© Entomologica Fennica. 26 October 1998

Boreal 1. {: ): New Synonymies and Holarctic Records

Jukka Jalava & William E. Miller

Jalava, J. & Miller, W. E. 1998: Boreal Olethreutini 1. (Lepidoptera, Tortrici­ dae): New Synonymies and Holarctic Records.- Entomol. Fennica 9: 137-142. We examined specimens of selected boreal palaearctic and nearctic taxa of Tortricidae to ascertain whether they might be holarctic conspecifics. Our findings confirm one putative synonymy: Tia enervana (Erschoff, 1877) = vulgana McDunnough, 1922, support four new synonymies: aquilonanus (Karvonen, 1932) = 0. kononenkoi Kuznetsov, 1991, 0. heinrichanus (McDunnough, 1927) = 0. hyperboreanus (Karvonen, 1932), 0 . turfosanus (Herrich-Schaffer, 1851) = 0 . intermistanus (Clemens, 1865), 0. septentrionanus (Curtis, 1835) = 0. schaefferanus (Herrich-Schiiffer, 1851) and resurrect one name from synonymy: the nearctic 0. kennethanus McDun­ nough, 1941 is not conspecific with the palaearctic 0. obsoletanus (Zetterstedt, 1840). Through synonymies and records, seven are new for the Nearctic, and ten are considered holarctic. These results increase the percentage of holarctic Beringian tortricid species from 48 to 61. Jukka Jalava, Finnish Museum ofNatural History, Zoological Museum, Division of Entomology, P. 0. Box 17 (P. Rautatiekatu 13) FIN-00014 University of Helsinki, Finland William E. Miller, Department ofEntomology, University ofMinnesota , 1980 Folwell Avenue, St. Paul, Minnesota 55108-6125, USA Received 23 May 1997, accepted 20 Apri/1998

1. Introduction 2. Materials and methods

Our collaboration started after Miller ( 1995) syn­ Much of our material was collected by various onymized the Nearctic E. arctica Miller of Alaska Finnish-Russian expeditions to Siberia starting in and Yukon with the Transpalaearctic Epiblema 1982 (Kuznetsov & Jalava 1988, Kuznetsov & simplonianum (Duponchel), thus making the lat­ Mikkola 1991). Material was also collected dur­ ter holarctic. It seemed possible that other boreal, ing a Finnish expedition to Yukon in 1985, as well boreo-montane, and arcto-alpine tortricid taxa of as during other Finnish collecting visits to Canada. Siberian Beringia and its Nearctic counterpart Material in several North American museums was across the Straits might prove to be synonymous, examined also. Some taxa discussed here are rare holarctic, or both. We focused the investigation on in museums, and the sexes of some have not yet species of Olethreutes and related genera of Olethreu­ been associated, hence the small numbers of speci­ tini having northeastern Siberian distributions. mens on which some conclusions are based. 138 Jalava & Miller: Boreal Olethreutini 1. • ENTOMOL. FENNICA Vol. 9

Nomenclature is reviewed only to the extent Olethreutes aquilonanus (Karvonen) necessary for our purposes. For the most part, mac­ ulation among the species treated here is so simi­ Argyroploce aquilonana Karvonen (1932:77) lar as to be oflittle value in diagnosis. Unless oth­ Olethreutes kononenkoi Kuznetsov, in Kuznetsov and Mikkola (1991:204). New Synonymy. erwise noted, all specimens listed by sex (d',9) in Material Examined sections were dissected for This species was long known only from north­ genitalia study. em Fennoscandia. In the Polar Ural of Russia, the Depository abbreviations are as follows: were recently collected commonly on ANSP, Academy of Natural Sciences, Philadel­ mountain tundra, which led to the recognition of the above synonymy. This synonymy makes the phia, Pennsylvania; CNC, Canadian National Col­ species transpalaearctic; we report it here as new lection, Ottawa; UCB, Essig Museum of Entomol­ for the nearctic, which makes it holarctic also. ogy, University of California, Berkeley; INHS, Illinois Natural History Survey, Champaign; Material examined. FINLAND: Malla,ld', 10.-18.VII.29 (aquilonanus holotype); Kilpisjarvi, 1d', 9.VII.76; 19, UMEM, University of Minnesota Entomology 8.VII.43. RUSSIA: Kola Pns., 1d', 18.VII.93; Polar Ural, Museum, St. Paul; USNM, U. S. National Mu­ 5d', 19, l.-15.VII.94; SW Altai, 19 exx., 20.-2l.VII.83 seum ofNatural History, Washington, D. C.; ZIN, (reported by Kuznetsov & Jalava 1988 as 0. aquilonanus Zoological Institute, St. Petersburg; MZH, Finn­ helvomaculanus Kostjuk, a larger and brighter colored sub­ species) (ld' dissected in this study); Chukotka, Chukchi ish Museum of Natural History, Helsinki. Pns., 1d', 29.Vl1.82 (kononenkoi paratype). CANADA: Ogilvie Mts., Yukon, 2d', 5.-6.VII.85 (MZH).

