, / R.Bastardo (d) (1 &, MHND); RD-176 Southern slopes of / Cerro Gordo, Baní Peravia prov., on way to / Salinas, 18o16.038'N 70o20. / 630'W, 23.xi.2003, D.Perez, / R.Bastardo, A. Francisco (d) (2 %, 8 &, USNM); RD-198 Matadero, start of / hills, Peravia prov., 450 m, / 18o24.464'N 70o25.736'W, / 14.xii.2003, D.Perez, B. / Hierro, R.Bastardo (d) (1 &, USNM). Remarks. Ommatius inflatus (Scarbrough & Perez-Gelabert, 2003) was originally reported from Montecristi province. Here we report it in Barahona, San Juan, Santiago, and Peravia provinces. It occurs in low and intermediate elevations on the island. The species is distinguished from congeners by the pale postocular bristles, wing venation (Fig. 48), mushroom- and pencil-like bristles of sternite 6 (Fig.54) and characters in the key.

Ommatius laticrus sp. nov. (Table 1, Figs. 49, 55, 67–74)

Male. Black, pleura entirely tomentose, hind tibia wide. Measurements, body 8.4–8.8 mm, wing 6.0–6.1 mm. Head: Face yellowish-gray to gray tomentose, setae dorsally sparse, shorter than scape and pedicel combined, about 1/2 as long as 6 brown bristles; face at antenna 1/ 6.7–1/7.4 times as wide as head. Palpus and proboscis whitish setose, latter with narrow base yellowish ventrally. Antenna black setose, one setae below pedicel about as long as flagellum; flagellum about as wide as pedicel; scape brownish-yellow. Frons yellow tomentose, brown setose. Ocellar tubercle oval, longer than wide, black setose, 2 setae slightly shorter than 3 antennal segments. Occiput yellowish tomentose with sparse setae dorsally, gray with abundant setae ventrally; 7–8 blackish, proclinate bristles dorsally, apex just beyond posterior margin of eye. Thorax. Prothorax yellowish tomentose, pale setae and bristles, sometimes blackish. Scutum brown tomentose, narrow sides and prescutellum brownish-yellow tomentose; setae sparse, brown dorsally, pale laterally; 4 lateral and 3 dorsocentral bristles black, notopleural bristle about 1/2 as long as supra-alar bristle, all thicker and most longer than dorsocentrals. Scutellum brownish-yellow tomentose, sparse brown setose, 2 blackish

58 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT marginal bristles. Pleura yellowish-gray to gray tomentose, anepisternum and ZOOTAXA katepisternum with thin, discontinous, black tomentose stripe; anepisternum sparsely 1381 setose; anepisternum and katepisternum with black bristles; anepimeron with white bristly seta; metacoxa blackish. Halter yellowish.

Wing (Fig. 49): Hyaline, apical 1/3 sparsely brownish microtrichose. Base of cell r4 beyond cell d, narrow, about 1/2 as wide as cell at wing margin. Crossvein r-m before just apical 1/3 of cell d. Cell m1 wide, base extremely narrow, about 1/5–1/12 as wide as at basal 1/3, much wider apically with vein M1 angular. Cell m3 petiolate, stalk 1/2 to about as long as rm, apex pointed. Legs. Coxae whitish tomentose with thin whitish vestiture, black except yellow fore coxae; hind coxa partly yellow. Trochanters pale yellow, except narrow apical margins and hind trochanter dorsally, latter yellow ventrally. Femora black setose, usually thin black bristles except pale yellowish seta-like bristles on hind femur ventrally. Fore and middle femora pale yellow except mostly blackish anteriorly and posterior 1/5–1/4; hind femur black except pale yellow base and incomplete wide preapical band; fore femur about as wide as hind femur, with row of 6–7 thin bristles dorsally, and shallow longitudinal groove posteriorly, about 1/2 as long as femur with pale yellow long, hooked, setae ventrally, and 2 long thin bristle basally; middle femur with thin anteroventral bristles on basal 1/2, 3 rows of short peg-like bristles on apical 1/2, and 4–5 long stout posteroventral yellowish bristles, about 1/2 as long as middle femur; hind femur narrow, about 8 times longer than wide. Fore and middle tibiae narrow, pale yellow, brownish anteriorly, apices brown, thin blackish bristles 1/2–2/3 as long as tibiae, blackish setose except yellowish ventrally; hind tibia brown, slightly narrower than hind femur laterally, black setose, short blackish bristles. Tarsi black except basal tarsomere of fore and middle basal tarsomere pale yellow, bristles and setae black; hind tarsus with basal tarsomere short, about as long as tarsomeres 2–3 combined, second tarsomere as long as wide, about same width as basal tarsomere, tarsomere 3–5 narrower. Abdomen: Black, apical margin of segments and lateral margins of tergites yellowish; black setose dorsally; sparse grayish or brownish gray tomentose, most dense on basal 4 segments, black setose except pale, often long, thin on basal segments; tergite 1, 3–5 laterally and apical corner sternite 3 with long black bristles. Sternite 3–5 (Fig. 55) with row of black bristles, those on 3 sparse, longer, those on 5–6 more abundant and shorter; apical margin of sternites 6–7 produced, with dense setae or thin bristles, those on 6 margined with sparse, apically hooked, thin bristles and tight cluster of apically angled bristles medially; sternite 6 (Fig 55) with sparse setae and 2 thin bristles laterally; cluster of abundant thin black setae basally. Sternite 8 black setose. Terminalia (Figs. 67–71): Black, black setose. Epandrium with ventral emargination apically. Aedeagus tubular apically, curved dorsally, prominent ventral spine present. Gonostylus minute. Hypandrium broadly triangular.

FIGURES 67–71. Ommatius laticrus sp. nov., terminalia. Male, 67–69. lateral, dorsal, and ventral views; 70. gonostylus, 71. aedeagus. Abbreviations: Ae=aedeagus, Ce=cercus, Ep=epandrium, Ea=ejaculatory apodeme, Gc=gonocoxa, Gs=Gonostylus, Hp=hypandrium.

Female. Measurements, body 8.1–14.0 mm, wing 7.3–9.8 mm. Face with 4–6 brown bristles; face at antenna 1/6.7–1/7.0 as wide as head. Thorax brownish-yellow tomentose;

prosternum partly to entirely yellow. Wing with base of cell m1 1/4 as wide as cell at basal 1/3. Femora and tibiae normal, not unusually wide or with unusual vestiture. Basal tarsomere of hind tarsus as long as those on anterior 2 tarsi, first and second tarsomeres

60 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT only slightly wider than third tarsomere. Abdomen with thin grayish tomentum except ZOOTAXA large oval spots laterally on tergites 3–5 and sternites 3–5 medially, black, atomentose; 1381 tergites 3–5 and sternites 4–7 with sparse, usually black, bristles laterally. Sternite 8 yellowish apically. Terminalia (Figs. 72–74).

FIGURES 72–74. Ommatius laticrus sp. nov., terminalia. Female, 72. sternite 8; 73. genital fork, 74. spermatheca. Abbreviations: me=membranous, thinly sclerotized.

Types. DOMINICAN REPUBLIC: RD-176 Southern slopes of / Cerro Gordo, Baní [Peravia prov.], on way to / Salinas, 18o16.038'N 70o20. / 630'W 23.xi.2003, D.Perez, R.Bastardo, A. Francisco (d) (holotype %, USNM). Paratypes, RD-229 –2 km road to San / Juan near bridge on Viajama / Azua prov., 294 m, 18o / 35.245'N 70o58.085'W, 20.iv. / 2004, D.Perez, B.Hierro, R. / Bastardo (1 %, USNM); RD-217-5 km S. Cabral / near road to Polo, Barahona prov.,411 m, 18o11.273.'N / 71o.14.970'W, 9.iv.2004, D. / Perez, B Hierro, R Bastardo (d) (1 &, USNM); same data as holotype (1 %, 9 &, USNM, MHND); RD-198 Matadero, start of / hills Peravia prov., 450 m / 18o24.464N 70o25.736'W / 14.xii.2003, D.Perez, B. / Hierro R.Bastardo (d) (4 &, USNM); RD-248 Entrance to Sabana / Iglesia Santiago prov., 372 m, / 19o20.196'N 70o45.483'W / 29.iv.2004, D.Perez, B.Hierro, / R.Bastardo (1 &, USNM). Distribution. Known only from the type locality. Etymology. Latin laticrus, from latus for wide and crur for shank, referring to the wide hind tibia. Remarks. Ommatius laticrus is distinguished from congeners by the characters in the

REVIEW OF ASILIDS © 2006 Magnolia Press 61 ZOOTAXA key, and combined characters of the terminalia (Figs. 67–74). The female is distinguished 1381 by the shining black spots on the sides of the tergites and long bristles along the lateral margin of the sternites.

