Introducción

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Introducción UNIVERSIDAD DE GUADALAJARA CENTRO UNIVERSITARIO DE CIENCIAS BIOLÓGICAS Y AGROPECUARIAS POSGRADO EN CIENCIAS BIOLÓGICAS PATRONES DE VARIACIÓN QUÍMICA DE POBLACIONES DEL COMPLEJO PINUS AYACAHUITE-PINUS STROBIFORMIS Por LETICIA MAYA ZEPEDA Tesis presentada como requisito parcial para obtener el grado de MAESTRO EN CIENCIAS BIOLÓGICAS (ÁREA DE ECOLOGÍA) LAS AGUJAS, ZAPOPAN, JALISCO MAYO DEL 2006 PATRONES DE VARIACIÓN QUÍMICA DE POBLACIONES DEL COMPLEJO PINUS AYACAHUITE-PINUS STROBIFORMIS Por LETICIA MAYA ZEPEDA Tesis presentada como requisito parcial para obtener el grado de MAESTRO EN CIENCIAS BIOLÓGICAS (ÁREA DE ECOLOGÍA) UNIVERSIDAD DE GUADALAJARA CENTRO UNIVERSITARIO DE CIENCIAS BIOLÓGICAS Y AGROPECUARIAS MAYO DE 2006 Aprobada por: ________________________________________________ _________________ Dra. Alma Rosa Villalobos Arámbula Fecha Asesor del Comité Particular __________________________________________________ _________________ M. en C. Antonio Rodríguez Rivas Fecha Asesor del Comité Particular __________________________________________________ _________________ Dr.a Anne Santerre-Lucas Fecha Sinodal del Comité Particular __________________________________________________ ________________ Dr. Jorge A. Pérez de la Rosa Fecha Sinodal del Comité Particular __________________________________________________ ________________ Dr. Fernando A. López Dellamary Fecha Presidente del Comité Particular Este trabajo se llevo a cabo en el Laboratorio de Cromatografía del Departamento de Madera Celulosa y Papel y en el Laboratorio de Bioprocesos del Departamento de Ingeniería de la División de Ingenierías del Centro Universitario de Ciencias Exactas e Ingenierías; Universidad de Guadalajara y forma parte del proyecto, “PINOS BLANCOS MEXICANOS: DIVERSIDAD GENÉTICA, QUÍMICA Y UBICACIÓN TAXONÓMICA”. financiado SEP-CONACYT (33129-B) (Responsable Dra. Alma Rosa Villalobos Arámbula), con la Beca de Proyectos de Investigación Número 3204 para Leticia Maya Zepeda. v CONTENIDO ÍNDICE DE FIGURAS..................................................................................................vi ÍNDICE DE TABLAS....................................................................................................ix RESUMEN.....................................................................................................................x SUMMARY...................................................................................................................xii INTRODUCCIÓN..........................................................................................................1 ANTECEDENTES..........................................................................................................3 1. Descripción del Complejo Pinus ayacahuite-P. strobiformis. 2. Variación de la Composición Química de los Pinos. 3. Quimiotaxonomía de Pinos. 4. Monoterpenos reportados en Pinos de México y Mesoamérica. 5. Biogeografía 6. Metabolómica OBJETIVO GENERAL.................................................................................................11 OBJETIVOS PARTICULARES....................................................................................11 HIPÓTESIS....................................................................................................................11 MATERIALES Y MÉTODOS......................................................................................12 RESULTADOS Y DISCUSIÓN...................................................................................15 CONCLUSIONES.........................................................................................................40 BIBLIOGRAFÍA...........................................................................................................42 ANEXOS ….………………………………………………………………………….45 vi ÍNDICE DE FIGURAS Figura 1. Características morfológicas de acículas, conos y semillas del complejo P. ayacahuite-P. strobiformis. ............................................................................17 Figura 2. Mapa de localización de los sitios de colecta ...............................................12 Figura 3. Perfiles cromatográficos en diferentes extractos de acículas de P. ayacahuite Ehrenberg ex Schlechtendal, localizado en el Departamento de Madera Celulosa y Papel ……….................................................................................15 Figura 4. Perfiles cromatográficos de ramas y acículas de P. ayacahuite y P. strobiformis ....................................................................................................16 Figura 5 . Perfiles cromatográficos de muestras de acículas extraídas sonicación ..........................................................................................................................17 Figura 6. Compuesto sugerido conforme a su índice de similitud ...................................................................................................................... 