3. Results Olethreutes heinrichanus (McDunnough) The following five species are holarctic by virtue of our confirming or proposing synonymies. Argyroploce heinrichana McDunnough (1927:33) Argyroploce hyperboreana Karvonen (1932:78) Lectotype designated here: Jenisej [Siberia], J. Sahib., coli. Tengstr., Zoo!. Mus. H:fors, spec. type No. 7173 Tia enervana (Erschoff) [male]; in MZH. New Synonymy. As 0. hyperboreanus, this species was re­ Penthina enervana (Erschoff 1877:341) ported to occur widely in the Palaearctic, includ­ Argyroploce vulgana McDunnough (1922:46) ing northeastern Siberia (Kuznetsov & Mikkola Kuznetsov (1987) reported Tia enervana to 1991). As 0. heinrichanus, it was previously known occur in northeastern Siberia and North America, in the Nearctic only from Labrador (McDunnough and McDunnough (1922) described Argyroploce 1927). The above synonymy makes the species vulgana from Nordegg, Alberta, Canada. Our find­ holarctic, and the Alaska and Yukon records be­ ings justify previous usage of the palaearctic senior low suggest it being transnearctic as well. synonym for Nearctic material (Kuznetsov 1987, Material examined. FINLAND: Lapland, several exx .; Dang 1990), thereby confirming that the species Li. Inari, 2d', 6.Vll.79. RUSSIA: Jenisej, 1d' (hyperborea­ is holarctic. nus lectotype); Polar Ural, 1d', 4.VII.94; Magadan, ld', 14.VII.87 (MZH). CANADA: "Ungava Bay, 76", ld' Material examined. RUSSIA: Kiiltuk, 19, 1l.VI.[18]69 (USNM); Yukon, 3d', 4.VIl.94 (MZH). USA: nr. Houston, (enervana holotype) (ZIN); Polar Ural, 45 exx. (1d' dis­ Alaska, ld', 26.VI.79 (UCB). sected), 6.-16.VII.94; Chukotka, ld', 3.VII.9l (MZH). CAN­ ADA: Nordegg, Alberta, 1d', 11.VII.21 ( vulgana holotype; previously published capture date corrected here), 1d', Olethreutes turfosanus (Herrich-Schaffer) 27.VI-14.VII.21 (vulgana paratype) (CNC); Aklavik, N.W.T., ld', 20.VII.31. USA: Peters Lk., Alaska, ld', Sericoris turfosana Herrich-Schaffer (1851 :217) 14.VII.73; SchraderLk., Alaska, 29, 4.- 27.VII.73 (USNM); Mixodia intermistana Clemens (1865:140). New Syn­ nr. Cantwell, Alaska, 2d', 27.VI.-2.VII.79 (UCB). onymy. ENTOMOL. FENNICA Vol. 9 • Jalava & Miller: Boreal Olethreutini 1. 139

Heinrich (1926) considered the above two taxa The following five species are holarctic by vir­ to be distinct, citing different lengths of spined tue of our confirming or newly discovering their oc­ lower margins of the valvae. In our material, this currence in the nearctic. difference is individual rather than taxonomic. As 0. tuifosanus, whose larvae probably live on Eri­ caceae and Vacciniaceae (Kuznetsov 1987), the atropunctana (Zetterstedt) species was reported to be transpalaearctic (Kuz­ netsov & Mikkola 1991); the above synonymy Penthina atro-punctana Zetterstedt (1840:977) makes it holarctic. Razowski (1983) photo-illus­ Kuznetsov and Jalava (1988) reported this spe­ trated the wings and line-illustrated male and fe­ cies to be transpalaearctic and possibly holarctic. male genitalia (as tuifosana), and Heinrich Our findings confirm that it is indeed western ne­ (1926) photo-illustrated male and line-illustrated arctic and thus holarctic also, vicariated by M. se­ female genitalia of 0. intermistanus. paratana (Kearfott, 1907) in the eastern N earctic. Material examined. FINLAND: > 100 exx. from The maculation of these two species is similar, throughout the country, the species being very common on but the male genitalia, line-illustrated by Miller peat bogs; Pellinge, ld', 19.VI.38. RUSSIA: Polar Ural, 25 1987 (as separatana), are different. The exx., VII.94; SW Altai, 10 exx., l0.