Ommatius lucidatus Scarbrough (Table 1, Figs. 50, 56)

Ommatius lucidatus Scarbrough, 1997: 13. Type locality: N. Cabo Rojo, Pedernales province, Dominican Republic, holotype: % (CMNH). Scarbrough & Perez-Gelabert, 2003: 53.

New records. DOMINICAN REPUBLIC: RD-229 ~2 km road to San / Juan, near bridge on Viajama / river, Azua prov., 294 m, 18o35.245'N 70o58.085'W, 20.iv. / 2004, D.Perez, B.Hierro, R. / Bastardo (d) (4 %, 11 &, USNM); RD-190 ~ 5 km hacia Caseta / No. 1, Parque Nacional Sierra de / Bahoruco, Independencia prov.,/ 18o18.302'N 71o34.219'W, / 11.xii.2003, D.Perez, / R.Bastardo, B.Hierro (d) (1 &, USNM); RD-078 El Cajuil, Oviedo, / Pedernales prov.,52 mm,/ 17o48.783'N 71o21.538'W / 2.xii.2002, D.Perez, B.Hierro, / R.Bastardo (night)(1 %, USNM); Pedernales prov., P N Jaragua / trail to Carlitos ca. 6 km S of / Highway 44 106 meters / 17o48.932'N 71o28.271'W. / 9 July 2004 / N.E.Woodley (1 %, USNM); Peravia prov. Bahia ~5k / W of Baní, 18o17.460'N / 70o23.090'W90 m, 8.viii. / 2006, D.Perez, R.Bastardo (1 %, 1 &, USNM); RD-247 Rd. Inoa – El Caimito, / nr San José de las Matas, San- / tiago prov., 552 m, 19o22.225'N / 71o00.661'W, 28.iv.2004, D. / Perez, B Hierro, R.Bastardo (d/n) (1 &, USNM). Distribution. Captured at low elevations (52–475 m) during April, July, October, November, and December in Azua, Bahoruco, Independencia, Montecristi, Pedernales, Peravia, and Santiago provinces. Vegetation patterns include pastures, scrublands, and deciduous forests. Remarks. Ommatius lucidatus is distinguished from congeners by the characters in the key, especially the row of stout bristles below the middle femur, wing venation (Fig. 50), rows of stout bristles on sternites 4–5, dense pad of fine setae bordered by hooked bristles on sternite 6, and raised sternite 7 which bears a tight cluster of apically angular bristles (Fig. 56). Its range is expanded to include Peravia and Santiago provinces and the month of April.

Ommatius maculosus sp. nov. (Table 1, Figs. 47, 57, 75–82)

Male. Black body, apical 1/3 of wing brownish, anepisternum partly shiny black. Measurements, body 9.8–10.3 mm; wing 6.9–7.1 mm. Head: Yellowish-gray or gray tomentose, white setose. Face gradually slanted forward ventrally, lower 1/2 moderately

62 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT exposed beyond margin of eye, setae about as long as pedicel and scape combined ZOOTAXA dorsally, much longer ventrally, about 1/2 as long as 6 blackish bristles; face at antenna 1/ 1381 6.1–1/7.0 as wide as head. Antenna black setose; flagellum wide, tear-shaped; style with 6 setae on apical 2/3, basal 3 sequentially shorter than apical setae. Frons yellowish-gray tomentose medially, wide basally, sides slightly convergent dorsally, only slightly wider than shiny black ocellar tubercle. Ocellar tubercle with 4 black setae, 2 stout and long, each about as long as 3 antennal segments combined. Occiput yellowish tomentose dorsally, white ventrally, pale setae, most abundant ventrally, 6–7 blackish postocular bristles dorsally, stout basally, thin apically, proclinate, apex just beyond posterior margin of eye. Thorax: Prothorax grayish tomentose, whitish setose, with 2 blakish bristles dorsally. Scutum brown tomentose dorsally, postpronotal spot and margins yellowish to grayish setose; setae sparse, sparse white setae, dorsocentrals short, brown; 4 lateral and 3 dorsocentral bristles blackish, each about 1/2 as long as scutum. Scutellum yellowish-gray tomentose with sparse black setae and 2 or 4 marginal bristles, 2 short and thin. Pleura gray tomentose, anepisternum partly shining black; setae sparse, whitish, bristles whitish or dull yellow; halter yellow, base narrowly dull brownish.

Wing (Fig. 47): Hyaline, apical 1/3 brownish and brown microtrichose. Cell r4 narrow, base V-shaped, beyond apex of cell d, about 1/5 as wide as cell at wing margin. Cross-vein r-m at apical 1/4–1/3 of cell d. Base of cell m1 1/3 as wide as width at basal 1/3, slightly constricted beyond. Cell m3 petiolate, about as long as rm, apex strongly pointed. Legs: Coxae black, whitish tomentose, whitish setae and bristles. Fore and middle trochanters yellow, narrow apices blackish; hind trochanter entirely blackish. Femora yellowish, black as follows: apical 1/4 or 1/3 of fore and middle femora, and dorsally almost to base, color extending onto anterior and posterior surfaces narrowly; hind femur with base and apical 1/3 banded yellow, bands narrow dorsally, much wider ventrally; setae mostly black; fore femur posteriorly with abundant, hooked, pale setae and ventral row of long setae, shallow groove about 1/2 as long as femur; middle femur with 3 unusually long bristles anteriorly, each about 1/3 as long as femur, ventral setae moderately long, thin; hind femur with pale bristles ventrally, 2 posteroventral bristles exceptionally long, setae black, pale dorsally, numerous bristly, black, posteroventral setae present on apical 2/3; hind femur narrow, 6.0–6.4 times longer than wide. Tibiae mostly yellow, apex and anteriorly brownish, middle and hind tibiae darkest anteriorly, setae and bristles black; fore tibia with sparse long setae laterally, longest and pale on apical 2/3, thin lateral bristles about 1/2 as long as tibia, those on middle tibia black and stout; hind tibia with much shorter bristles and dense yellowish setae ventrally. Tarsi with black vestiture, bristles about 1/2 as long as tarsi, basal tarsomere on fore and middle tarsi mostly brownish-yellow, base of hind tarsus narrowly reddish; second and third basal tarsomere produced, spur-like process lateroapically.

FIGURES 75–79. Ommatius maculosus sp. nov., terminalia. Male, 75–77. lateral, dorsal, and ventral views; 78. gonostylus, 79. aedeagus. Abbreviations: Ae=aedeagus, Ce=cercus, Ep=epandrium, Ea=ejaculatory apodeme, Gc=gonocoxa, Gs=Gonostylus, Hp=hypandrium.

Abdomen: Black, apex and lateral margin narrowly yellowish; tergites 1–6 brownish- yellow or brownish tomentose, narrow apices, side of tergite 1, tergites 2–4 basally

64 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT grayish, tergites 7–8 black, sparsely tomentose; tergites 2–6 with shiny black spot laterally, ZOOTAXA spot atomentose; tergites black setose except white on tergites 1–2 laterally; bristles 1381 usually pale on tergites 1–5 laterally. Sternites (Fig 57) shiny black, atomentose or sparsely tomentose, narrow lateral margins brownish or pale tomentose, setae and bristles black; sternite 1 grayish tomentose, small oval shiny black spot medially. Sternites 3–4 slightly concave, former scattered, short setae, long black bristle in each apical corner, latter sternite with narrow plate anteromedially, reclinate setae medially. Sternite 5 with broad oval concavity apically, M-shaped ridge basally with fine setae, transverse row of stout, twisted bristles more anteriorly, often capitate; oval plate anteromedially. Sternite 6 with dense vestiture laterally, long stout bristles anteriorly, thin pencil-like bristles medially, acutely pointed thin bristles basally. Terminalia (Figs. 75–79): Black, gonostylus minute, distiphallus curved dorsally, apex tubular with flange ventrally. Hypandrium broadly rounded apically. Female. Body 10.3–10.9 mm; wing 7.3–8.0 mm. Head: Ocellar tubercle with 4–6 setae, longest 3/4 as long as 3 antennal segments combined. Middle femur with bristles short, less than 1/3 its length. Fore tibiae with short lateral fringe, much shorter than in male. Hind tarsus with basal 1/3–2/3 brownish-yellow. Abdomen with tergites 2–7 laterally and sternites 3–7 with large atomentose spots; vestiture normal, without unusual patterns. Terminalia (Figs. 80–82).

FIGURES 80–82. Ommatius maculosus sp. nov., terminalia. Female, 80. sternite 8; 81. genital fork, 82. spermatheca. Abbreviations: me=membranous, thinly sclerotized.