18 Figura 7. Perfil cromatográfico de P. strobiformis 138, localidad de Creel en Chihuahua .......................................................................................................24 Figura 8. Perfil cromatográfico de P. ayacahuite, muestra 45b, comparado con el perfil cromatográfico de P. oocarpa, muestra 151 ..................................................25 Figura 9. Perfil cromatográfico de P. ayacahuite, muestra 45b (arriba), comparado con el perfil cromatográfico del P. ayacahuite localizado en el DMCyP (abajo) ..........................................................................................................................26 vii Figura 10. Perfil cromatográfico de acículas de P. ayacahuite, localidad: Las Palmas, municipio de Los Reyes, Michoacán ..........................................................27 Figura 11. Perfil cromatográfico de acículas de P. ayacahuite, localidad: Comunidad de Palo Hueco, municipio de Real del Monte, en el Edo. de Hidalgo ..........................................................................................................................28 Figura 12. Perfil cromatográfico de acículas de de P. ayacahuite, de la localidad Km. 228 carretera 186 Palenque-San Cristóbal, municipio de San Cristóbal Chiapas .........................................................................................................................28 Figura 13. Perfil cromatográfico de acículas de P. ayacahuite, de la localidad: Km. 4.5, 6, 9, 31 y 32 del camino Talpa-Cuale, municipio de Talpa de Allende, en el Edo. de Jalisco ................................................................................................29 Figura 14. Comparación de los perfiles cromatográficos de acículas de P. ayacahuite muestreado en diferentes localidades ............................................................29 Figura 15. Perfil cromatográfico de acículas de P. ayacahuite var. Veitchii ...............30 Figura 16. Perfil cromatográfico de acículas de P. ayacahuite var. veitchii comparado con el perfil cromatográfico de acículas P. ayacahuite ..........................................................................................................................31 Figura 17. Perfil cromatográfico de acículas de P. strobiformis, de la localidad: km 130 de la carretera 40 Durango-Mazatlán, 3 km al S por el camino a Chavaría, municipio de Pueblo Nuevo, en el Edo. de Durango ……………………………………………………...........................................32 Figura 18. Perfil cromatográfico de acículas de P. strobiformis, de la localidad: Observatorio INAOE, en el municipio de Cananea, en el Edo. de Sonora ..........................................................................................................................33 viii Figura 19. Perfil cromatográfico de acículas de P. strobiformis, de la localidad: Carretera Creel-Divisadero, en el municipio de Boycona, Chihuahua ...........33 Figura 20. Comparación de los perfiles cromatográficos de acículas de P. strobiformis muestreado en diferentes localidades ..............................................................34 Figura 21. Perfil cromatográfico de acículas de P. strobus var. chiapensis Martínez, de la localidad: Parada La Reforma, 8 km al oeste de Tlapacoyan, municipio de Tlapacoyan , Edo. de Veracruz .......................................................................35 Figura 22. Perfil cromatográfico de P. strobus var. chiapensis Martínez, de la localidad: Km. 200 carretera 195 en el municipio de Bochil, Estado de Chiapas ..........................................................................................................................35 Figura 23. Identificación de los compuestos elegidos ..........................................................................................................................36 Figura 24. Correlación de los pinos blandos analizados en el estudio ………………………......................................................................................37 ix ÍNDICE DE TABLAS Tabla 1. Compuestos sugeridos en ramas del complejo P. ayacahuite-P. strobiformis.......................................................................................................19 Tabla 2. Compuestos sugeridos para acículas del complejo P. ayacahuite-P. strobiformis ......................................................................................................21 Tabla 3. Resumen de las funciones canónicas discriminante ....................................38 Tabla 4. Coeficientes estandarizados de las funciones discriminantes canónicas ……………………………………………………………………………......38
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