-14.VII.83 (3d' dissected larvae of M. atropunctana feed in webbed leaves in tills study); Irkutskaya obl., 16 exx., VII.84 (1d' dissected of species of and other in this study); Provideniya, Chukchi Pns., ld', 7.VII.91 Betula, Salix, Tilia, Fagus, (MZH). CANADA: Bilby, Alberta, 1d', 23.VI.24; Church­ genera (Bentinck & Diakonoff 1968, Razowski ill, Manitoba, 1d', 5.VII.83; Hopedale, Labrador, 1<:;?, 1983, Kuznetsov 1987). Bentinck and Diakonoff 24.VIII.3l. USA: Mt. Katahdin, Maine, 1d', 8.VII.39; Mt. (1968) photo-illustrated the wings and genitalia Washington, N. H., 1d', (no date) (USNM); nr. Anderson, of both sexes (as Hedya atropunctana), and Ra­ Alaska, 1d', 28.VI.79; nr. Cantwell, Alaska, 1d', 1<:;?, zowski (1983) photo-illustrated the wings and 27.VI.79 (UCB); Mt. Baker Natl. For., Wyoming, 2<:;? , I.VII1.36 (ANSP); Bangor, Maine, 1<:;?, 15.VI.38 (INHS). line-illustrated the genitalia of both sexes. Material examined. FINLAND: Several exx. from throughout the country. RUSSIA: Kola Pns., 2d', 1<:;?, Olethreutes septentrionanus (Curtis) 22.VI.93; Prov. Petropolis, 1d' (no date), Coil. Duske; CAN­ ADA: Ogilvie Mts., Yukon, ld', 12.VII.85 (MZH). USA: Dietrich River, Alaska, 1d', 19.VII.1971 (UMEM). septentrionana Curtis (1835:74) Sericoris schaefferana Herrich-Schaffer (1851 :211), New Synonymy. Olethreutes dalecarlianus (Guenee) Kuznetsov and Mikkola (1991) reported the palaearctic taxon from several areas including dalecarliana (Guenee 1845: 160) northeastern Siberia. The nearctic taxon was pre­ viously known only from Labrador (Heinrich 1926). Kuznetsov (1987) reported this species to be The above synonymy makes the species holarctic. transpalaearctic; we report it here as new for Ne­ Razowski (1983) photo-illustrated the wings and arctic, which thus makes it holarctic also. The lar­ vae feed in tied leaves of species of Pyrola (Ben­ line-illustrated the male and female genitalia (as tinck & Diakonoff 1968, Razowski 1983, Kuz­ Phiaris schaefferana), and Heinrich (1926) photo­ netsov 1987). Bentinck and Diakonoff (1968) pho­ illustrated the male genitalia of 0. septentrionanus. to-illustrated the wings and genitalia of both sexes Material examined. FINLAND: Lapland, where the (as Selenodes dalecarliana ), and Razowski ( 1983) species is locally common, many exx.; Malia, ld', l.VII.36. photo-illustrated the wings and line-illustrated the RUSSIA: KolaPns., many exx., 16.VII.91 (1d'dissected in genitalia of both sexes (as A rgyroploce dalecarlia­ this study); Magadan obl., 8d', 3.VII.90 (ld' dissected in na). The maculation is very similar to that of the this study) (MZH). CANADA: Labrador, 1<:;?, "609"; Bra­ dare Bay, Quebec, ld', 14.VII.29 (USNM). USA: nr. Cant­ nearctic 0. costimaculanus, which is discussed well, Alaska, ld', !<:;?, 27.VI.79; nr. Anderson, Alaska, ld', in another context further on, but these two taxa 18 .VI.79 (UCB). are distinct structurally. 140 Jalava & Miller: Boreal Olethreutini 1. • ENTOMOL. FENNICA Vol. 9