REVIEW OF ASILIDS © 2006 Magnolia Press 65 ZOOTAXA Types. DOMINICAN REPUBLIC: RD-215 – 2 km on trail to / Carlitos, Parque 1381 Nacional / Jaragua, Pedernales prov., 124 m, 18o06.591'N 71o37.234'W, 7.iv.2004, D.Perez, B.Hierro, R.Bastardo (holotype %, USNM). Paratypes, same data as holotype (1 %, 2 &, USNM); Pedernales prov., PN Jaragua / trail to Carlitos, 6 km. S. of / Highway 44, 106 meters, / 17o48.932'N 71o28.271'W, 9 July, 2004 N.E.Woodley (err., id. O. lucidatus) (1 %, USNM); Pedernales prov., 0.7 km N. / Cruce Aguas Negras — Mencia / 18o06.956'N 71o43.388'W, 326 m / 17.viii.2006, D. Perez, R. Bastardo (1 &, USNM). Distribution. Captured at low elevations (106–124 m) from April to August in Pedernales province. Etymology. Latin maculosus, referring to the black, oval, atomentose spot on the anepisternum. Remarks. Ommatius maculosus is distinguished from congeners by the characters in the key, vestiture pattern of the sternites (Fig. 57), and combined characters of the terminalia (Figs. 75–82). Additionally, the scutellum has thin marginal bristles in addition to a thicker, longer pair.

Ommatius nigellus Scarbough (Table 1)

Ommatius nigellus Scarbrough, 1984: 144. Type locality: Furcy, Haiti, Holotype & (AMNH); 1997: 15; Scarbrough & Perez-Gelabert, 2003: 54.

New records. DOMINICAN REPUBLIC: RD-266 Las Abejas, P.N, / Sierra de Bahoruco, Peder- / nales prov., 1,310 m, / 18o09.011'N 71o37.342'W. / 11.vii 2004, D.Perez (d/n) (10 %, 4 &, USNM, MHND). Distribution. Collected in July from 350 to 1310 m in Ouest District, Haiti, and Pedernales and Barahona provinces, Dominican Republic, in southern Hispaniola. These areas are predominately composed of scrubthorn vegetation and mixed forest. Remarks. The abundant, long, black setae of the face, scutum, scutellum, and fore coxae, wholly black legs, absence of scutellar bristles, and combined characters of the terminalia distinguish Ommatius nigellus (Scarbrough 1997)

Ommatius praelongus sp. nov. (Table 1, Figs. 51, 58, 83–90)

Male. Black, pleura entirely tomentose, yellow setose, unsually long discal and m3 cells. Measurements, body 11.8–13.5 mm; wing 9.5–10.5 mm. Head: Yellowish tomentose, yellow setose. Face narrow, sparsely setose dorsally, mystax dense, about as long as proboscis, 4–6 black bristles present; face at antenna 1/9 as wide as head. Palpus yellow

66 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT apically, brown basally. Proboscis black, narrow base yellow ventrally, sparsely setose ZOOTAXA basally. Antenna with scape and pedicel yellowish, black setose, scape ventrally, 1381 sometimes yellow setose partially dorsally; flagellum blackish, about as wide as pedicel; style with several ventral setae extending from base to apex, none exceptionally short. Frons parallel sided, not convergent dorsally. Ocellar tubercle, longer than wide, about 1/2 as wide as vertex, with 5–6 short yellowish or yellowish and black mixed setae, 2 blackish setae slightly longer than scape and pedicel combined. Occiput yellowish-gray tomentose, vestiture pale, setae most abundant ventrally, postocular bristles thin, yellowish, sometimes one or 2 blackish, apices just beyond posterior margin of eye. Thorax: Prothorax grayish tomentose, vestiture pale, one or 2 bristles blackish; prosternum yellow. Scutum brownish or brownish-yellow tomentose dorsally, wide apical and lateral margins yellowish-gray or grayish; setae sparse, blackish dorsally, pale laterally; 4 lateral bristles and 3 dorsocentral bristles blackish, long, thin, notopleural bristle short, about 1/3 or 1/2 as long as supra-alar and postalar bristles, dorsocentral bristles, thinner, shorter than lateral bristles. Scutellum sparsely yellow setose, 2 marginal bristles yellow or mixed brown, sparse, thin, seta sometimes present. Pleura entirely tomentose, yellowish or white, vestiture white or yellowish, anepisternum with abundant setae posteriorly, anepimeron with 2–3 bristly setae; metapleuron and metacoxa yellow. Halter yellowish.

Wing (Fig. 51): Hyaline, cells r4 and m1 short; base of r4 at apex of cell d, about 1/2 as wide as cell at wing margin, veins divergent just before apex; cell m1 divergent apically, much wider at wing margin, base of M1 curved or angular. Cells d and m3 narrow, unusually long, former slightly less than 1/2 as long as wing, latter about 1/3; r-m crossvein beyond middle of cell d; cells d and m3 and narrow base of m1 opaque; M2 short, joining base of cell m3, Cu joins M3 slightly beyond base of cell m3. Legs: Coxae yellow, base of middle coxa narrowly blackish, grayish tomentose, vestiture yellowish, apical bristles stout. Trochanters yellow, narrow margins brown. Femora and tibiae mostly yellow, largely blackish setose, bristles largely thin, yellowish. Fore femur slightly brownish anteriorly; apical 1/3 twice as wide as base, somewhat flat, concave anteriorly with abundant short pale setae, swollen posteriorly with 3 black bristles dorsally, abundant whitish hooked setae ventrally, one blackish basomedially. Middle and hind femora with long blackish spot dorsally, color extending ventrally onto anterior and ventral surfaces, spot about 1/2 as long as femora; middle femur with row of 4–5 black bristles anteriorly, basal 2 contrastingly long. Hind femur narrow, 6–7 times as long as wide; 2 long yellowish posteroventral bristles basally. Tibiae with long yellowish bristles, sparse short bristles black; fore and middle tibiae yellow, basal 1/3 and apical 1/4 of hind tibia yellow, blackish medially. Tarsi black setose, most bristles black, 2–3 bristles on fore and middle tarsi yellowish; fore and middle tarsi yellowish, darker brownish-yellow or brownish apically; hind tarsus yellowish-brown or blackish basally, basal tarsomere swollen, about 2 times as wide as second tarsomere, stout capitate bristle apically.

FIGURES 83–87. Ommatius praelongus sp. nov., terminalia. Male, 83–85. lateral, dorsal, and ventral views; 86. gonostylus, 87. aedeagus. Abbreviations: Ae=aedeagus, Ce=cercus, Ep=epandrium, Ea=ejaculatory apodeme, Gc=gonocoxa, Gs=Gonostylus, Hp=hypandrium.

Abdomen: Black, narrow apical margins of basal 4–5 segments and lateral margins of tergites yellow, sparse yellowish or yellowish-gray tomentose, mostly yellow setose; tergites dorsally black setose, segments 6–8 black. Sternites 1–4 and 7–8 yellow setose, thin long bristles on apical 1/2 of sternites 3–4; apical 1/2 of sternite 5 with short, thin, brown bristles, each slightly bent apically, diameter about same except slightly pointed at

68 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT apex. Sternite 6 with abundant short setae, slightly bent apically, apex blunt medially, latter ZOOTAXA margined with thinner, longer, yellow setae; apical 1/2 extremely dense mass of long, thin, 1381 brown or brownish-yellow bristles laterally, apices strongly curved medially, mass often concealing base of sternite 7. Sternite 7 with long thin yellow bristles laterally, apical bristles shorter. Terminalia (Figs. 83–87): Black, yellow and black setose. Epandrium narrow, apex round, not pointed. Gonostylus yellow, with large spatulate lobes basally. Aedeagus with distiphallus curved dorsally, tubular, with prominent ventral flange. Hypandrium broadly rounded. Female. Measurements, body 11.3 mm; wing 9.5 mm. Face with sparse vestiture. Pedicel brownish apically. Frons slightly convergent dorsally. Ocellar tubercle with 2 setae as long as pedicel and scape combined. Pronotum with thin, white bristles. Marginal bristles black. Pleura grayish tomentose. Wing entirely hyaline, veins Cu and M fused just before apex of cell d. Legs normal with only thin bristles; brown color darker, often extending completely across anterior surface or connecting ventrally as band on hind femur. Tibiae with only black bristles, narrow apex of fore and middle tibiae brownish, hind tibia with apex narrowly brownish-yellow or reddish. Tarsi darker brown, hind tibiae almost entirely brown or blackish, bristles black, basal tarsomere of hind tarsus wider than second tarsomere, stout, apically capitate bristles absent. Abdomen normal, without unusual vestiture; apical sternites with scattered black setae. Terminalia (Figs. 88–90). Sternite 8 longer than wide. Genital fork mostly membranous, narrow dorsal arms sclerotized. spermatheca oval.

FIGURES 88–90. Ommatius praelongus sp. nov., terminalia. Female, 88. sternite 8; 89. genital fork, 90. spermathecae. Abbreviations: me=membranous, thinly sclerotized.