Material examined. FINLAND: Flaka, 4 exx., reared and Diakonoff (1968) photo-illustrated the wings [19]48; Perna, 19, 23.VI.42; Sortavala, 19, 29.VI.20; Ka­ and genitalia of both sexes (as Argyroploce pa­ rislojo, 19, "897"; Tervola, ld', 10.VII.71 ; Suojarvi, ld', lustrana ), and Razowski (1983) photo-illustrated 1939. RUSSIA: Magadan obi., ld', 15.-18.VII.87; Prov. the wings and line-illustrated the genitalia of both Petropolis, (no date), ld', Coli. Duske. CANADA: Ogilvie Mts., Yukon, ld', 4.VII.85 (MZH). Wellington, British Co­ sexes (as Phiaris palustrana). lumbia, "vi" (misidentified in 1925 as 0 . costimaculanus) Material examined. FINLAND: Ca. 100 exx. from (USNM). throughout the country (MZH); Nurmijarvi, 1d', 7. VIII.71; Pellinge, ld', 20.VII.36; N. Esbo, ld', 2l.VIII.65; Helsinki, 19, 10.VII.2l (UMEM). RUSSIA: SW Altai, ld', 9-14.VII.83 Olethreutes exaridanus Kuznetsov (MZH). USA: Galbraith Lk., Alaska, ld', 9.VII.79 (UCB); Schrader Lk., Alaska, 5d', 19, 3.-26.VII.73 (USNM). Olethreutes exaridanus Kuznetsov, in Kuznetsov and Based on our studies, the species discussed Mikkola (1991:202) below, although Beringian, do not appear thus far This species was previously known only from to be holarctic. the Chukchi Peninsula type locality (Kuznetsov & Mikkola 1991). We report it here as new for the nearctic, which thus makes it holarctic also. Olethreutes kennethanus McDunnough, New Material examined. RUSSIA: nr. Provideniya, Chukchi Status. Pns., 2d', 9.-14.VII.91 ; Anadyr River, Chukchi Pns., 59 (para types, not dissected), 1d', 2l.VII.89. CANADA: Ogil­ 0/ethreutes kennethana McDunnough ( 1941 :99) vie Mts., Yukon, ld', 7.VII.85 (MZH). Olethreutes obsoletana (not Zetterstedt 1840:980); Miller (1985:414) We resurrect the name 0. kennethanus because Olethreutes concretanus (Wocke) the taxon differs from 0. obsoletanus in number, size, and arrangement of comuti. In 0. (or Phiaris) Penthina concretana Wocke (1862:57) obsoletanus, there are three groups of comuti: one This boreal species, described from N-Nor­ group of one to three long ones at the tip of the way, whose larvae are said to feed on Ericaceae aedeagus, and two separate vesical plates each and Vacciniaceae, was previously reported to be with six or more short ones (Razowski 1983, Kuz­ transpalaearctic (Kuznetsov 1987, Kuznetsov & netsov 1987). In 0 . kennethanus, there are up to Mikkola 1991). We report it here as new for the five separate vesical com uti, all long, and no plates nearctic, which thus makes it hoi arctic also. Kuz­ of short cornuti. Also the sacculus differs: in 0. ken­ netsov (1987) line-illustrated the male genitalia. nethanus it is evenly curved and with rows of numerous spines; in 0. obsoletanus it is more an­ Material examined. FINLAND: Middle parts of the gulate and contains only a group of spines (Kuz­ country north to Lapland, 27 exx .. RUSSIA: Kola Pns., 13 exx.; Polar Ural, 29, l.-6.Vll.94; Upper Kolyma River, Ma­ netsov 1987). Staining of earlier genital prepara­ gadanobl., ld', 19, 8.-18.VII.87; ChukchiPns., 1'¥, 7.Vll.91 tions obscured the plates of short cornuti in 0 . ob­ (MZH). USA: Eagle Summit, Alaska, ld', 2.VII.79; nr. Fair­ soletanus and thus led to the synonymical error. banks, Alaska, 1"?, l.VII.79 (UCB). Material examined. 0. kennethanus: CANADA: Ed­ monton, Alberta, ld', l.VIII.40 (holotype) (CNC); Church­ ill, Manitoba, ld', 4.VII.37 (USNM); Lk. Abitibi, Ontario, Olethreutes palustranus (Lienig & Zeller) ld', 28 .VJJ .92 (MZH). USA: Bear Creek, Alaska, ld', 28.VI.70; Mt. McKinley, Alaska, ld', 3.VII.38; Peters Lk., Sericoris palustrana Lienig & Zeller (1846:230) Alaska, ld', 19, 14.VII.73 (USNM); nr. Cantwell, Alaska, 2d', 27 .VI.79 (UCB). 0 . obsoletanus: FINLAND: Lapland, This species was previously reported to be where the species is one of the most abundant lepidopterans, transpalaearctic (Kuznetsov and Mikkola 1991). many exx. (8d' dissected for this study); Kilpisjarvi, ld', We report it here as new for the nearctic, which ll.VII.34. RUSSIA: Polar Ural, 36 exx., 7 .-13.VII.94 (2d' thus makes it holarctic also. The larvae feed in­ dissected in this study); SW Altai, 7d', 22.- 27 .VI.83 (ld' side tubes constructed of moss stems (Bentinck dissected in this study) ; Irkutskaya obi., ld', 16-18.VII.84 & Diakonoff 1968, Kuznetsov 1987). Bentinck (MZH). ENTOMOL. FENNICA Vol. 9 • Jalava & Miller: Boreal Olethreutini 1. 141

Olethreutes mengelanus (Fernald) Olethreutes bowmananus (McDunnough)

Sericoris mengelana Fernald (1894:131) Argyroploce bowman ana McDunnough (1923: 165) This nearctic species was previously known Although the line-illustration of the valva of only from Greenland and Southampton Island, this species (McDunnough 1923) is similar to that N. W. T. (Heinrich 1926, 1935). The new records of 0. exaridanus Kuznetsov (Kuznetsov & Mik­ below from Alaska and Yukon suggest a trans­ kola 1991), specimen comparisons showed that nearctic distribution. the uncus and socii are much smaller in 0 . bowma­ nanus. Material examined. GREENLAND: 1d', 26. Vl.21; 1d', (no date); l<.;l , "29.7" (USNM). CANADA: Ogilvie Mts., Material examined. 0 . bowmananus: CANADA: Nord­ Yukon, ld', 5.-6.VII.85 (MZH). USA: Barter Is., Alaska, egg, Alberta, ld', 7.VIII.23 (paratype); Banff, Alberta, ld', 1d', 25 .VI.71 (UMEM). 4.VII.25; Moraine Lk., Alberta, 1d', 7.VIII.23 (USNM); OgilvieMts., Yukon, 3d', 3.-19.VII.85 (MZH). 0. exarida­ Besides being Beringian and apparently con­ nus: as enumerated in a previous discussion. fined to the Nearctic, the following three species have similar but distinct counterparts on the palae­ arctic side of the Straits. These pairs of species Olethreutes minaki McDunnough are examples of zoogeographic vicariants in the sense of Kuznetsov and Mikkola (1991). Argyroploce minaki McDunnough (1929:270) Although the line-illustration of the valva Olethreutes costimaculanus (Fernald) (McDunnough 1929) is similar to that of 0 . disso­ lutanus (Stange) (Kuznetsov 1987), specimen Penthina costimaculana Fernald (1882:70) comparisons showed that the uncus differs in out­ line and the corn uti are much longer in 0. minaki. Besides superficially resembling the trans­ Olethreutes dissolutanus, whose larvae live be­ pa1aearctic 0. dalecarlianus, as discussed earlier, tween tied moss-stems (Kuznetsov 1987), is dis­ the nearctic 0 . costimaculanus is also similar to tributed widely in the palaearctic, including north­ 0 . ledianus (L.), another transpalaearctic species eastern Siberia (Kuznetsov & Mikkola 1991). (Kuznetsov & Mikkola 1991 ). We confirmed that Material examined. 