REVIEW OF ASILIDS © 2006 Magnolia Press 69 ZOOTAXA Types. DOMINICAN REPUBLIC: RD-266 Las Abejas, P.N. / Sierra de Bahoruco, 1381 Peder- / nales prov., 1,310 m, / 18o09.011'N 71o37.342'W, / 11.vii.2004, D.Perez (d/n) (holotype %, USNM). Paratypes, same data holotype (8 %, 1 &, USNM, MHND). Distribution. Captured at intermediate elevations (1310 meters) during July in Pedernales province. Etymology. Latin adjective praelongus for 'elongated', referring to the long, narrow d

and m3 cells of the wing. Remarks. Ommatius praelongus is distinguished from congeners by the characters in the key, the yellowish metapleuron and metacoxa, the posterior patch of long setae on the anepisternum, the exceptionally enlarged basal tarsomere and capitate bristle on the hind tarsus, distinctive vestiture of sternites 4–7 (Fig. 58), and combined characters of the terminalia (Figs. 83–90).

Ommatius lucidatus species group

The 4 new species, together with 4 earlier species (Scarbrough & Perez-Gelabert, 2003) and 2 extinct species (Scarbrough & Poinar, 1993; Scarbrough, 2001), form a distinct cluster of species here designated the lucidatus species group. The group is distinguished by the following combination of characters; 1) slender body with thin legs; 2) face at the base of the antenna >1/7 as wide as the head; 3) the fore femur (%) with pale yellow hooked setae posteriorly; 4) the tibial and tarsal bristles unusually long, at least 2/3 as long as the leg part; 5) the hind basotarsomere, sometimes the second basotarsomere, is swollen, about 1/4 wider than tarsomeres 3–5; 6) sternites 6–7 (%) with complex pattern of rows and clusters of setae and bristles, bristles often compressed or close spaced forming dense mass or angular structures; 7) setae and bristles are often modified apically, i.e. pencil-shaped, only slightly tapered apically; hooked or angled about 45 degrees; apex round, not acuminate; capitate as a knob or flat horizontal plate; apex flattened vertically, wider than shaft and with notched apex; 8) gonostylus minute, aedeagus tubular, horizontal, slightly curved dorsally; 9) sternite 8 (&) longer than wide with medial fissure apically; and 10) the oval spermatheca. The following key will separate the known species of this group from the remaining Hispaniolan Ommatius species [see Scarbrough & Perez- Gelabert, 2003 for key]. Although members of this group are presently known only from the Dominican Republic, it is reasonable to assume that they occur across Hispaniola.

Key to the known species of the Ommatius lucidatus species group from Hispaniola

1. Anepisternum with stout bristles; prosternum yellow; cell m3 long, cu-m basally (Figs. 45, 46, 51); ocellar tubercle narrow, 1/4 or 1/3 longer than wide; fore femur (%) wider

70 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT than hind femur vertically; body >12 mm...... 2 ZOOTAXA - Anepimeron without stout bristles thin, if present seta-like; prosternum usually black; 1381

cell m3 short, petiolate basally (Figs. 48–50); ocellar tubercle about as wide as long; fore femur narrow (%), about as wide as hind femur vertically; body <12 mm ...... 4 2. Palpus brown, proboscis entirely dark, narrow base yellowish-brown ventrally; hind basotarsomere black; fore femur (%) with wide, scar-like, vertical groove posteriorly; hind tibia black, clavate (%), apex twice as wide as base; hind basotarsomere (%) with 3, black, blunt-tipped bristles apically; tergites 2–6 with abundant, erect, yellow bristles laterally (%); sternite 6 with dense mass of long thin bristles (Fig. 53), sternites otherwise with simple vestiture; sternite 8 strongly produced; wing (%) and terminalia as in Figs. 46, 59–66 ...... geminus sp. nov. - Palpus at least partly yellow; narrow base of proboscis yellow ventrally; fore femur (%) without wide scar-like, vertical groove posteriorly; hind tibia slender, only slightly wider apically than basally; hind basotarsomere and abdomen without combination of characters above ...... 3 3. Palpus yellowish apically; hind tibia partially brown, basal 1/3 and apical 1/4 yellow; % cell r4 at apex of cell d, m1 unusually short, cells d and m3 unusually long, opaque ( ) (Fig. 51); fore tibia with several short, spur-like bristles laterally, long bristles absent; hind basotarsomere with capitate bristle apically; sternites 4–7 with abundant, long thin bristles, sometimes forming dense masses, and flat, apically angled, bristles (Fig. 58); terminalia as in Figs. 83–90...... praelongus sp. nov. - Palpus entirely yellow; hind tibiae mostly dark, base yellowish posteriorly; costal cell

entirely and apical 1/3 of wing brownish, base of cell r4 beyond apex of cell d, cells m1,

d, and m3 not unusually short or long (Fig. 45); fore tibia with long bristles laterally, sometimes twisted (%); hind basal tarsomere without capitate bristle apically; sternite 6 with dense mass of short capitate bristles medially and long, thin, brown bristles in apical corner, apex flat, wider than base, notched (Fig. 52)...... elusivus Scarbrough 4. Palpus brown setose; metacoxa entirely and hind basotarsomere partly yellow basally; tergites densely yellowish tomentose; fore femur (%) swollen with wide, shallow, vertical groove posteriorly, dorsal row of bristles pale, flat basally; middle femur with wide, shallow, vertical groove anteriorly; sternite 6 (%) with thin, brown, pencil-like and short, yellow, mushroom-like bristles (Fig. 54); terminalia (%) yellow or brownish-yellow; fore femur (&) yellow posteriorly, narrow apex sometimes blackish, fore coxa black anteriorly, yellow posteriorly… ...... inflatus Scarbrough - Palpus pale setose; metacoxa partly and hind basotarsomere dark; tergites mostly sparsely tomentose, mostly subshiny or shiny, if tomentum dense then limited to basal 3 tergites; fore and middle femora (%) without shallow vertical grooves; sternite 6 (%) without combination of characters above; terminalia (%) black; fore femur (&) in part black posteriorly, fore coxa variable ...... 5 5. Anepisternum entirely tomentose; wings hyaline; fore coxae usually yellow [lucidatus

REVIEW OF ASILIDS © 2006 Magnolia Press 71 ZOOTAXA sometimes blackish]; middle femur (%) with 4–5 unusually long, thick, posteroventral 1381 bristles; fore tibia sometimes with fringe of thin setae laterally; hind basotarsomere blackish; apical corners of sternite 5 (%) with sparse, black, hooked bristles ...... 6 - Anepisternum partly atomentose with black oval spot; wings blackish apically; middle femur of male with only thin posteroventral bristles; fore tibia with fringe of thin setae laterally; fore coxa black; hind basotarsomere yellowish-red; sternite 6 (%) with abundant bristles laterally, those apically contrastingly long and stout (Fig. 57); terminalia as in Figs. 75–82 ...... maculosus sp. nov. 6. Fore and middle femora mostly yellow, apical 1/3 diagionally black; fore coxa usually black, sometimes partly yellow anteriorly, prosternum black; middle femur (%) with long, stout bristles ventrally; sternite 7 (%) with narrow, apically angled, cluster of compressed bristles arising from raised medioapical tubercle, sternites 4–5 with long bristles laterally, usually pale yellow (Fig. 56) ...... lucidatus Scarbrough - Fore and middle femora anteriorly and apical 1/3 posteriorly black; fore coxa and prosternum yellow; middle femora with or without stout bristles ventrally...... 7 7. Tarsi black, at most basotarsomere of fore and middle tarsi yellow; middle femur (%) with long, stout bristles and numerous, black, spur-like bristles ventrally; fore tibia (%) with sparse long setae; hind tibia as wide as hind femur in lateral view; apical margin of sternites 6–7 (%) strongly produced and with abundant bristles (Fig. 55); epandrium emarginate ventroapically; terminalia as in Figs. 67–74) ...... laticrus sp. nov. - Fore tarsus with basal two tarsomeres and middle tarsus with all tarsomeres yellow, narrow apices black; middle femur (%) with only thin, short bristles ventrally, spur- like bristles absent; fore tibiae (%) with fringe of mumerous long, wavy black and yellow setae; hind tibia narrow, less than 2/3 as wide as hind femur; apical margin of sternites 6–7 (%) normal, not strongly produced or with abundant bristles; epandrium angular apically, without an emargination ...... flavescens Scarbrough

Subfamily Stenopogoninae

Tribe Stenopogonini

Plesiomma Macquart

Plesiomma Macquart, 1838: 54. Type species: testacea Marquart, designated by Back, 1909: 306.

Remarks. This genus is limited to the New World with at least 14 species of which at least 5 are found in the Greater Antilles. The species are black, yellowish, or reddish with elongated, slender, wasp-like bodies with red or brown wings and are usually associated with low growing vegetation at low altitude. The genus is distinguished by the unusually wide head and converging face and frons so that the vertex is quite narrow. The male

72 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT terminalia is rotated about 90–120 degrees. The female terminalia have long spoon-shaped ZOOTAXA spines; sternite 8 has a median cleft apically. 1381

Key to Plesiomma in Hispaniola

1. Oral margin with 3 or 4 rows of abundant bristles; scape swollen, 3/4 as wide as long; 4 marginal scutellar bristles usually present; hind femur brownish-red or brownish- yellow; terminalia (Figs. 98–104); blackish species ...... inflatum Hull - .Oral margin with single row of bristles; scape not swollen, about 1/2 as wide as long; 2–3 marginal scutellar bristles present; hind femur mostly yellow, brownish-yellow or red; yellow and reddish species...... 2 2. . Flagellum black; frons dark brown tomentose; vertex wide, about 1/2 as wide as face at antenna; fore coxae blackish anteriorly, vestiture black, red or yellow; hind femur and tibiae brownish-yellow or yellow; pulvilli sooty gray; terminalia (Figs. 91–97) ...... angustum (Macquart) - .Flagellum reddish; frons yellow or yellowish-gray tomentose; vertex narrow, 1/6–1/4 as wide as face at antenna; fore coxae yellow, vestiture only reddish or yellow; hind femur and tibiae red; pulvilli yellow; terminalia (Figs. 105–111, 122) ...simile sp. nov.

Plesiomma angustum (Macquart) (Table 1, Figs. 91–97)

Dasypogon angustum Macquart, 1848: 20. Type locality: Haiti, type: ? Plesiomma angustum (Macquart). Hull, 1962: 219; Martin & Papavero 1970: 21. Catalogue.

Redescription, male. Brownish-yellow or yellow. Measurements, body 11.6–13.1mm; wing 8.9–9.0 mm. Head: Black dorsally, brown or brownish yellow ventrally. Face brownish-yellow tomentose; oral margin with one row of yellow or reddish bristles. Palpus yellow, yellow setose. Proboscis black, middle 1/2 ventrally yellowish; yellowish setose. Antennae black setose; scape and pedicel yellow, about same length, scape not unusually swollen; flagellum black with sparse black setae dorsally, about as long as pedicel and scape combined, about twice as long as either scape or pedicel. Frons and ocellar tubercle dark brown tomentose, slightly yellowish at some light angles; ocellar setae black. Vertex wide, about 1/2 as wide as face at antennae; vertex posteriorly yellowish tomentose, narrow brown tomentose stripe from base of ocellar tubercle posteriorly sometimes present. Occiput yellow tomentose, grayish medially and ventrally; postocular bristles and setae yellow, ventral setae pale yellow. Thorax: Prothorax mostly black, brown or yellowish-brown tomentum, propleuron yellow, bright yellow tomentose; stout yellow bristles dorsally, whitish long setae laterally.

REVIEW OF ASILIDS © 2006 Magnolia Press 73 ZOOTAXA Scutum with 3 black stripes dorsally, blackish or brown tomentose, lateral stripe shorter 1381 than median stripe and divided transversally and separated from median stripe by wide yellow or golden yellow tomentose line; median stripe extends forward to pronotum and divided longitudinally by faint, narrow, yellowish-brown tomentose line; ground color yellow on each side of median stripe anteriorly and wide margin laterally; vestiture mostly yellow or reddish, stout bristles mostly black or black tipped; setae scattered anteriorly and laterally in yellow areas, longitudinal row of short acrostichal setae and bristly dorsocentral setae present, 2–5 of latter long, often blackish. Scutellum and mediotergite black, brown tomentose; 2–3 marginal scutellar bristles present, reddish or black tipped. Mesopleuron mostly black, dorsoposterior corner of katepisternum, anepisternum posteriorly, anatergite, katatergite, and metapleuron yellow, latter usually bright yellow tomentose; wide dark brown tomentose stripe extends across the anepisternum and katepisternum anteriorly, yellowish-gray tomentose posteriorly; katatergal bristles thin, either yellow or black. Halter yellow. Wing: Dense blackish microtrichose, darkest anteriorly; microtrichia brownish present medially. Veins mostly brown, costa black, veins at base of wing yellowish, color often extending toward middle of wing. Legs: Mostly yellow, black setose, bristles yellow or reddish. Coxae mostly yellow and yellowish tomentose, fore and middle coxae anteriorly and hind coxa laterally black except narrow apex; fore coxa blackish tomentose anteriorly; vestiture pale with sparse black setae. Trochanters blackish apically, yellow or yellowish brown basally. Basal 1/2 of fore and middle femora with blackish anteroventral stripe; hind femur and hind tibia with preapical bands or anterior black spot. Tarsi yellow, apical tarsomeres slightly brownish- yellow; black setose, and black and reddish bristles. Claws black, pulvilli gray. Abdomen: Spatulate, tergites 6–7 only slightly wider than junction of 2–3 yellow, black setose. Tergites 1–4 or 5 sooty black basally; narrow lateral margins of tergites 2–8 black; tergites 6–8 with short black line medially. Setae short dorsally, long on sternites and tergites 1–2 laterally, latter bristly. Terminalia (Figs. 91–95): Rotated 90o, yellow with hypandrium black basally, black setose. Female. Measurements, body 13.3–15.1 mm; wing 9.9–11.2 mm. Identical to male except narrow apical margins of segments 6–8 black. Terminalia (Figs. 96–97). Sternite 8 medially with long, thin setae. Genital fork Y-shaped. Spermathecae long, extending to base of segment 5, each narrow, darkly sclerotized, tightly coiled 7 times apically. New records. DOMINICAN REPUBLIC: prov. La Altagracia, / Nisibon, 4–7.v.78 / beach under coconuts ~ G.B.Fairchild & / R.E.Woodruff / insect light trap (1 %, 1 &, FSCA); RD-187 near Uvero, Rd. to / Playa Buen Hombre [Montecristi province], 200 m. / 7.xii.2003, D.Perez, R. Bas- / tardo, A. Marmolejos (7 %, 4 &, MHND, USNM); RD-264 Km. 25 rd Cabo Rojo / - Aceitillar, Pedernales prov. / 678 M, 10.vii.2004, D.Perez (1 &, USNM).

FIGURES 91–95. Plesiomma angustum (Macquart), terminalia. Male, 91. dorsal view; 92. gonocoxa, 93. hypandrium; 94 gonostylus; 95. aedeagus. Abbreviations: Ce=cercus, E p=epandrium, Ea=ejaculatory apodeme, Gc=gonocoxa, Gs=Gonostylus.

Distribution. Captured during May, July, and December in La Altagracia, Montecristi, and Pedernales provinces, Dominican Republic, and in Haiti.

FIGURES 96–97. Plesiomma angustum (Macquart), terminalia. Female, 96–97. dorsal and ventral views. Abbreviations: Ac Sp= acanthophorite, Ce=cercus, Gf=genital fork, S8=sternite 8, S10=sternite 10, me=membranous, thinly sclerotized, T8=tergite 8, T9=tergite 9, T10=tergite10.

Remarks. Plesiomma angustum is distinguished from congeners by the characters in the key, the proboscis is mostly yellow ventrally, and the combined characters of the terminalia (Figs. 91–97). This is the first report of P. angustum from Hispaniola since its original description in 1848. Although, the type specimen has been 'lost' (Martin & Papavero, 1970), these specimens have a quite narrow abdomen as its name implies. A comparison of these Hispaniolan specimens with valid West Indian Plesiomma species suggests that it is endemic to Hispaniola. Though we are uncertain of the type locality in Haiti (possibly Port-au-Prince vicinity), new material adds La Altagracia, Montecristi, and Pedernales provinces, Dominican Republic, to its distribution. Collecting records suggest that it occurs at low elevations (sea level to 700 m).

Plesiomma inflatum Hull (Table 1, Figs. 98–104)

Plesiomma inflatum Hull, 1962: 219. Type locality: Haiti, holotype: % (MCZ). Martin & Papavero, 1970: 22. Catalogue.