0 . minaki: CANADA: Minaki, On­ the latter two taxa differ structurally, most nota­ tario, ld', 8.VII.28 (paratype) (USNM). 0 . dissolutanus: bly by 0. costimaculanus male genitalia having FINLAND: Many exx., the species being moderately com­ large, broad and blunt socii (small, narrow and mon in spruce forests. RUSSIA: SW Altai, 9 exx. (ld' dis­ sharp-tipped in 0. ledianus). In female genitalia sected), 23.-25.Vll.83; Upper Kolyma River, Magadan obi., the papillae anales are several times larger than 12 exx. (2d' dissected), 14.-28 .VII.87 (MZH). those of 0. ledianus, as can be seen in published illustrations (Heinrich 1926, Bentinck & Diako­ noff 1968, Razowski 1983). Olethreutes costima­ 4. Discussion culanus was previously known to occur no fur­ The holarctic taxa discussed here are also Berin­ ther west in the nearctic than Alberta (Heinrich gian. The number of Beringian tortricid species 1926), but the Yukon record below suggests it is that are also holarctic previously stood at 48% transnearctic. (Miller 1995). With the net addition or confirma­ Material examined. 0. costimaculanus: USA: Orono, tion here of 10 more holarctic species, the figure Maine, 1<_fl , V1.17.[18]81 (lectotype) (USNM); Bangor, rises to 61% (42/69). We suspect this percentage Maine, 2<_fl , Vl.15.38 (INHS). CANADA: Meach Lk., Que­ will increase even more with further collecting bec, 1d', 25.V.03; Mer Bleue, Ontario, 1<.fl, (no date) and identification of unsorted nearctic and pa­ (USNM); Ogilvie Mts., Yukon, 1d', 4.VII.85; Newfound­ laearctic material from Beringia. land, 1d' 1949 (MZH). 0. ledianus: FINLAND: Many exx. from throughout the country, where the species is common Acknowledgement. We thank D. Azuma, R. L. Brown, on peat bogs. RUSSIA: Magadan obi., 19, 3d', 2.VII.90; P. 1. Clausen, P. T. Dang, R. W. Hodges, V.I. Kuznetsov, Kola Pns. , ld', 3. VII.93, 19, 2.VII.90 (MZH). K. Methven, and J. A. Powell for the loan of specimens in 142 Jalava & Miller: Boreal Olethreutini 1. • ENTOMOL. FENNICA Vol. 9 their care, and R. L. Brown and J. A. Powell for reviewing Ziinsler und Wickler.- Manz, Regensburg, 218 pp. the manuscript. Special thanks to all the participants of the Karvonen, V. J. 1932: Vier neue Kleinschmetterlinge aus expeditions and to the Academy of Finland and Russian Finnland.- Not. Entomol. 12: 77-81. Academy of Sciences for granting and to J. Kullberg and Kuznetsov, V.I. 1987: Tortricidae (tortricid moths). L. Kaila for using their e-mail. -In: Medvedev, G. S. (ed.), Keys to the of the European part of the USSR [translation].-U.S. Dept. Agr. & Nat!. Sci. Found, pp. 279-956. Kuznetsov, V.I. & Jalava, J. 1988: Soviet-Finnish entomo­ References logical expeditions to southern Siberia 1982-1984. 2. Tortricidae (Lepidoptera).- Nota Lepid. 