Hull (1962) description of the species is quite good but was derived from a single male. Here we add notes from Dominican Republic specimens, report the female of the species

76 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT for the first time, and illustrate the terminalia. ZOOTAXA Male. Measurements, body 13.5–13.7 mm; wing 9.0–9.3 mm. Head: Ground color 1381 mostly blackish, brownish-yellow as follows: face at antenna continuing ventrally along margin of eye and below oral margin medially, and scape and pedicel. Tomentum usually brown, appearing black at some angles on frons, vertex, oral margin medially, and occiput dorsally; occiput mostly yellowish-gray, below oral margin and base of proboscis below head gray. Color of bristles and setae variable, black, reddish, mixed red and black, or bicolored with red and black. Oral margin swollen with 3–4 irregular rows of stout bristles. Palpus yellow or brownish-yellow, apex black, setae yellowish or pale reddish. Antenna usually black setose; scape swollen, slightly longer than wide, about as long as pedicel; basal flagellomere flat, wide, base about 1/2 as greatest width, 1.5–1.7 times as long as scape; apical flagellomere usually slightly less than 1/2 as long as basal flagellomere. Vertex narrow, 1/4–1/3 as wide as face at antenna. Occiput sparsely setose, most abundant along margin of eye. Thorax: Prothorax, scutum, and scutellum mostly black with black tomentum, sides of scutum including postpronotum and postalar callus and corners of scutellum pale brownish-yellow with grayish tomentum. Black tomentose dorsum of scutum divided longitudinally by yellowish lines. Vestiture usually black, sometimes bicolored or reddish; scutum with dorsocentral, lateral, and marginal scutellar bristles stout; setae sparse, scattered. Scutellum usually with 4 marginal bristles, sometimes only 2. Mediotergite black with blackish tomentum, anatergite pale brownish-yellow. Meso- and metapleura brownish-yellow, anepisternum anteriorly blackish with black tomentose, katepisternum brownish-yellow with similar colored tomentum; immediately below wing base ventrally to the middle and hind coxae gray tomentose, metapleuron and katepisternum brownish- yellow. Wing: Brown microtrichose, pale brown posteriorly; yellowish spot from Rs ventrally across apex of bm and base of discal cells. Legs: Vestiture mostly black, stout bristles often red. Coxae and trochanters brownish- yellow, brownish-yellow tomentose, except middle coxae gray; scattered setae anteriorly, stout bristles apically. Femora yellowish-brown or brownish-red, narrow apices yellow; fore and middle femora ventrally black, hind femur with small preapical blackish spot ventrally; sparse bristles present. Tibiae yellowish-brown, narrow apices yellow, bases yellowish to brownish-yellow, extending to basal 1/5–1/4 anteriorly; several stout bristles present. Tarsi yellowish-brown, narrow apices of basal 3–4 tarsomeres yellow; bristles black and reddish. Claws and pulvilli blackish. Abdomen: Slightly spatulate, junction of segments 2–3 narrowest, segments 5–6 only slightly wider, by 1/5. Ground color sooty brownish-yellow, wide base of tergite 2, narrow margins of 2–4 yellow; tergite 1, narrow median stripe of tergites 2–7, and tergites laterally, black; setae mostly black, short on tergites dorsally, thin and long on basal 2 tergites laterally and on all sternites.

FIGURES 98–102. Plesiomma inflatum Hull, terminalia. Male, 98. dorsal view; 99. gonocoxa, 100. hypandrium; 101 gonostylus; 102. aedeagus. Abbreviations: Ce=cercus, Ep=epandrium, Ea=ejaculatory apodeme, Gc=gonocoxa, Gs=Gonostylus.

Terminalia (Figs. 98–102): Rotated 90o, sooty brownish-yellow, black setose. Female. Pale ground color in male distinctly yellow or much vibrant brownish-yellow, sooty color generally scarce in females; stout bristles mostly red or bicolored. Measurements, body 13.7–17.0 mm; wing 10.0–12.1 mm. Face at antenna 2.4–3.0 times

78 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT diameter of vertex immediately behind ocellar tubercle. Antenna with scape 1.4–1.5 ZOOTAXA longer than wide, about as long as pedicel; basal flagellomere 1.5–1.7 times as long as 1381 scape; apical flagellomere usually 2/5 as long as basal flagellomere. Prescutellum brownish-yellow or yellow. Marginal scutellal bristles 2–6, usually 4. Coxae, femora, tibiae, and tarsi pale brownish-yellow ground color; fore and middle femora black ventrally; hind femur without preapical blackish spot. Slightly spatulate, segments 6–7 about 1/6 as wide as junction of 2–3. Segments 2–4 yellow with sides black, median black stripes on tergites absent; segments 5–8 becoming increasingly brown, narrow apical margin black, segment 8 dark brown or black. Terminalia (Figs. 103–104). Sternite 8 with short, thick setae, medially. Genital fork V-shaped; spermatheca long, extending to segment 4, extremely narrow apically, coiled 5 times.

FIGURES 103–104. Plesiomma inflatum Hull, terminalia. Female, 103–104. dorsal and ventral views. Abbreviations: Ac Sp=acanthophorite, Ce=cercus, Gf=genital fork, S8=sternite 8, S10=sternite 10, me=membranous, thinly sclerotized, T8=tergite 8, T9=tergite 9, T10=tergite10.

New records. DOMINICAN REPUBLIC: La / Vega [province], Cordillera Central / 4.1 km SW El Convento / 18-50-37N, 70-42-48 W [18o50.37'N 70o42.48'W] / 1730 m, 31 May 2003 ~ J.Rawlings, R.Davidson, / C.Young, C.Nuñez, P. / Acevedo, dense secondary / evergreen forest with pine, hand collected / Sample 22242 (3 %, CMNH); La / Vega [province], Cordillera Central / 4.1 km SW El Convento / 18-50-37N, 70-42-48W [18o50.37'N, 70o42.48'W] / 1730 m, 31 May 2003 ~ J.Rawlings, R.Davidson / C.Young, C.Nuñez, P. / Acevedo, dense secondary / evergreen forest with / pine, hand collected. / Sample 22242 (1 &, CMNH); RD-275 La Ciénaga – Los / Tablones, PN Armando / Bermúdez La Vega prov./ 19o04.044'N 70o51.789'W / 1100–1270 m 17.vii.2004 / D.Perez (d) La Vega prov. PN Armando / Bermúdez trail / La Ciénaga de / Manabao Los Tablones, 20. / viii.2006, D.Perez, Ruth Bastardo, B.Hierro (1 %, USNM); prov. La Vega / Constanza 31.v.1986 / R.Miller & L.Stange (1 %, FSCA); San / Juan prov. Sierra de

REVIEW OF ASILIDS © 2006 Magnolia Press 79 ZOOTAXA Neiba, trail to Sabana del Silencio, 11 km / SSW of El Cercado, 1700–1900 m / 1381 19o39'.427'N, 71o32.475'W / 10 July 2006 N.E. Woodley (1 %, USNM); San / Juan [province], Sierra de Neiba, / Sabana del Silencio, / 10.0 km SSW El Cercado / 18-39-07 N, 71-33-21W [18o39.07'N, 70o33.21'W] / 2009 m, 20 June 2003 ~ J. Rawlings, C.Nuñez, R. / Davidson, C.Young, P. / Acevedo, N. de la Cruz / cloud forest along Dan- / thonia savannah, hand / collected, sample 33242 (2 &, CMNH); Peravia prov. Bahia, ~ 5 km / of Baní 8o16.460'N 70o23.090'W 90 m, 8.viii. / 2006 D.Perez, R.Bastardo (1 &, USNM); 2123, 2133, 2135, 2144, 2149 [no data] (3 %, 2 &, IIBZ); prov. Santiago / Par. Nac. Armando Bermudez / trail to Pico Duarte, / 1780 m, 29JULY1993 / D. Sikes & R. Rosenfeld (1 %, MAIC). Distribution. Captured at intermediate and high elevations during May to July in La Vega, Preavia, San Juan, and Santiago provinces in the Dominican Republic and Haiti. Remarks. As in the previous species, P. inflatum is endemic to Hispaniola. In contrast, it occurs at higher elevations (900–2009 m). The type locality, Mount Basil in northern Haiti, is approximately 1500 meters in elevation. We record the species in the Dominican Republic for the first time. Here it is associated with dense secondary evergreen forests and cloud forest in the Cordillera Central and the Sierra de Neiba mountains. Plesiomma inflatum is distinguished from congeners by the dark, sooty body, swollen scape, and terminalia (Figs. 98–104). However, the male is usually significantly darker than the female. Abdominal tergites 2–4 of females are brownish-yellow or yellow with narrow black lateral margins. Tergites 5–8 are usually progressively darker from yellowish-brown to reddish-brown with the terminal segment darkest. The sides and narrow apical margins are black. Plesiomma inflatum is most similar to P. funestum Loew in the black body. However, it is distinguished from the latter by the multiple rows of bristles on the oral margin, the blackish legs, and combined characters of the terminalia (Figs. 98–104). Plesiomma funestum has a single row of sparse bristles along the oral margin, the legs are entirely yellow, and the terminalia of the male is distinctive. These new records extend the distribution of P. inflatum across Hispaniola.

Plesiomma lineata (Fabricius) (Table 1)

Asilis lineatum Fabricius, 1781: 465. Type locality: St. Croix, Virgin Islands, type: &? (ZMUC). Dasypogon lineatus Fabricius, 1805: 13. Plesiomma macra Loew, 1861: 35. Type locality: Cuba, type: %. Plesiomma lineata (Fabricius). Back, 1909: 307; Bromley 1929: 276; Hull, 1962: 219; Martin & Papavero, 1970: 22. Catalogue.

Remarks. The identity of P. lineata is problematic because the type specimen is essentially destroyed, with only the thoracic dorsum surviving (Martin & Papavero, 1970). The

80 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT species has three easily recognized black stripes on the dorsum of the scutum. Thus, ZOOTAXA museum specimens and authors usually identified specimens with such markings as P. 1381 lineata. Unfortunately, most of the West Indian species have these stripes. Hull (1962) grouped Plesiomma into the subgenera Boropis Hull, Dapsilochaetus Hull, and Cystoprosopa Hull. Plesiomma lineata was designated the type species of Boropis Hull to include those species that lacked marginal scutellum bristles. Marginal scutellar bristles in this genus are often thin [with small sockets], readily subject to being broken. All Antillean species that we examined have marginal scutellar bristles. Bromley (1929) reported P. lineata from Cuba. Later, Farr (1963) reported it from Jamaica based on Bromley's paper. Yet the type locality of Plesiomma lineata is St. Croix, Virgin Islands, 'Habitat in Insula St. Crux Americes. Dom. Vahl.' (Fabricius, 1781; see Fabrician dipterous types in Thompson, 1981; Zimsen, 1964). Bromley (1929) listed Plesiomma macra Loew as a synonym of P. lineata. The type locality of the species is Cuba but its identity is also uncertain because the type is lost (Martin & Papavero, 1970). Furthermore, we have not located Bromley's specimens of P. lineata (=P. m a c ra ) from Cuba or Farr's P. li ne a ta from Jamaica for study. Illustrations of the male terminalia of the Jamaican P. li ne a ta distinguish it from the 3 Hispaniolan species. Since there are no authenticated records of P. lineata from Cuba, Jamaica, or Hispaniola and its distribution is thus far limited to St. Croix, P. m a cr a is removed from synonymy. Distribution. St. Croix, U.S. Virgin Islands.

Plesiomma simile sp. nov. (Table 1, Figs. 105–111, 122)

Male. Reddish. Measurements 21.3 mm; wing 14.5 mm. Head: Face yellow, yellowish- gray tomentum, oral margin with single row of stout reddish-yellow or yellow bristles. Palpus yellow, yellow setose. Proboscis reddish, yellow setose. Antenna reddish-yellow, flagellum and microsegments darkest, yellow setose; pedicel slightly longer than scape, basal flagellomere slightly longer than scape and pedicel combined, apical flagellomeres 1/2 as long as basal flagellomere. Frons reddish, tomentum yellow. Ocellar tubercle with sparse, short, reddish setae. Vertex narrow, 1/5 as wide face at antennae. Occiput yellowish, yellowish setose with reddish-yellow postocular bristles and yellow setae. Thorax: Dull brownish-yellow or dull reddish, brownish tomentose, with thin yellow bristles dorsally and yellow setae laterally. Scutum reddish, 3 black stripes dorsally with reddish-brown tomentum, median stripe extends anteriorly to pronotum; tomentum yellow anteriorly, laterally, and between longitudinal black stripes, dull reddish in some light; vestiture reddish, setae most abundant anteriorly, row of thin dorsocentral bristles extend anteriorly to level of postpronotum, lateral bristles longer and thicker. Scutellum and mediotergite reddish, anatergite and katatergite yellow, yellow tomentose; 2 marginal

REVIEW OF ASILIDS © 2006 Magnolia Press 81 ZOOTAXA scutellar bristles present. Pleura mostly dull brownish-yellow, anepisternum, 1381 katepisternum, epimeron, and katepimeron partly blackish, metapleuron yellowish; tomentum mostly bright yellow and yellowish-grayish, anepisternum anteriorly reddish- brown or brown; katatergal bristles thin, reddish. Halter yellow.

FIGURES 105–109. Plesiomma simile, sp. nov,, terminalia. Male, 105. dorsal view; 106. gonocoxa, 107. hypandrium; 108 gonostylus; 109. aedeagus. Abbreviations: Ce=cercus, Ep=epandrium, Ea=ejaculatory apodeme, Gc=gonocoxa, Gs=Gonostylus.

82 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT Wing: Venations reddish-brown or reddish-yellow, microtrichia reddish-yellow. ZOOTAXA Legs: Coxae yellowish-gray or grayish tomentose, vestiture yellowish or reddish- 1381 yellow. Fore coxa yellow; middle coxa entirely and hind coxa brownish or reddish-yellow. Trochanters yellowish, apical margins partly black. Fore and middle legs yellow, fore and mid femora anteriorly except apices and apical tarsomere dark red; hind leg dark red entirely; bristles and setae yellow or red. Tarsal claws black, reddish-yellow basally; pulvilli yellow to reddish-yellow. Abdomen: Slightly spatulate, mostly brownish-red or reddish, base yellowish, apical tergites slightly sooty. Vestiture reddish or yellow; tergite 1 laterally with sparse stout bristles. Terminalia (Figs. 105–109): Red with reddish vestiture. Female. Measurements, body 16.3–28.1 mm; wing 10.9–18.0 mm. Frons yellow or brownish-yellow tomentose. Apical flagellomeres red or black. Vertex 1/6–1/4 as wide as face at antenna. Hind legs pale or dark red. Abdomen slightly spatulate; mostly dark brownish-red. Terminalia (Figs. 110–111).

FIGURES 110–111. Plesiomma simile, sp. nov,, terminalia. Female, 110–111. dorsal and ventral views. Abbreviations: Ac Sp= acanthophorite, Ce=cercus, Gf=genital fork, S8=sternite 8, S10=sternite 10, me=membranous, thinly sclerotized, T8=tergite 8, T9=tergite 9, T10=tergite10.

Types. DOMINICAN REPUBLIC: prov./Barahona, nr. Filipinas / Larimar Mine: 20–26.vi.1992: R.E. Woodruff / P.E. Skelley, at light (holotype %, FSCA). Paratypes, prov. / Barahona, nr. Filipinas / Larimar Mine: 26.vi–7.vii.1992: R.E. Woodruff / P.E. Skelley (1 &, FSCA); DOMINICAN REP.: prov. / Barahona, nr. Filipinas / Larimar Mine: 6–11.vii.1993: R.E. Woodruff / mercury vap. light (1 &, FSCA); La Toma, N. of /

REVIEW OF ASILIDS © 2006 Magnolia Press 83 ZOOTAXA San Cristóbal [province] / 9–10 June 1969 / Flint & Gomez (1 &, USNM). 1381 Distribution. Captured at low elevations during June and July in Barahona and San Cristóbal provinces. Etymology. Latin simile for 'likeness or resemblance' referring to the appearance of the species in this genus. Remarks. Plesiomma simile is distinguished from congeners by the characters in the key, the reddish ground color and yellowish tomentum of the head, reddish wings, and terminalia (Figs. 105–111). It is further recognized by the absence of black vestiture.

Subfamily Trigonomiminae

Tribe Damalini

Holcocephala Jaennicke

Discocephala Macquart, 1838: 50 (1839: 166) (preocc. Laporte, 1832). Type species: rufiventris Macquart, (Coquillett 1910: 534) = abdominalis (Say). Holcocephala Jaennicke, 1867: 359 (nom, nov. for Discocephala Macquart). Type species: Disco- cephala rufiventris Macquart (aut.) = abdominalis (Say).

Remarks. This largely Neotropical genus is composed of small [4–8 mm], broad-winged, weak flying species. Species are distinguished by the wide, flattened head, reduced face and frons, prominent goggle-like eyes and the thorax is strongly arched (Hull, 1962). They usually inhabit moist habitats, usually in diffuse light or shades of nearby forest margins, perching on the tips of vegetation. Here we report the first species in the West Indies.