11: 126--138. Bentinck, G. A. & Diakonoff, A. 1968: De nederlandse Blad­ Kuznetsov, V.I. &Mikkola, K. 1991: The leaf-roller fauna rollers (Tortricidae). -Monogr. Nederl. Entomol. Ver. 3, of north-eastern Siberia, USSR, with descriptions of 200 pp. three new species (Lepidoptera, Tortricidae).- Nota Clemens, B. 1865: North American micro-lepidoptera.­ Lepid. 14: 194--219. Proc. Entomol. Soc. Philadelphia 5: 133-147. Lienig, F. & Zeller, P. C. 1846: Lepidopterologische Fauna Curtis, J. 1835: Insects- In: Ross, J. C. (ed.), Appendix to von Lievland und Curland.- Isis 1846: 175-302. the narrative of a second voyage in search of a north­ McDunnough, J. 1922: Undescribed Lepidoptera in theCa­ west passage, and of a residence in the arctic regions nadian National Collection. -Can. En to mol. 54: 34--4 7. during the years 1829, 1830, 1831, 1832, 1833. Lon­ McDunnough, J. 1923: New Canadian Lepidoptera. -Can. don, A. W. Webster, pp.lix-lxxx. [Pagination not con­ Entomol. 55: 163-168. tinuous.] McDunnough, J. 1927: Two new Canadian tortricids (Lepid.). Dang, P. T. 1990: Redefinition of Bactrini Falkovitsh -Can. Entomol. 49: 33-34. and revised status of genera Taniva Heinrich and Hulda McDunnough, J. 1929: Some apparently new microlepi­ Heinrich (Tortricidae: ).-J. Lepid. Soc. doptera.- Can. Entomol. 61:266--271. 44:77-87. McDunnough, J. 1941: A new Albertan olethreutid. -Can. Erschoff, N. 1877: Diagnosen neuer Lepidopteren aus den Entomol. 73: 98-99. verschiedenen Provinzen des Russichen Reiches. - Miller, W. E. 1985. Nearctic Olethreutes: Five new synony­ Horae Soc. Entomol. Ross. 12: 336-348. mies, two revised statuses, and notes (Lepidoptera: Tor­ Fernald, C. H. 1882: Descriptions of new species ofTortri­ tricidae). -Proc. Entomol. Soc. Washington 87:408- cidae.- Trans. Amer. Entomol. Soc. 10: 65-72. 417. Fernald, C. H. 1894: North Greenland microlepidoptera. Miller, W. E. 1987: Guide to the Olethreutine Moths of -Entomol. News 5: 129-132. Midland North America (Tortricidae).-U.S. Depart­ Guenee, M.A. 1845. Essai sur une nouvelle classification ment of Agriculture, Forest Service, Agriculture Hand­ des microlepidopteres et catalogue des especes book 660, 104 pp. europeennes connues jusqu'a ce jour. - Ann. Soc. Entomol. France (ser. 2) 3: 105-192, 297-344. Miller, W. E. 1995: Beringian Tortricidae: A new synonymy Heinrich, C. 1926: Revision of the North American moths in Epiblema and a recount of holarctic species. - J. of the subfamilies Laspeyresiinae and Olethreutinae. Lepid. Soc. 49: 250--252. - U. S. Nat!. Mus. Bull. 132, 207 pp. Razowski, J. 1983: Motyle (Lepidoptera) Polski, 6- Ole­ Heinrich, C. 1935: Heterocera.- In: Sutton, G. M. (ed), threutinae: Olethreutidii.- Monogr. Fauny Polski 13, The exploration of Southampton Island, Hudson Bay. 177 pp. Pt. 2, Zoology. Sec. 5, Insects (Lepidoptera) and inver­ Wocke, M. F. 1862: Reise nach Finnmarken II. Microlepi­ tebrates.- Mem. Carnegie Mus. 12, pp. 27-29. doptera.- Stett. Entomol. Zeit. 23: 30-78, 233-257. Herrich-Schaffer, G. A. W. [1849]1851: Systematische Be­ Zetterstedt, J. H. 1840: Insecta lapponica. - L. Voss, Leip­ arbeitung der Schmetterlinge von Europa. Vol. 4, Die zig, 1139 pp.