Holcocephala indigena sp. nov. (Table 1, Figs. 112–117, 123)

Male. Black body, wings, and legs; dark brown tomentum dorsally. Measurements, body 5.0–5.3 mm, wing 5.0–6.3 mm. Head: Face with dorsal 1/3 grayish or sooty black tomentose, sometimes pale with only narrow margin along eye gray; blackish ventrally to or just beyond transverse facial groove, and narrowly along margin of eye and gena; ventral 1/3 yellow or yellowish-gray medially below facial groove and along oral margin; setae yellow, sparse above facial groove, more abundant below; facial width medially slightly greater than diameter of eye. Palpus long, slender, black with yellow setae apically, sometimes mixed with black. Proboscis black, white setose ventrally. Antenna black, black setose; flagellum with short, conical microsegment and seta-like apical style, style subequal or slightly longer than microsegment. Frons, ocellar tubercle, and vertex dark brown or blackish tomentose, tomentum extending narrowly along eye margin

84 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT dorsally; setae sparse, usually blackish, sometimes yellow. Ocellar tubercle prominent, ZOOTAXA unusually wide, with large ocelli. Occiput mostly gray tomentose, setae whitish, sparse, 1381 most abundant ventrally. Thorax: Pronotum blackish or dark brown tomentose, gray with sparse white setae laterally. Scutum dark brown or blackish tomentose, narrow anterolateral and lateral margins posteriorly to transverse suture brownish-yellow tomentose; prescutellum, postalar callus, and dorsal spot above callus brownish-yellow tomentose, often prescutellum and dorsal spot connected by narrow brownish-yellow tomentose stripe; setae sparse, acrostichial setae short, thin, brown anteriorly, sometimes yellow posteriorly; bristles absent. Scutellum with large brown spot dorsally, narrow apical margin, often basal margin, brownish-yellow tomentose; setae sparse, usually yellowish; marginal bristles absent. Mediotergite dark brown tomentose, anatergite usually pale brownish- yellow or grayish. Pleura gray tomentose, anepisternum sometimes yellowish-gray, setae sparse, whitish or yellow. Halter yellow, base and knob usually partially or entirely sooty. Wing: Blackish microtrichose. Legs: Black, usually yellow setose, sometimes partly white; stout bristles sparse, black; tibiae narrowly brownish or yellowish basally. Hind tibiae clubbed, gradually enlarged apically, apex about twice as wide as base. Tarsi black; middle and hind tarsi ventrally dense setose, color usually brassy. Basal tarsomere of hind tarsus slightly shorter than tarsomeres 2–4 combined, about as wide as hind tibia apically. Abdomen: Tergites dark brown or blackish tomentose dorsally with sparse brown setae, narrow sides and sternites entirely gray with white setae. Terminalia (Figs. 112–117): Black, white setose, membranous basally. Epandrium narrow apically, curved dorsally, base membranous dorsally. Gonostylus wider basally than apically, apex horizontal, pointed anteriorly. Aedeagus sigmoid in lateral view, long, and slender, apex trifid with short apical processes; horizontal and ventral apodemes present. Hypandrium swollen, triangular in ventral view, base notched medially. Female. Measurements, body 8.3–8.4 mm, wing 6.3–7.3 mm. Terminalia not dissected. Types. DOMINICAN REPUBLIC: RD-256 La Compartición - / Los Tablones, PN Armando / Bermúdez, La Vega prov. / 2,450–1,110 m, 3.vii.2004, / D.Perez (d) (holotype %, USNM). Paratypes, Bara- / hona [province] 6 km NW Paraiso, Rio / Nizao. 18-02N, 71-12W [18o02'N, 71o12'W], 170 m / 25–26 July 1990, C.Young, / J.E.Rawlins, S.A.Thompson (2 %, CMNH); RD-163 Rio Limpio, Elías / Piña prov., by river, 19o14. / 221'N, 71o31.643'W, 770 m / 24.vii.2003, D.Perez, R.Bas- / tardo, B.Hierro (day/night) (1 &, MHND); Independencia prov. Road / 47, between Los Pinos and / Angel Félix, 760 meters / 18o36.966'N, 71o46.556'W / 20 June 2005 N.E.Woodley (1 &, USNM); Independencia prov. Road / 47, between Los Pinos and / Angel Félix, 847 meters / 18o36.966'N, 71o46.556'W / 16 July 2006 N.E.Woodley (2 %, 2 &, USNM); same data as holotype (2 %, 2 &, USNM); RD-251 Los Tablones – La / Laguna, P N Armando Ber- /

FIGURES 112–117. Holcocephala indigena sp. nov., male terminalia. 112–114. Lateral, dorsal, ventral views; 115. gonostylus; 116–117. aedeagus, dorsal and lateral views. Abbreviations: Ae=aedeagus, Ce=cercus, Ep=epandrium, Ea=ejaculatory apodeme, Gc=gonocoxa, Gs=Gonostylus, H=horizontal apodeme, Hp=hypandrium, V=ventral apodeme, me=membranous, thinly sclerotized.

múdez, La Vega prov. 1270 / -1,980 m, 30.vi.2004, D. / Perez (d) (4 %, 4 &, USNM); Peder- / nales prov. 'Las Abejas' / 7.3 km NNE Las Mercedes / 18o19'N, 71o38'W 1100- / 1150 m 20 August 1983 ~ F.M.Harrington / J. D. Weintraub / collectors (1 %, 2 &, FSCA);

86 © 2006 Magnolia Press SCARBROUGH & PEREZ-GELABERT prov. Pedernales / ca. 35 km N Cabo Rojo / El Aceitillar – Las Abejas / 1250–1430 m, 23 ZOOTAXA Aug 1988 / M.Ivie, T.K.Phillips / & K.A.Johnson (1 &, MAIC); prov. Pedernales / ca. 35 1381 km N Cabo Rojo / Las Abejas, 1250 m / 26 AUG – 09 SEP 1988 ~ M.A.Ivie, T.K.Phillips / & K.A.Johnson colrs. ~ Malaise Trap (2 &, MAIC); Pedernales [province], 23.5 km / N Cabo Rojo. / 18-06N, 71-38W [18o06'N, 71o38'W]. 540 m ~ 13–19 July 1990 / L.Masner, J.Rawlins / C.Young. Deciduous / forest: intercept trap (1 &, CMNH); RD-072 ca. 5 km N. La Colonia, / San Cristóbal prov., 566 m, / 18o31.167'N, 70o16.740'W, 26.xi.2002, D.Perez, B.Hierro, R.Bastardo (night) (1 &, MHND); RD-242 ca. 2 km N. La Colonia / San Cristóbal prov., 24.iv.2004, / D.Perez, B.Hierro. (d) (1 %, MHND).

FIGURES 118–123. Six asilids from Hispaniola. 118. Proctacanthella taina sp. nov., 119. Atomosia ciguaya sp. nov., 120. Cerotainia sola sp. nov., 121. Rhopalogaster albidus sp. nov., 122. Plesiomma simile sp. nov., 123. Holcocephala indigena sp. nov.

Distribution. Captured between 170 and 2450 m during April, June-September, and November in deciduous forests of Barahona, Independencia, La Vega, Elías Piña, San

REVIEW OF ASILIDS © 2006 Magnolia Press 87 ZOOTAXA Cristóbal, and Pedernales provinces. 1381 Etymology. Latin indigena for native, alluding to its distribution being limited to Hispaniola. Remarks. Holcocephala indigena is distinguished by its male terminalia (Figs. 112–117) and is the first species of the genus reported in the Greater Antilles.

Acknowledgments

We thank our colleagues who collected numerous asilids and made them available to us during this study; Chen Young, J. E. Rawlins, and associates (CMNH), Norman E. Woodley (Systematic Entomology Laboratory, PSI, ARS, USDA, National Museum of Natural History, Smithsonian Institution — USNM), Mike Ivie, Montana State University (MAIC), Robert Woodruff (FSCA), Ruth Bastardo (Universidad Autónoma de Santo Domingo and Fundación Moscoso Puello, Inc.) and Brígido Hierro (Departmento de Vida Silvestre, Secretaría de Medio Ambiente, Santo Domingo). Brian D. Farrell and Stefan Cover (MCZ) are also thanked for the loan of type specimens. F. Christian Thompson (Systematic Entomology Laboratory, PSI, ARS, USDA, at the National Museum of Natural History, Smithsonian Institution (USNM), Wayne Mathis and Holly Williams (Department of Entomology, National Museum of Natural History, Smithsonian Institution, USNM), and David Grimaldi (AMNH) are thanked for permitting access to Bromley's and Curran's asilid types, respectively. Special thanks are extended to Chen Young, Gary Steck, (FSCA), F. Christian Thompson, and David Grimaldi for their help during AGS's visits to the museums. Eric Fisher, Plant Pest Diagnostics California Department of Food & Agriculture, Sacramento, California, is thanked for calling our attention to the status of Andrenosoma, allowing us to review a manuscript of the Central American Asilidae, and numerous helpful suggestions during the course of this study. We thank Francisco Núñez and Andrés Ferrer (Fundación Moscoso Puello, Inc.) for logistical support with the field vehicle and Ramón Elías Castillo (Fundación Progressio, Inc.) for access to the Reserva Científica Ebano Verde. Matilde Mota, Ramón Ovidio Sánchez and José Ramón Albaine (Dirección Nacional de Parques and Departamento de Vida Silvestre, Santo Domingo) assisted with collecting and export permits. Field work by DEPG in the Dominican Republic was supported by National Science Foundation grant DEB-0103042 and a grant from the S. W. Williston Diptera Research Fund, Smithsonian Institution. We also thank Drs. Eric Fisher and Neil Evenhuis, Department of Natural Sciences, Bishop Museum for their critical reviews of the manuscript.

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